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<document id="8D4AD1CA4EA99614D428F7A761E82FB4" ID-DOI="10.3897/zookeys.1208.116083" ID-ZooBank="0215503D-E8B4-4179-89F6-FB2CCC59F37E" ID-publisher-id="116083" URI-arpha="CF5F342A-04A4-57D6-860A-D48A80ACC24A" URI-zoobank="http://zoobank.org/0215503D-E8B4-4179-89F6-FB2CCC59F37E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1722361871078" checkinUser="pensoft" docAuthor="Man, Nem Sian, Ablett, Jonathan D., Lwin, Ngwe, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2024" docId="76E4022A8AEF5B1DBF5C3AC31C442CA9" docLanguage="en" docName="ZooKeys 1208: 173-239" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/116083/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Glessula ponsiensis Godwin-Austen 1920" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="239" masterDocId="CF5F342A04A457D6860AD48A80ACC24A" masterDocTitle="Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar" masterLastPageNumber="239" masterPageNumber="173" pageNumber="173" updateTime="1722370176574" updateUser="admin">
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<mods:title id="263EFCC2079D541E8BDBAC1EA2D512F8">Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar</mods:title>
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<mods:namePart id="65BBA56AC2C73ACD8DC8316E2E19A8B5">Man, Nem Sian</mods:namePart>
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<mods:affiliation id="0595EF20CFFF226E36CBFD8151775CEE">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Department of Zoology, University of Yangon, University Avenue Road, Kamayut Township 11041, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="6CD479B3D663975F15AA20A563C58E77">Lwin, Ngwe</mods:namePart>
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<mods:affiliation id="1D7247EF43F86491D6AA0B3E6E52E628">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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<mods:affiliation id="4C229701B42F9EE33BB1532ECE2F6F30">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:affiliation id="0948F819DEA8E7E493C9AE9B86E9919F">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<subSubSection id="AD304E6DD039264DEA1BA9ABDC823C82" type="nomenclature">
<paragraph id="32FD2E233B0AA77B507CCACD1684AC63">
<heading id="8434D64608609F6A719BEDF33EF6A55F" reason="label">18</heading>
<taxonomicName id="27A24604D8E261278E7EDCAE8CFDB0F3" authority="Godwin-Austen, 1920" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Subulinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ponsiensis">
<emphasis id="87C21D5E07B5F593487B6F0C456B2540" italics="true">Glessula ponsiensis</emphasis>
<bibRefCitation id="BD8E05CFEA35D3B6E1207FB2190F01BB" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="BD825C5F08DFBC6C421A9CD0349EE1EE" type="description">
<paragraph id="3C266917A114F25BCC81758E9077977C">
<figureCitation id="CFBF3D4429BFCA4ED8BE8A8B60A329E2" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Glessula orophila, specimen NHMUK ex. Blanford collection from Mahableshwar B, C Glessula peguensis, specimen NHMUK 1909.3. 15.8 from Thyet-Myo D, E Glessula perlevis, syntypes NHMUK 1906.5. 5.89 from Shan Hills, Burma F Glessula ponsiensis, syntypes NHMUK 1989042 from Ponsee." figureDoi="10.3897/zookeys.1208.116083.figure10" httpUri="https://binary.pensoft.net/fig/1104508">Figs 10 F</figureCitation>
,
<figureCitation id="2E9EC63B21A02C660F298797DB8DBBA7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">11 A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="231C598B5667D7F3E1063C05C8AB6C96" type="reference_group">
<paragraph id="C5CE8E4E03975334E8DB5BC283432A92">
<treatmentCitationGroup id="0BF87D689283BE5D82DD60700E84BF5F">
<treatmentCitation id="EE29C52AD71A13A4F5B892FFBBB745CE" author="Nevill G" page="169" year="1878">
