430 lines
61 KiB
XML
430 lines
61 KiB
XML
<document id="F0426A3CA2FEB913C5CDCCD6A05862D2" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald" docDate="2022" docId="039487D1FFBBFF93FFBAFDC16DF1FF3A" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Granulatocincta theoderichi Harzhauser & Landau & Janssen 2022, nov. sp." docType="treatment" docVersion="1" lastPageNumber="60" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="58" updateTime="1709123294374" updateUser="karina">
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<mods:title id="F07296CF4E3A8532D189F94839C8926E">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="B990ADD52D141AFC69591654EAA8713F">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="A4C98A9DA80E65B4E733E25CFF188E82">Landau, Bernard</mods:namePart>
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<mods:namePart id="95922F118C697AEBE8684F83747F6337">Janssen, Ronald</mods:namePart>
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<taxonomicName id="4C3D4D44FFBBFF91FFBAFDC1699CFD45" ID-CoL="8YMMY" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[151,494,618,645]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="theoderichi" status="sp. nov.">
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<emphasis id="B949EAD5FFBBFF91FFBAFDC1699CFD45" bold="true" box="[151,494,618,645]" italics="true" pageId="57" pageNumber="58">Granulatocincta theoderichi</emphasis>
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<paragraph id="8B8236C7FFBBFF91FFBAFD246AE7FD69" blockId="57.[151,661,618,687]" box="[151,661,654,687]" pageId="57" pageNumber="58">
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<figureCitation id="13062A42FFBBFF91FFBAFD246889FD69" box="[151,251,655,681]" captionStart="FIGURE 19" captionStartId="58.[151,250,1792,1816]" captionTargetBox="[274,1314,181,1767]" captionTargetId="figure-26@58.[274,1314,181,1767]" captionTargetPageId="58" captionText="FIGURE 19 A1–A3. Granulatocincta schreibersi (Hörnes, 1854), lectotype, NHMW 1846/0037/0315, Steinebrunn (Austria). B1–B3. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW 2021/0130/0001, Enzesfeld (Austria). C1–C3. Granulatocincta schreibersi (Hörnes, 1854), NHMW 1863/0015/1131, Gainfarn (Austria). D1–D3. Granulatocincta schreibersi (Hörnes, 1854), NHMW 2021/0126/0007, Gainfarn (Austria). E1–E3. Granulatocincta theoderichi nov. sp., holotype, NHMW 2016/0026/0320, Ritzing (Austria). F1–F3. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). G1–G3. Granulatocincta theoderichi nov. sp., paratype, NHMW 1848/0003/0022a, Ritzing (Austria). H1–H3. Granulatocincta theoderichi nov. sp., paratype, NHMW 1848/0003/0022b, Ritzing (Austria)." figureDoi="http://doi.org/10.5281/zenodo.6399875" httpUri="https://zenodo.org/record/6399875/files/figure.png" pageId="57" pageNumber="58">Figs 19E</figureCitation>
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<figureCitation id="13062A42FFBBFF91FFD1FD0A6951FD69" box="[252,291,655,687]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="57" pageNumber="58">
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–E
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<figureCitation id="13062A42FFBBFF91FE0EFD0A6959FD6F" box="[291,299,673,687]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="57" pageNumber="58">
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, F
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–F
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<figureCitation id="13062A42FFBBFF91FE43FD0A6904FD6F" box="[366,374,673,687]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="57" pageNumber="58">
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, G
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–G
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<figureCitation id="13062A42FFBBFF91FEEEFD0A69B9FD6F" box="[451,459,673,687]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="57" pageNumber="58">
|
||
<subScript id="17B93482FFBBFF91FEEEFD0A69B9FD6F" attach="left" box="[451,459,673,687]" fontSize="6" pageId="57" pageNumber="58">3</subScript>
|
||
</figureCitation>
|
||
, H
|
||
<figureCitation id="13062A42FFBBFF91FEC0FD0A6A6AFD69" box="[493,536,655,687]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="57" pageNumber="58">
