The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera Author Harzhauser, Mathias 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at Author Landau, Bernard 0000-0002-7768-8494 bernardmlandau@gmail.com Author Janssen, Ronald 0000-0002-4471-6655 mathias.harzhauser@nhm-wien.ac.at text Zootaxa 2022 2022-03-30 5123 1 1 172 http://dx.doi.org/10.11646/zootaxa.5123.1.1 journal article 10.11646/zootaxa.5123.1.1 1175-5326 6399827 036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A Granulatocincta theoderichi nov. sp. Figs 19E 1 –E 3 , F 1 –F 3 , G 1 –G 3 , H 1 –H 3 , 3 Aa, 5, 6 Type material. Holotype : NHMW 2016 /0026/0320a, SL: 34.4 mm , MD : 13.0 mm, Ritzing ( Austria ), figs 19E 1 –E 3 . Paratypes : NHMW 2016 /0026/0320b, SL: 33.1 mm , MD : 13.2 mm , Ritzing ( Austria ) , figs 19F 1 –F 2 , 3Aa; NHMW 1848 /0003/0022a, SL: 25.2 mm , MD : 10.3 mm , Ritzing ( Austria ) , figs 19G 1 –G 3 ; NHMW 1848 /0003/0022b, SL: 20.3 mm , MD : 8.7 mm , Ritzing ( Austria ) , figs 19H 1 –H 3 ; NHMW 2013 /0297/0012, 3 spec. , Ritzing ( Austria ) . Type locality. Ritzing ( Austria ); Oberpullendorf Basin . Type stratum. Silty sand of the Ritzing Formation. Age. Middle Miocene, late Badenian (Serravallian). Etymology. Referring to Theodoric the Great (454–526), king of the Ostrogoth, who was born in the area, which today belongs to the Burgenland province in Austria . Diagnosis. Granulatocincta species of medium size, solid, slender fusiform shell, strongly coronate, weakly gradate spire, tripartite early spire whorls. Last whorl with broad tuberculate subsutural collar. Narrow subsutural ramp bearing fine spiral cord. About 12 cords below ramp bearing coarse beaded sculpture. Description. Shell medium-sized, solid, fusiform, with strongly coronate, weakly gradate spire; apical angle ~40–45°. Protoconch not preserved. Teleoconch of nine whorls. Earliest teleoconch whorls flat-sided with tripartite sculpture; indistinctly beaded subsutural cord, weakly beaded central cord. Coarsely beaded suprasutural cord. Sculpture changing gradually from fourth whorl; prominent, swollen subsutural collar with pointed tubercles (14– 16 on last whorl) developed on adapical half of collar, central row of beads coalesce to form smooth, narrow central cord. Suprasutural row of beads further strengthens, overrun by one or two weak, narrow cords. Whorl profile on later whorls subcylindrical with narrow central concavity filled by central cord. Suture narrowly incised, undulating below beads. Last whorl ~60% of total height. Subsutural collar broad, swollen, sharply delimited, bearing small, pointed tubercles. Subsutural ramp narrow, concave, bearing single smooth, narrow cord, shoulder delimited by beaded cord, broadly concave below, weakly constricted at base. Siphonal fasciole strongly swollen, twisted. Sculpture below ramp of about 12 beaded spiral cords, separated by narrow grooves at mid-whorl, becoming wider spaced towards fasciole. Narrow, unbeaded cords over fasciole. Prominent axial growth lines. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately narrow and deep, symmetrically Ushaped, with apex on central cord. Siphonal canal moderately long, moderately narrow, straight, shallowly notched at tip. Columella strongly excavated in upper third, straight below, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim and pseudumbilical chink. Discussion. This species becomes gradually more elongate due to the ontogentic increase of the height of the last whorl. It is quite variable in sculpture, especially concerning the strength of the tubercles. This variability might be an expression of the unstable ecological conditions in the intertidal to shallow sublittoral zone. Granulatocincta theoderichi is unique within the Paratethyan Clavatulidae due to its coarse sculpture. Olegia agathae ( Hoernes & Auinger, 1891 ) is superficially similar, but differs in its less protruding subsutural collar and much wider central concavity of the spire whorls. In addition, O. agathae has fewer spiral cords on the base. The same features distinguish Olegia mandici nov. sp. from G. theoderichi . Olegia szokolyensis ( Strausz, 1960 ) differs in its more conical spire whorls and the much weaker spiral sculpture on the base. A placement in Olegia is excluded based on the tripartite sculpture of early teleoconch whorls. FIGURE 19 A 1 –A 3 . Granulatocincta schreibersi ( Hörnes, 1854 ) , lectotype, NHMW 1846/0037/0315, Steinebrunn (Austria). B 1 –B 3 . Granulatocincta schreibersi ( Hörnes, 1854 ) , paralectotype, NHMW 2021/0130/0001, Enzesfeld (Austria). C 1 –C 3 . Granulatocincta schreibersi ( Hörnes, 1854 ) , NHMW 1863/0015/1131, Gainfarn (Austria). D 1 –D 3 . Granulatocincta schreibersi ( Hörnes, 1854 ) , NHMW 2021/0126/0007, Gainfarn (Austria). E 1 –E 3 . Granulatocincta theoderichi nov. sp. , holotype, NHMW 2016/0026/0320, Ritzing (Austria). F 1 –F 3 . Granulatocincta theoderichi nov. sp. , paratype, NHMW 2016/0026/0320b, Ritzing (Austria). G 1 –G 3 . Granulatocincta theoderichi nov. sp. , paratype, NHMW 1848/0003/0022a, Ritzing (Austria). H 1 –H 3 . Granulatocincta theoderichi nov. sp. , paratype, NHMW 1848/0003/0022b, Ritzing (Austria). Paleoenvironment. Coastal, intertidal to inner neritic sandy bottoms in a restricted lagoon based on lithology and molluscan assemblages (own data). Distribution in Central Paratethys. Badenian (middle Miocene): Oberpullendorf Basin : Ritzing ( Austria ).