<taxonomicName id="C15A45C5EE8B7207E64CD9AF84CC8DC2" ID-CoL="8ZFKQ" authority="Nevill, 1878: 169" authorityName="Nevill" authorityPageNumber="169" authorityYear="1878" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="pyramis" subGenus="Glessula" variety="major">
<emphasis id="FC29C5162C24B777E76BE733656365A1" italics="true">Stenogyra</emphasis>
(
<emphasis id="349EC34B2D7E06E7E5DC34C1F61ADD90" italics="true">Glessula</emphasis>
)
<emphasis id="37A3C788CA457346390D7E111CB768D6" italics="true">pyramis var. major</emphasis>
<comment id="9C1EE1760F70388AE56A205D4BD98028">
<bibRefCitation id="10E8286FAC4DEE2AAF37931F2119DD2D" DOI="10.5962/bhl.title.11957" author="Nevill G" journalOrPublisher="Indian Museum, Calcutta" refId="B69" refString="Nevill G (1878) Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Indian Museum, Calcutta, 338 pp. https: // doi. org / 10.5962 / bhl. title. 11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda." year="1878">Nevill, 1878: 169</bibRefCitation>
[nomen nudum]. Type locality: Ponsee.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="D62DA8A9BB6466794B735C8EB164CB4F">
<treatmentCitationGroup id="EFFB802EB957EB09C1A564F5A83A20A5">
<taxonomicName id="79E73E1A1FBC545351589E1F31D5480A" ID-CoL="7RFKY" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ponsiensis">
<emphasis id="0C41555FCC2D62E0E65D932517FA8258" italics="true">Glessula ponsiensis</emphasis>
</taxonomicName>
<comment id="15A21C058E2850EB331F4F8FB3C17AEE">
<treatmentCitation id="1C4CDB7E97C7887E3DE28C6642151B0A" author="Godwin-Austen HH" page="56" year="1920">
<bibRefCitation id="A7979D78477AE39D701BFE157A556BA7" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920: 56</bibRefCitation>
</treatmentCitation>
, pl. 164, fig. 19. Type locality: Ponsee,
<collectingRegion id="C4FBA288B547DA43C347853DF696E6F2" country="China" name="Yunnan">Yunnan</collectingRegion>
.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDZAK" type="Type specimens">
<paragraph id="B71351886F8E3F45766B9BAB7EAADD57">
<heading id="89CB9C049C2C00394B1EA901A346720F" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="3B0209912FF4B42A03555AC70838294C">
<emphasis id="31DE5E73867AEFE05F7CD592242CAD3C" bold="true" italics="true">
<typeStatus id="859F10A465EF77F51580217A82748302" type="syntype">Syntypes</typeStatus>
</emphasis>
<named-content id="01919FC1D6511F5EDCBABF1B0E2B4568" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="D2B916E913987AF19093D4E39A26A42E" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1989042 (1 shell; Fig.
<figureCitation id="7913B4882FE6CE46593A776828EEBF9C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Glessula orophila, specimen NHMUK ex. Blanford collection from Mahableshwar B, C Glessula peguensis, specimen NHMUK 1909.3. 15.8 from Thyet-Myo D, E Glessula perlevis, syntypes NHMUK 1906.5. 5.89 from Shan Hills, Burma F Glessula ponsiensis, syntypes NHMUK 1989042 from Ponsee." figureDoi="10.3897/zookeys.1208.116083.figure10" httpUri="https://binary.pensoft.net/fig/1104508">10 F</figureCitation>
) and
<named-content id="45F9E34674F497A539527D8DA91D846A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="4E6E25793D5A93C40C847794E9361E4E" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1989043 (3 shells; Fig.
<figureCitation id="C0B06E3701699127D32F71E57BD8C7A5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">11 A</figureCitation>
), both specimen lots collected by J. Anderson from Ponsee.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EA1AK" type="material">
<paragraph id="2DE44AAD0A89F1AF733A0CFA1DB4ED45">
<heading id="6133B5CFE43C5A7383E3E246535EAAC2" reason="title">Other material.</heading>
</paragraph>
<paragraph id="2B5B6FE971943177A2B3F3CFF408964D">
<named-content id="161A62E70AFC318E27B2AEB15A0212F6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="E82DF5CC22E72953C34CBFB5B3ECBD0E" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1912.4. 16.747 (2 shells; Fig.