|
||
<subScript id="17B93482FFBBFF91FEC0FD0A6987FD6F" attach="left" box="[493,501,673,687]" fontSize="6" pageId="57" pageNumber="58">1</subScript>
|
||
–H
|
||
</figureCitation>
|
||
<figureCitation id="13062A42FFBBFF91FD35FD0A6A49FD69" box="[536,571,655,687]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="57" pageNumber="58">
|
||
<subScript id="17B93482FFBBFF91FD35FD0A6A52FD6F" attach="left" box="[536,544,673,687]" fontSize="6" pageId="57" pageNumber="58">3</subScript>
|
||
, 3
|
||
</figureCitation>
|
||
Aa, 5, 6
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFBBFF91FFBAFD7C6B7FFC0A" pageId="57" pageNumber="58" type="materials_examined">
|
||
<paragraph id="8B8236C7FFBBFF91FFBAFD7C6D13FC9E" blockId="57.[151,1437,726,2049]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FFBAFD7C6936FD30" bold="true" box="[151,324,726,753]" pageId="57" pageNumber="58">Type material.</emphasis>
|
||
<materialsCitation id="3B553C9AFFBBFF91FE67FD7C6DE7FD37" box="[330,1429,727,759]" collectionCode="MD" country="Austria" location="Ritzing" pageId="57" pageNumber="58" specimenCode="NHMW 2016" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54868865FFBBFF91FE67FD7C69C5FD31" box="[330,439,727,753]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DB9B9EBCFFBBFF91FEECFD7C6A28FD31" box="[449,602,727,753]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="57" pageNumber="58" type="Museum">NHMW 2016</specimenCode>
|
||
/0026/0320a, SL:
|
||
<quantity id="4CC59B22FFBBFF91FC0FFD7C6BF5FD31" box="[802,903,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.44" pageId="57" pageNumber="58" unit="mm" value="34.4">34.4 mm</quantity>
|
||
,
|
||
<collectionCode id="ED2CAE02FFBBFF91FCBFFD7C6BB1FD31" box="[914,963,727,753]" country="Germany" name="Museum Donaueschingen" pageId="57" pageNumber="58">MD</collectionCode>
|
||
: 13.0 mm,
|
||
<location id="8EE2601CFFBBFF91FB10FD7C6CFDFD31" LSID="urn:lsid:plazi:treatment:039487D1FFBBFF93FFBAFDC16DF1FF3A:8EE2601CFFBBFF91FB10FD7C6CFDFD31" box="[1085,1167,727,753]" country="Austria" name="Ritzing" pageId="57" pageNumber="58">Ritzing</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFBBFF91FBB3FD7C6C84FD32" box="[1182,1270,727,754]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
|
||
), figs 19E
|
||
<subScript id="17B93482FFBBFF91FA4BFD426D1CFD37" attach="right" box="[1382,1390,745,759]" fontSize="6" pageId="57" pageNumber="58">1</subScript>
|
||
–E
|
||
<subScript id="17B93482FFBBFF91FAA0FD426DE7FD37" attach="left" box="[1421,1429,745,759]" fontSize="6" pageId="57" pageNumber="58">3</subScript>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B553C9AFFBBFF91FFBAFD506A93FCD5" box="[151,737,763,789]" collectionCode="NHMW" country="Austria" pageId="57" pageNumber="58" specimenCode="NHMW 2016" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54868865FFBBFF91FFBAFD50697AFCD5" box="[151,264,763,789]" pageId="57" pageNumber="58" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCode id="DB9B9EBCFFBBFF91FE3EFD5069DCFCD5" box="[275,430,763,789]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="57" pageNumber="58" type="Museum">NHMW 2016</specimenCode>
|
||
/0026/0320b, SL:
|
||
<quantity id="4CC59B22FFBBFF91FD57FD506A93FCD5" box="[634,737,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="3.31" pageId="57" pageNumber="58" unit="mm" value="33.1">33.1 mm</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B553C9AFFBBFF91FDC0FD506C2CFCD6" box="[749,1118,763,790]" collectionCode="MD" country="Austria" location="Ritzing" pageId="57" pageNumber="58" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2CAE02FFBBFF91FDC0FD506B6CFCD5" box="[749,798,763,789]" country="Germany" name="Museum Donaueschingen" pageId="57" pageNumber="58">MD</collectionCode>
|
||
:
|
||
<quantity id="4CC59B22FFBBFF91FC04FD506BE2FCD5" box="[809,912,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="57" pageNumber="58" unit="mm" value="13.2">13.2 mm</quantity>
|
||
,
|
||
<location id="8EE2601CFFBBFF91FCB1FD506B9CFCD5" LSID="urn:lsid:plazi:treatment:039487D1FFBBFF93FFBAFDC16DF1FF3A:8EE2601CFFBBFF91FCB1FD506B9CFCD5" box="[924,1006,763,789]" country="Austria" name="Ritzing" pageId="57" pageNumber="58">Ritzing</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFBBFF91FCD3FD506C24FCD6" box="[1022,1110,763,790]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
, figs 19F
|
||
<subScript id="17B93482FFBBFF91FBEAFCA66CBDFCDB" attach="left" box="[1223,1231,781,795]" fontSize="6" pageId="57" pageNumber="58">1</subScript>
|
||
–F
|
||
<subScript id="17B93482FFBBFF91FBC1FCA66C86FCDB" attach="left" box="[1260,1268,781,795]" fontSize="6" pageId="57" pageNumber="58">2</subScript>
|
||
, 3Aa;
|
||
<materialsCitation id="3B553C9AFFBBFF91FA6CFD506A76FCF9" collectionCode="NHMW" country="Austria" pageId="57" pageNumber="58" specimenCode="NHMW 1848" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9B9EBCFFBBFF91FA6CFD5068BCFCF9" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="57" pageNumber="58" type="Museum">NHMW 1848</specimenCode>
|
||
/0003/0022a, SL:
|
||
<quantity id="4CC59B22FFBBFF91FEB1FCB56A76FCF9" box="[412,516,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="57" pageNumber="58" unit="mm" value="25.2">25.2 mm</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B553C9AFFBBFF91FD3CFCB46BF5FCFA" box="[529,903,799,826]" collectionCode="MD" country="Austria" location="Ritzing" pageId="57" pageNumber="58" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2CAE02FFBBFF91FD3CFCB46A30FCF9" box="[529,578,799,825]" country="Germany" name="Museum Donaueschingen" pageId="57" pageNumber="58">MD</collectionCode>
|
||
:
|
||
<quantity id="4CC59B22FFBBFF91FD62FCB46AC5FCF9" box="[591,695,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="57" pageNumber="58" unit="mm" value="10.3">10.3 mm</quantity>
|
||
,
|
||
<location id="8EE2601CFFBBFF91FDE9FCB46B64FCF9" LSID="urn:lsid:plazi:treatment:039487D1FFBBFF93FFBAFDC16DF1FF3A:8EE2601CFFBBFF91FDE9FCB46B64FCF9" box="[708,790,799,825]" country="Austria" name="Ritzing" pageId="57" pageNumber="58">Ritzing</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFBBFF91FC0AFCB46B0DFCFA" box="[807,895,799,826]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
, figs 19G
|
||
<subScript id="17B93482FFBBFF91FCDAFC9A6B8DFCFF" attach="left" box="[1015,1023,817,831]" fontSize="6" pageId="57" pageNumber="58">1</subScript>
|
||
–G
|
||
<subScript id="17B93482FFBBFF91FB0FFC9A6C58FCFF" attach="left" box="[1058,1066,817,831]" fontSize="6" pageId="57" pageNumber="58">3</subScript>
|
||
;
|
||
<materialsCitation id="3B553C9AFFBBFF91FB17FCB4688FFC9D" collectionCode="NHMW" country="Austria" pageId="57" pageNumber="58" specimenCode="NHMW 1848" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9B9EBCFFBBFF91FB17FCB46CA6FCF9" box="[1082,1236,799,825]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="57" pageNumber="58" type="Museum">NHMW 1848</specimenCode>
|
||
/0003/0022b, SL:
|
||
<quantity id="4CC59B22FFBBFF91FFBAFCE8688FFC9D" box="[151,253,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="57" pageNumber="58" unit="mm" value="20.3">20.3 mm</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B553C9AFFBBFF91FE24FCE86A18FC9E" box="[265,618,835,862]" collectionCode="MD" country="Austria" location="Ritzing" pageId="57" pageNumber="58" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2CAE02FFBBFF91FE24FCE86948FC9D" box="[265,314,835,861]" country="Germany" name="Museum Donaueschingen" pageId="57" pageNumber="58">MD</collectionCode>
|
||
:
|
||
<quantity id="4CC59B22FFBBFF91FE68FCE869EFFC9D" box="[325,413,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="57" pageNumber="58" unit="mm" value="8.7">8.7 mm</quantity>
|
||
,
|
||
<location id="8EE2601CFFBBFF91FE85FCE86988FC9D" LSID="urn:lsid:plazi:treatment:039487D1FFBBFF93FFBAFDC16DF1FF3A:8EE2601CFFBBFF91FE85FCE86988FC9D" box="[424,506,835,861]" country="Austria" name="Ritzing" pageId="57" pageNumber="58">Ritzing</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFBBFF91FD24FCE86A13FC9E" box="[521,609,835,862]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
, figs 19H
|
||
<subScript id="17B93482FFBBFF91FDFBFCFE6AACFCA3" attach="right" box="[726,734,853,867]" fontSize="6" pageId="57" pageNumber="58">1</subScript>
|
||
–H
|
||
<subScript id="17B93482FFBBFF91FC2DFCFE6B7AFCA3" attach="left" box="[768,776,853,867]" fontSize="6" pageId="57" pageNumber="58">3</subScript>