<figureCitation id="B58E87A465EACFDD07276A619CB6362B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">11 B</figureCitation>
) ex. Beddome collection from Ponsee,
<collectingRegion id="303D7008C0A86A37E59D50DB070B8E87" country="China" name="Yunnan">Yunnan</collectingRegion>
.
</paragraph>
<caption id="CDDEF077ECA5CFEE8EC3599385A5F9AD" ID-DOI="10.3897/zookeys.1208.116083.figure11" ID-arpha="EF10C7A6-8C41-5611-BE9F-84D6E320EC8D" httpUri="https://binary.pensoft.net/fig/1104509" startId="F11">
<paragraph id="D9154D0951A5086D06108F8825E3B464">
<label id="0AF8D9C9DF5CCA4C53272BF684067D11">Figure 11.</label>
</paragraph>
<paragraph id="F7B4E925E0B9A77731E4C8F40C669852">
<emphasis id="A152E5D5C2ACE990FBA10FE2F1EBAAA6" bold="true">A, B</emphasis>
<taxonomicName id="6E5C1C1D9C75CCD98DCEFB4A32B35679" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ponsiensis">
<emphasis id="260389FD6E419D462A86886012717C23" italics="true">Glessula ponsiensis</emphasis>
</taxonomicName>
<emphasis id="9BDC112BC5F201790CB28C344398D61C" bold="true">A</emphasis>
syntypes
<named-content id="834AEC17EDC32140BF821A2A225BB0A2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1989043 from Ponsee and
<emphasis id="B7B555DC51BDEF3696E79684BE923CDF" bold="true">B</emphasis>
specimen
<named-content id="5BC8BC8FDAB76C5D4201B95151B5CDFD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1912.4. 16.747 from Ponsee, Yunnan
<emphasis id="D77FE91EE91B0A9F5147326E2569FA2A" bold="true">C E</emphasis>
<taxonomicName id="47A813454F991515EB5027A6D93FC020" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="woodthorpei">
<emphasis id="873BDFB24FD7B1698582DF135B44BD0C" italics="true">Glessula woodthorpei</emphasis>
</taxonomicName>
, syntypes
<named-content id="2D2286117AD0E42EDBED3765252DD5BC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1903.7. 1.1628 from Shan States
<emphasis id="052A21D00477DBD9277DB2A42A8C3016" bold="true">F</emphasis>
<taxonomicName id="B83ABBF696C4B1D2739C6BCEE87C4241" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="yuangensis">
<emphasis id="0CBD1D386D7FC0B1C727B1CAADFAF938" italics="true">Glessula yuangensis</emphasis>
</taxonomicName>
, holotype
<named-content id="D7D4F11443C8477A919F3A9810ADA67D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1903.7. 1.1312 from Yuang Ha, Shan States.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E23AK" type="diagnosis">
<paragraph id="AE68F53790182477B7F7132420593883">
<heading id="35437B947B712FE15388F817FF28E9E5" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="8FCDF3AB3A8C48061E80AED4EA9F5DCE">Shell oblong turreted and regularly attenuated; spire high conical; apex rounded and blunt; subsequent whorls nearly smooth with fine growth lines, and strong, equally spaced radial ridges near suture. Suture impressed and whorls slightly convex. Aperture ovate; columella strong, concave, and truncated.</paragraph>
</subSubSection>
<subSubSection id="SECID0EA4AK" type="distribution">
<paragraph id="3C9E8841DDC334F90955D053A2FDAA1D">
<heading id="E059B44AB7A60C9FBA3FD70D42BE04D9" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="E74D6626626301745C1BDFD27E6EB19B">
<taxonomicName id="E447B5BDFC234097C59BDDA9C7B9B2DB" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ponsiensis">
<emphasis id="B0D03471FD01CC1FA8AA0183E8098E4B" italics="true">Glessula ponsiensis</emphasis>
</taxonomicName>
was first described from Ponsee,
<collectingRegion id="877A1081CE722D6AB025E57935BCB32A" country="China" name="Yunnan">Yunnan</collectingRegion>
, and later discovered in Bhamao,
<collectingRegion id="5AED9718E1695A85A9EAAA5893FADCA1" country="Myanmar" name="Kachin">Kachin State</collectingRegion>
, situated on the upper part of
<collectingRegion id="FD95371FEBB2D7FA95773C1663408FE9" country="Myanmar" name="Ayeyawady">Ayeyarwady</collectingRegion>
River.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ER4AK" type="remarks">
<paragraph id="35A781EAE97135016A4B855B3E6F0AA4">
<heading id="9DA5F8E7CE1311966A915121D3B2BCB0" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="4F80FF7268D8A854BB5C9996103ED649">
The original description included a set of shell measurements and an illustration of the apex.