|
||
;
|
||
<materialsCitation id="3B553C9AFFBBFF91FC3AFCE86D2FFC9E" box="[791,1373,835,862]" collectionCode="NHMW" country="Austria" location="Ritzing" pageId="57" pageNumber="58" specimenCode="NHMW 2013" specimenCount="3" typeStatus="paratype">
|
||
<specimenCode id="DB9B9EBCFFBBFF91FC3AFCE86BC0FC9D" box="[791,946,835,861]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="57" pageNumber="58" type="Museum">NHMW 2013</specimenCode>
|
||
/0297/0012,
|
||
<specimenCount id="9D3BFD4EFFBBFF91FB12FCE86CE2FC9D" box="[1087,1168,835,861]" pageId="57" pageNumber="58" type="generic">3 spec.</specimenCount>
|
||
,
|
||
<location id="8EE2601CFFBBFF91FBB6FCE86C9FFC9D" LSID="urn:lsid:plazi:treatment:039487D1FFBBFF93FFBAFDC16DF1FF3A:8EE2601CFFBBFF91FBB6FCE86C9FFC9D" box="[1179,1261,835,861]" country="Austria" name="Ritzing" pageId="57" pageNumber="58">Ritzing</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFBBFF91FBD1FCE86D27FC9E" box="[1276,1365,835,862]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFBBFF91FFEAFCCC6B47FC41" blockId="57.[151,1437,726,2049]" box="[199,821,870,898]" pageId="57" pageNumber="58">
|
||
<materialsCitation id="3B553C9AFFBBFF91FFEAFCCC6B42FC41" box="[199,816,870,898]" country="Austria" location="Ritzing" municipality="Oberpullendorf Basin" pageId="57" pageNumber="58" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAFCCC6914FC40" bold="true" box="[199,358,870,897]" pageId="57" pageNumber="58">
|
||
<typeStatus id="54868865FFBBFF91FFEAFCCC6970FC41" box="[199,258,871,897]" pageId="57" pageNumber="58">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EE2601CFFBBFF91FE41FCCC69CCFC41" LSID="urn:lsid:plazi:treatment:039487D1FFBBFF93FFBAFDC16DF1FF3A:8EE2601CFFBBFF91FE41FCCC69CCFC41" box="[364,446,871,897]" country="Austria" municipality="Oberpullendorf Basin" name="Ritzing" pageId="57" pageNumber="58">Ritzing</location>
|
||
(
|
||
<collectingCountry id="F32A7657FFBBFF91FEE3FCCC6A55FC42" box="[462,551,871,898]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
|
||
);
|
||
<collectingMunicipality id="6BE6ACBDFFBBFF91FD17FCCC6B42FC41" box="[570,816,871,897]" pageId="57" pageNumber="58">Oberpullendorf Basin</collectingMunicipality>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFBBFF91FFEAFC206B7BFC65" blockId="57.[151,1437,726,2049]" box="[199,777,907,933]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAFC20691DFC65" bold="true" box="[199,367,907,933]" pageId="57" pageNumber="58">
|
||
<typeStatus id="54868865FFBBFF91FFEAFC206970FC65" box="[199,258,907,933]" pageId="57" pageNumber="58">Type</typeStatus>
|
||
stratum.
|
||
</emphasis>
|
||
Silty sand of the Ritzing Formation.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFBBFF91FFEAFC056B7FFC0A" blockId="57.[151,1437,726,2049]" box="[199,781,942,970]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAFC05688FFC08" bold="true" box="[199,253,942,968]" pageId="57" pageNumber="58">Age.</emphasis>
|
||
Middle Miocene, late Badenian (Serravallian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFBBFF91FFEAFC786B41FBD1" pageId="57" pageNumber="58" type="etymology">
|
||
<paragraph id="8B8236C7FFBBFF91FFEAFC786B41FBD1" blockId="57.[151,1437,726,2049]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAFC78693CFC2D" bold="true" box="[199,334,979,1005]" pageId="57" pageNumber="58">Etymology.</emphasis>
|
||
Referring to Theodoric the Great (454–526), king of the Ostrogoth, who was born in the area, which today belongs to the
|
||
<collectingRegion id="49F9F825FFBBFF91FEE1FC5C6AC5FBD1" box="[460,695,1015,1041]" country="Austria" name="Burgenland" pageId="57" pageNumber="58">Burgenland province</collectingRegion>
|
||
in
|
||
<collectingCountry id="F32A7657FFBBFF91FDF4FC5C6B5DFBD1" box="[729,815,1015,1041]" name="Austria" pageId="57" pageNumber="58">Austria</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFBBFF91FFEAFBB16CE4FBBD" pageId="57" pageNumber="58" type="diagnosis">
|
||
<paragraph id="8B8236C7FFBBFF91FFEAFBB16CE4FBBD" blockId="57.[151,1437,726,2049]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAFBB16930FBF4" bold="true" box="[199,322,1050,1076]" pageId="57" pageNumber="58">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C3D4D44FFBBFF91FE67FBB06A2CFBF5" box="[330,606,1051,1077]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FE67FBB06A71FBF5" box="[330,515,1051,1077]" italics="true" pageId="57" pageNumber="58">Granulatocincta</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of medium size, solid, slender fusiform shell, strongly coronate, weakly gradate spire, tripartite early spire whorls. Last whorl with broad tuberculate subsutural collar. Narrow subsutural ramp bearing fine spiral cord. About 12 cords below ramp bearing coarse beaded sculpture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFBBFF91FFEAFB2D6C5BF97D" pageId="57" pageNumber="58" type="description">
|
||
<paragraph id="8B8236C7FFBBFF91FFEAFB2D6C5BF97D" blockId="57.[151,1437,726,2049]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAFB2D692BFB60" bold="true" box="[199,345,1158,1184]" pageId="57" pageNumber="58">Description.</emphasis>
|
||
Shell medium-sized, solid, fusiform, with strongly coronate, weakly gradate spire; apical angle ~40–45°. Protoconch not preserved. Teleoconch of nine whorls. Earliest teleoconch whorls flat-sided with tripartite sculpture; indistinctly beaded subsutural cord, weakly beaded central cord. Coarsely beaded suprasutural cord. Sculpture changing gradually from fourth whorl; prominent, swollen subsutural collar with pointed tubercles (14– 16 on last whorl) developed on adapical half of collar, central row of beads coalesce to form smooth, narrow central cord. Suprasutural row of beads further strengthens, overrun by one or two weak, narrow cords. Whorl profile on later whorls subcylindrical with narrow central concavity filled by central cord. Suture narrowly incised, undulating below beads. Last whorl ~60% of total height. Subsutural collar broad, swollen, sharply delimited, bearing small, pointed tubercles. Subsutural ramp narrow, concave, bearing single smooth, narrow cord, shoulder delimited by beaded cord, broadly concave below, weakly constricted at base. Siphonal fasciole strongly swollen, twisted. Sculpture below ramp of about 12 beaded spiral cords, separated by narrow grooves at mid-whorl, becoming wider spaced towards fasciole. Narrow, unbeaded cords over fasciole. Prominent axial growth lines. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately narrow and deep, symmetrically Ushaped, with apex on central cord. Siphonal canal moderately long, moderately narrow, straight, shallowly notched at tip. Columella strongly excavated in upper third, straight below, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim and pseudumbilical chink.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFBBFF93FFEAF96D6A26FF16" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58" type="discussion">
|
||
<paragraph id="8B8236C7FFBBFF91FFEAF96D6CAEF8E9" blockId="57.[151,1437,726,2049]" pageId="57" pageNumber="58">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAF96D693EF920" bold="true" box="[199,332,1734,1760]" pageId="57" pageNumber="58">Discussion.</emphasis>
|
||
This species becomes gradually more elongate due to the ontogentic increase of the height of the last whorl. It is quite variable in sculpture, especially concerning the strength of the tubercles. This variability might be an expression of the unstable ecological conditions in the intertidal to shallow sublittoral zone.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFBBFF91FFEAF8986B43F7C1" blockId="57.[151,1437,726,2049]" pageId="57" pageNumber="58">
|
||
<taxonomicName id="4C3D4D44FFBBFF91FFEAF8986A7AF88D" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[199,520,1843,1869]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="theoderichi">
|
||
<emphasis id="B949EAD5FFBBFF91FFEAF8986A7AF88D" box="[199,520,1843,1869]" italics="true" pageId="57" pageNumber="58">Granulatocincta theoderichi</emphasis>
|
||
</taxonomicName>
|
||
is unique within the Paratethyan
|
||
<taxonomicName id="4C3D4D44FFBBFF91FCA8F8986C61F88D" authorityName="Gray" authorityYear="1853" box="[901,1043,1843,1869]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="57" pageNumber="58" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
due to its coarse sculpture.