<bibRefCitation id="76F261EAAC518636982E93FDA881F106" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="56" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920: 56)</bibRefCitation>
clearly states that this species was described based on specimens from the Indian Museum collection (now Zoological Survey of
<collectingCountry id="929A96C32870455C965EEF60E58A2677" name="India">India</collectingCountry>
(
<collectionCode id="57854E34E08F1A359026FCAE6F4B9F0D">ZSI</collectionCode>
), Kolkata) collected by J. Anderson from Ponsee, which was misidentified as
<taxonomicName id="F3FBB29505E2F9DBD61E9EAE3C372BDF" authority="Bs." authorityName="Bs." class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="pyramis" subGenus="Glessula">
<emphasis id="914981F2FEECBB8EE99C95BC97062BBD" italics="true">Stenogyra</emphasis>
(
<emphasis id="FD92E08D23815A7F1EAAFE9A25E0F1DD" italics="true">Glessula</emphasis>
)
<emphasis id="B84B2E6816DBF0EE0C02323E2968BC14" italics="true">pyramis</emphasis>
Bs.
</taxonomicName>
,
<taxonomicName id="C5CC666B892CE43DDB1A7B2F3685C9BE" authority="Nevill, 1878" authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="pyramis" variety="major">
var.
<emphasis id="CF0355C605E0403EF6226105E2B47C23" italics="true">major</emphasis>
<bibRefCitation id="3944966FA4134E342BEED738BE544AB1" DOI="10.5962/bhl.title.11957" author="Nevill G" journalOrPublisher="Indian Museum, Calcutta" refId="B69" refString="Nevill G (1878) Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Indian Museum, Calcutta, 338 pp. https: // doi. org / 10.5962 / bhl. title. 11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda." year="1878">Nevill, 1878</bibRefCitation>
</taxonomicName>
. The name
<taxonomicName id="0FD9C10CA7C7FCEDB13F11B8201996D4" authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="pyramis" variety="major">
var.