|
||
<taxonomicName id="4C3D4D44FFBBFF91FA7DF8986A42F8B1" authority="(Hoernes & Auinger, 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="agathae">
|
||
<emphasis id="B949EAD5FFBBFF91FA7DF8986883F8B1" italics="true" pageId="57" pageNumber="58">Olegia agathae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFBBFF91FE2CF8FC6A5AF8B1" author="Hoernes, R. & Auinger, M." box="[257,552,1879,1905]" pageId="57" pageNumber="58" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is superficially similar, but differs in its less protruding subsutural collar and much wider central concavity of the spire whorls. In addition,
|
||
<taxonomicName id="4C3D4D44FFBBFF91FC71F8D06BA8F855" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[860,986,1915,1941]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="agathae">
|
||
<emphasis id="B949EAD5FFBBFF91FC71F8D06BA8F855" box="[860,986,1915,1941]" italics="true" pageId="57" pageNumber="58">O. agathae</emphasis>
|
||
</taxonomicName>
|
||
has fewer spiral cords on the base. The same features distinguish
|
||
<taxonomicName id="4C3D4D44FFBBFF91FE97F8346A1AF879" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[442,616,1951,1977]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="mandici" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFBBFF91FE97F8346A1AF879" box="[442,616,1951,1977]" italics="true" pageId="57" pageNumber="58">Olegia mandici</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFBBFF91FD42F8346AB4F879" box="[623,710,1951,1977]" pageId="57" pageNumber="58" rank="species">nov. sp.</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName id="4C3D4D44FFBBFF91FC26F8346BDCF879" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[779,942,1951,1977]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="theoderichi">
|
||
<emphasis id="B949EAD5FFBBFF91FC26F8346BDCF879" box="[779,942,1951,1977]" italics="true" pageId="57" pageNumber="58">G. theoderichi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C3D4D44FFBBFF91FC96F8346D3EF87A" authority="(Strausz, 1960)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Strausz" baseAuthorityYear="1960" box="[955,1356,1951,1978]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="szokolyensis">
|
||
<emphasis id="B949EAD5FFBBFF91FC96F8346CEBF879" box="[955,1177,1951,1977]" italics="true" pageId="57" pageNumber="58">Olegia szokolyensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFBBFF91FB84F8346D36F87A" author="Strausz, L." box="[1193,1348,1951,1978]" pageId="57" pageNumber="58" pagination="348 - 354" refId="ref144947" refString="Strausz, L. (1960) Uj nevek es uj alakok a Miocen Puhatestuek Kozt. Foldtani Kozlony, 1960, 348 - 354. [http: // epa. niif. hu / 01600 / 01635 / 00154 / pdf / EPA 01635 _ foldtani _ kozlony _ 1960 _ 3 _ 348 - 354. pdf]" type="journal article" year="1960">Strausz, 1960</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
differs in its more conical spire whorls and the much weaker spiral sculpture on the base. A placement in
|
||
<taxonomicName id="4C3D4D44FFBBFF91FBE0F8686D6BF81D" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[1229,1305,1987,2013]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949EAD5FFBBFF91FBE0F8686D6BF81D" box="[1229,1305,1987,2013]" italics="true" pageId="57" pageNumber="58">Olegia</emphasis>
|
||
</taxonomicName>
|
||
is excluded based on the tripartite sculpture of early teleoconch whorls.
|
||
</paragraph>
|
||
<caption id="DF42664FFFB8FF92FFBAF8AB6C3BF830" ID-DOI="http://doi.org/10.5281/zenodo.6399875" ID-Zenodo-Dep="6399875" httpUri="https://zenodo.org/record/6399875/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,250,1792,1816]" targetBox="[274,1314,181,1767]" targetPageId="58">
|
||
<paragraph id="8B8236C7FFB8FF92FFBAF8AB6C3BF830" blockId="58.[151,1437,1792,2032]" pageId="58" pageNumber="59">
|
||
<emphasis id="B949EAD5FFB8FF92FFBAF8AB691AF8D8" bold="true" box="[151,360,1792,1821]" pageId="58" pageNumber="59">
|
||
FIGURE 19 A
|
||
<subScript id="17B93482FFB8FF92FE18F8A4694EF8DD" attach="right" box="[309,316,1807,1821]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–A
|
||
<subScript id="17B93482FFB8FF92FE76F8A46910F8DD" attach="left" box="[347,354,1807,1821]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FE5DF8AB6B41F8D8" authority="(Hornes, 1854)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" box="[368,819,1792,1816]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="schreibersi">
|
||
<emphasis id="B949EAD5FFB8FF92FE5DF8AB6AFDF8D8" box="[368,655,1792,1816]" italics="true" pageId="58" pageNumber="59">Granulatocincta schreibersi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFB8FF92FDB3F8AB6B5EF8D8" author="Hornes, M." box="[670,812,1792,1816]" pageId="58" pageNumber="59" pagination="297 - 382" refId="ref135553" refString="Hornes, M. (1854) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der K. K. Geologischen Reichsanstalt, 3, 297 - 382, pls. XXXIII - XL (1854). [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1854">Hörnes, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype, NHMW 1846/0037/0315, Steinebrunn (Austria).
|
||
<emphasis id="B949EAD5FFB8FF92FFBAF88F68A8F8FC" bold="true" box="[151,218,1828,1857]" pageId="58" pageNumber="59">
|
||
B
|
||
<subScript id="17B93482FFB8FF92FF85F89868DDF881" attach="left" box="[168,175,1843,1857]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–B
|
||
<subScript id="17B93482FFB8FF92FFE0F89868A6F881" attach="left" box="[205,212,1843,1857]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FFCAF88F6AC4F8FC" authority="(Hornes, 1854)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" box="[231,694,1828,1852]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="schreibersi">
|
||
<emphasis id="B949EAD5FFB8FF92FFCAF88F6A78F8FC" box="[231,522,1828,1852]" italics="true" pageId="58" pageNumber="59">Granulatocincta schreibersi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFB8FF92FD30F88F6ADDF8FC" author="Hornes, M." box="[541,687,1828,1852]" pageId="58" pageNumber="59" pagination="297 - 382" refId="ref135553" refString="Hornes, M. (1854) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der K. K. Geologischen Reichsanstalt, 3, 297 - 382, pls. XXXIII - XL (1854). [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1854">Hörnes, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, paralectotype, NHMW 2021/0130/0001, Enzesfeld (Austria).
|
||
<emphasis id="B949EAD5FFB8FF92FA7BF88F6DEEF8FC" bold="true" box="[1366,1436,1828,1857]" pageId="58" pageNumber="59">
|
||
C
|
||
<subScript id="17B93482FFB8FF92FA45F8986D1DF881" attach="left" box="[1384,1391,1843,1857]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–C
|
||
<subScript id="17B93482FFB8FF92FAA2F8986DE4F881" attach="right" box="[1423,1430,1843,1857]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FFBAF8E36A24F8A0" authority="(Hornes, 1854)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" box="[151,598,1864,1888]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="schreibersi">
|
||
<emphasis id="B949EAD5FFB8FF92FFBAF8E369C7F8A0" box="[151,437,1864,1888]" italics="true" pageId="58" pageNumber="59">Granulatocincta schreibersi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFB8FF92FEEFF8E36A3DF8A0" author="Hornes, M." box="[450,591,1864,1888]" pageId="58" pageNumber="59" pagination="297 - 382" refId="ref135553" refString="Hornes, M. (1854) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der K. K. Geologischen Reichsanstalt, 3, 297 - 382, pls. XXXIII - XL (1854). [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1854">Hörnes, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1863/0015/1131, Gainfarn (Austria).
|
||
<emphasis id="B949EAD5FFB8FF92FB1EF8E36C0BF8A0" bold="true" box="[1075,1145,1864,1893]" pageId="58" pageNumber="59">
|
||
D
|
||
<subScript id="17B93482FFB8FF92FB68F8FC6C3EF8A5" attach="left" box="[1093,1100,1879,1893]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–D
|
||
<subScript id="17B93482FFB8FF92FB46F8FC6C00F8A5" attach="left" box="[1131,1138,1879,1893]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FB52F8E36941F844" authority="(Hornes, 1854)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="schreibersi">
|
||
<emphasis id="B949EAD5FFB8FF92FB52F8E36DEFF8A0" box="[1151,1437,1864,1888]" italics="true" pageId="58" pageNumber="59">Granulatocincta schreibersi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFB8FF92FFB3F8C7695EF844" author="Hornes, M." box="[158,300,1900,1924]" pageId="58" pageNumber="59" pagination="297 - 382" refId="ref135553" refString="Hornes, M. (1854) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der K. K. Geologischen Reichsanstalt, 3, 297 - 382, pls. XXXIII - XL (1854). [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1854">Hörnes, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 2021/0126/0007, Gainfarn (Austria).
|
||
<emphasis id="B949EAD5FFB8FF92FC38F8C76B2AF844" bold="true" box="[789,856,1900,1929]" pageId="58" pageNumber="59">
|
||
E
|
||
<subScript id="17B93482FFB8FF92FC0BF8D06B5FF849" attach="left" box="[806,813,1915,1929]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–E
|
||
<subScript id="17B93482FFB8FF92FC66F8D06B20F849" attach="left" box="[843,850,1915,1929]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FC72F8C76CF3F844" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[863,1153,1900,1924]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="theoderichi" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB8FF92FC72F8C76CF3F844" box="[863,1153,1900,1924]" italics="true" pageId="58" pageNumber="59">Granulatocincta theoderichi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB8FF92FBA5F8C76CAAF843" box="[1160,1240,1900,1923]" pageId="58" pageNumber="59" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, holotype, NHMW 2016/0026/0320, Ritzing (Austria).