<emphasis id="9424D25BECCFA6BEF3994D110092E019" italics="true">major</emphasis>
</taxonomicName>
by
<bibRefCitation id="97D44BE5B67162728ABDA71CB0993858" DOI="10.5962/bhl.title.11957" author="Nevill G" firstAuthor="Nevill" journalOrPublisher="Indian Museum, Calcutta" page="169" refId="B69" refString="Nevill G (1878) Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Indian Museum, Calcutta, 338 pp. https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda." year="1878">Nevill (1878: 169)</bibRefCitation>
is unavailable (
<collectionCode id="32190037FA0169AFAC0104515D590726">ICZN</collectionCode>
1999: Art. 12). Under the note of the species, Godwin-Austen states, I am fortunate in getting the type-shells for examination. It seemed that he received on loan the type specimen (or specimens that were examined by
<bibRefCitation id="19BDB3CD0DDE534E0D354168568A237A" DOI="10.5962/bhl.title.11957" author="Nevill G" firstAuthor="Nevill" journalOrPublisher="Indian Museum, Calcutta" page="169" refId="B69" refString="Nevill G (1878) Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda. Indian Museum, Calcutta, 338 pp. https://doi.org/10.5962/bhl.title.11957" title="Hand list of Mollusca in the Indian Museum, Calcutta. Part I. Gastropoda." year="1878">Nevill (1878: 169 ))</bibRefCitation>
. The
<named-content id="FCB0156B6E3F22440584E402970C4879" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="F152759B0221342BA261F47EC858AED0" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
type collections contain a specimen lot with two registration numbers, M 10905 / 2 and 10906 / 2, which are ex-Indian Museum collection numbers where the original label states
<taxonomicName id="288C0DD2E4662244EF42AA56225B8EF8" authority="Benson" authorityName="Benson" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="pyramis" subGenus="Glessula">
<emphasis id="4746216F57DCC46B28EAAB8055E86BB9" italics="true">Stenogyra</emphasis>
(
<emphasis id="E1EB43306F70E23F03441BCEFA058F48" italics="true">Glessula</emphasis>
)
<emphasis id="DEC1F888D420E54D05781F337064B170" italics="true">pyramis</emphasis>
Benson
</taxonomicName>
,
<taxonomicName id="53CEA88B26FE20CAE490A7C9038E2B6A" authorityName="Nevill" authorityYear="1878" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="pyramis" variety="major">
var.
<emphasis id="786890ECDFE65AF818C89FDADD98CE3E" italics="true">major</emphasis>
</taxonomicName>
, collection of Dr. J. Anderson and with collection locality from Ponsee. Interestingly, a species-group name was struck through and rewritten with red ink by Godwin-Austen, handwritten as
<taxonomicName id="64EE9E6BB8EAC6BEF7524AD2A360C9BB" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ponsiensis">
<emphasis id="9284BC6B9A7B8C40E496DCEE84D5CA04" italics="true">ponsiensis</emphasis>
</taxonomicName>
, G-A. and with Type and three other one takes for
<abbrev content-type="institution" id="ABBRID0EMABK" xlink_title="used in old labels for the British Museum">
<collectionCode id="1CD36256B5CAD0AF593F0D8DE3462659">B</collectionCode>
. M
</abbrev>
. . It is probable that these specimens were presented to Godwin-Austen for examination. The specimen
<named-content id="D8194CB53CED24F76C54636CA52D6B46" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="35EB413221954CEE5E295D5D8642A106" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1989042 (1 shell has blue wool inside the shell; Fig.
<figureCitation id="CC566C8D851063102A5EA17DD413DA82" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Glessula orophila, specimen NHMUK ex. Blanford collection from Mahableshwar B, C Glessula peguensis, specimen NHMUK 1909.3. 15.8 from Thyet-Myo D, E Glessula perlevis, syntypes NHMUK 1906.5. 5.89 from Shan Hills, Burma F Glessula ponsiensis, syntypes NHMUK 1989042 from Ponsee." figureDoi="10.3897/zookeys.1208.116083.figure10" httpUri="https://binary.pensoft.net/fig/1104508">10 F</figureCitation>
) has a subsequent label stating holotype, and
<named-content id="E162F8A943A740126B31AD3C565FD198" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="0F5BE3025717298E7D1DDA8E812CD6EA" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1989043 (3 shells; Fig.
<figureCitation id="647B92D24E5F951F8D00644D3F9B9171" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">11 A</figureCitation>
) originates from the same Nevill type lot and has a subsequent label stating paratypes. This is not a valid
<typeStatus id="6658BC6011DE51ED03E098F931636974" type="holotype">holotype</typeStatus>
designation (
<collectionCode id="D84E205B44651E7B79E326D238A4430B">ICZN</collectionCode>
1999: Arts 73.1, 73.2 and Recommendation 73 F) and these specimens are considered as the
<typeStatus id="C45A0C8A881AAFB913266B8D1E74D27F" type="syntype">syntypes</typeStatus>
, illustrated herein for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>