|
||
<emphasis id="B949EAD5FFB8FF92FD2BF83B6A34F868" bold="true" box="[518,582,1936,1965]" pageId="58" pageNumber="59">
|
||
F
|
||
<subScript id="17B93482FFB8FF92FD38F8346A6EF86D" attach="left" box="[533,540,1951,1965]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–F
|
||
<subScript id="17B93482FFB8FF92FD14F8346A32F86D" attach="right" box="[569,576,1951,1965]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FD60F83B6B1DF868" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[589,879,1936,1960]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="theoderichi" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB8FF92FD60F83B6B1DF868" box="[589,879,1936,1960]" italics="true" pageId="58" pageNumber="59">Granulatocincta theoderichi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB8FF92FC5AF83B6BB5F867" box="[887,967,1936,1959]" pageId="58" pageNumber="59" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, paratype, NHMW 2016/0026/0320b, Ritzing (Austria).
|
||
<emphasis id="B949EAD5FFB8FF92FE2BF81F693DF80C" bold="true" box="[262,335,1972,2001]" pageId="58" pageNumber="59">
|
||
G
|
||
<subScript id="17B93482FFB8FF92FE37F8686953F811" attach="left" box="[282,289,1987,2001]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–G
|
||
<subScript id="17B93482FFB8FF92FE6FF868693BF811" attach="left" box="[322,329,1987,2001]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FE70F81F6AF7F80C" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[349,645,1972,1996]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="theoderichi" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB8FF92FE70F81F6AF7F80C" box="[349,645,1972,1996]" italics="true" pageId="58" pageNumber="59">Granulatocincta theoderichi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB8FF92FDBEF81F6A98F80B" box="[659,746,1972,1995]" pageId="58" pageNumber="59" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, paratype, NHMW 1848/0003/0022a, Ritzing (Austria).
|
||
<emphasis id="B949EAD5FFB8FF92FA7EF81F6DEEF80C" bold="true" box="[1363,1436,1972,2001]" pageId="58" pageNumber="59">
|
||
H
|
||
<subScript id="17B93482FFB8FF92FA4AF8686D1CF811" attach="left" box="[1383,1390,1987,2001]" fontSize="6" pageId="58" pageNumber="59">1</subScript>
|
||
–H
|
||
<subScript id="17B93482FFB8FF92FAA2F8686DE4F811" attach="left" box="[1423,1430,1987,2001]" fontSize="6" pageId="58" pageNumber="59">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB8FF92FFBAF87369CAF830" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[151,440,2008,2032]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="58" pageNumber="59" phylum="Mollusca" rank="species" species="theoderichi" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB8FF92FFBAF87369CAF830" box="[151,440,2008,2032]" italics="true" pageId="58" pageNumber="59">Granulatocincta theoderichi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB8FF92FE92F8736A7DF82F" box="[447,527,2008,2031]" pageId="58" pageNumber="59" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, paratype, NHMW 1848/0003/0022b, Ritzing (Austria).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8236C7FFB9FF93FFEAFF3D6A26FF16" blockId="59.[151,1436,150,250]" pageId="59" pageNumber="60">
|
||
<emphasis id="B949EAD5FFB9FF93FFEAFF3D69DAFF70" bold="true" box="[199,424,150,176]" pageId="59" pageNumber="60">Paleoenvironment.</emphasis>
|
||
Coastal, intertidal to inner neritic sandy bottoms in a restricted lagoon based on lithology and molluscan assemblages (own data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFB9FF93FFEAFF756DF1FF3A" box="[199,1411,222,250]" pageId="59" pageNumber="60" type="distribution">
|
||
<paragraph id="8B8236C7FFB9FF93FFEAFF756DF1FF3A" blockId="59.[151,1436,150,250]" box="[199,1411,222,250]" pageId="59" pageNumber="60">
|
||
<emphasis id="B949EAD5FFB9FF93FFEAFF756A18FF39" bold="true" box="[199,618,222,249]" pageId="59" pageNumber="60">Distribution in Central Paratethys.</emphasis>
|
||
Badenian (middle Miocene):
|
||
<emphasis id="B949EAD5FFB9FF93FC94FF746CDDFF38" box="[953,1199,223,249]" italics="true" pageId="59" pageNumber="60">Oberpullendorf Basin</emphasis>
|
||
: Ritzing (
|
||
<collectingCountry id="F32A7657FFB9FF93FA32FF746D0AFF3A" box="[1311,1400,223,250]" name="Austria" pageId="59" pageNumber="60">Austria</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |