199 lines
19 KiB
XML
199 lines
19 KiB
XML
<document id="6DAF010887E316CFE5F15263517E9E33" ID-DOI="10.5281/zenodo.3677235" ID-GBIF-Dataset="1ee19d5d-27cf-4c93-989c-3fe3eb467203" ID-ISSN="1942-1354" ID-Zenodo-Dep="3677235" ID-ZooBank="BA35690A-FC73-4E5A-A805-FE9550275FEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582218729488" checkinUser="felipe" docAuthor="Cong, Qian, Zhang, Jing, Shen, Jinhui & Grishin, Nick V." docDate="2019" docId="038E29227E7EFFF2FF3F7031DCDBFA0D" docLanguage="en" docName="InsectaMundi.0731.1-56.pdf.imf" docOrigin="Insecta Mundi 731 (731)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Incisus Grishin 2019, new genus" docType="treatment" docUuid="2ED74AC8-4CCF-4796-952B-0292620F0855" docUuidSource="ZooBank" docVersion="11" lastPageNumber="11" masterDocId="FFB7515A7E72FFFEFFFF732ADF1EFFFB" masterDocTitle="Fifty new genera of Hesperiidae (Lepidoptera)" masterLastPageNumber="56" masterPageNumber="1" pageNumber="11" updateTime="1698720609646" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="460A3B47EBA675EC9EC70A0C7B4F867C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1344E47340CCD22193BD75C0419E3056">
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<mods:title id="C2CB90BD320F4C97107D83BF0EE3C290">Fifty new genera of Hesperiidae (Lepidoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="4537C870BE957B0C9FAE76CE2EE0B4B3" type="personal">
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<mods:roleTerm id="9719EB3A08FD0271BFF1F36602302176">Author</mods:roleTerm>
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<mods:namePart id="05DADDE2984E07E1164E922890E4B684">Cong, Qian</mods:namePart>
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<mods:namePart id="8D638186AE2C2294827B87D4DEA8B4C7">Zhang, Jing</mods:namePart>
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<mods:namePart id="17AFE5ECB6A3FC5D11A8CD4723AC1EBA">Shen, Jinhui</mods:namePart>
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</mods:name>
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<mods:name id="E313EF79F14B5537BBFF82C071457F93" type="personal">
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<mods:roleTerm id="1B2FEFF362EA468C9D87914F8F9A1F78">Author</mods:roleTerm>
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<mods:namePart id="9B2AA04EFA5FEFB8584F671DE886E477">Grishin, Nick V.</mods:namePart>
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<mods:typeOfResource id="6A012A8A0703D89E4A7F4873A72E9C52">text</mods:typeOfResource>
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<mods:title id="7992F5CA67F9A6A9BE8AA70D3CB832A5">Insecta Mundi</mods:title>
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<mods:part id="1E76EB1610E6BBB3FA1FF6FCBD7DB323">
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<mods:date id="EEE428A7D1408D4B84525A8E19D14B27">2019</mods:date>
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<mods:number id="901F7939D934B06928DB85EFD58C714C">2019-10-11</mods:number>
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<mods:identifier id="A1AAF7A1A5D4B4C592E2EC07CA2B46A8" type="ISSN">1942-1354</mods:identifier>
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<treatment id="038E29227E7EFFF2FF3F7031DCDBFA0D" ID-DOI="http://doi.org/10.5281/zenodo.3681703" ID-GBIF-Taxon="162238025" ID-Zenodo-Dep="3681703" LSID="urn:lsid:zoobank.org:act:2ED74AC8-4CCF-4796-952B-0292620F0855" httpUri="http://treatment.plazi.org/id/038E29227E7EFFF2FF3F7031DCDBFA0D" lastPageNumber="11" pageId="12" pageNumber="11">
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<subSubSection id="C33DCBBF7E7EFFF2FF3F7031DD2DFCCF" box="[192,563,795,820]" pageId="12" pageNumber="11" type="nomenclature">
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<paragraph id="8B9898347E7EFFF2FF3F7031DD2DFCCF" blockId="12.[192,563,795,820]" box="[192,563,795,820]" pageId="12" pageNumber="11">
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<heading id="D0D02F587E7EFFF2FF3F7031DD2DFCCF" bold="true" box="[192,563,795,820]" fontSize="10" level="2" pageId="12" pageNumber="11" reason="3">
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<emphasis id="B95344267E7EFFF2FF3F7031DD2DFCCF" bold="true" box="[192,563,795,820]" pageId="12" pageNumber="11">
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<taxonomicName id="4C27E3B77E7EFFF2FF3F7031DE8DFCC8" authority="Grishin" authorityName="Grishin" authorityYear="2019" box="[192,403,795,819]" class="Insecta" family="Hesperiidae" genus="Incisus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B95344267E7EFFF2FF3F7031DE3CFCC8" bold="true" box="[192,290,795,819]" italics="true" pageId="12" pageNumber="11">Incisus</emphasis>
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Grishin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A260F95D7E7EFFF2FE5E7036DD2DFCCF" box="[417,563,796,820]" pageId="12" pageNumber="11" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33DCBBF7E7EFFF2FF3F7062DCDDFC9B" box="[192,963,840,864]" pageId="12" pageNumber="11" type="description">
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<paragraph id="8B9898347E7EFFF2FF3F7062DCDDFC9B" blockId="12.[192,963,840,864]" box="[192,963,840,864]" pageId="12" pageNumber="11">
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http://zoobank.org/
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<uuid id="FF81A2E17E7EFFF2FE607062DCDDFC9B" box="[415,963,840,864]" pageId="12" pageNumber="11">2ED74AC8-4CCF-4796-952B-0292620F0855</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33DCBBF7E7EFFF2FF3F70A4DB82FB0D" pageId="12" pageNumber="11" type="type_taxon">
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<paragraph id="8B9898347E7EFFF2FF3F70A4DC12FC5D" blockId="12.[192,780,910,934]" box="[192,780,910,934]" pageId="12" pageNumber="11">
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<emphasis id="B95344267E7EFFF2FF3F70A4DE68FC5D" bold="true" box="[192,374,910,934]" pageId="12" pageNumber="11">
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<typeStatus id="549C26967E7EFFF2FF3F70A4DE1AFC5D" box="[192,260,910,934]" pageId="12" pageNumber="11">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C27E3B77E7EFFF2FE8170A4DC17FC5D" ID-CoL="95WVC" authority="Mabille, 1878" authorityName="Mabille" authorityYear="1878" box="[382,777,910,934]" class="Insecta" family="Hesperiidae" genus="Antigonus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="incisus">
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<emphasis id="B95344267E7EFFF2FE8170A4DD49FC5E" box="[382,599,910,933]" italics="true" pageId="12" pageNumber="11">Antigonus incisus</emphasis>
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Mabille, 1878
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B9898347E7EFFF2FF3F7094DB82FB0D" blockId="12.[192,1440,958,1270]" pageId="12" pageNumber="11">
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<emphasis id="B95344267E7EFFF2FF3F7094DE4CFC2D" bold="true" box="[192,338,958,982]" pageId="12" pageNumber="11">Definition.</emphasis>
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A genus without close relatives. Formerly placed in
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<taxonomicName id="4C27E3B77E7EFFF2FC357094DA84FC2D" ID-CoL="92N65" authority="Godman and Salvin, [1896]" authorityName="Godman and Salvin" authorityYear="1896" box="[970,1434,958,982]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B95344267E7EFFF2FC357094DB59FC2D" box="[970,1095,958,982]" italics="true" pageId="12" pageNumber="11">Staphylus</emphasis>
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Godman and Salvin, [1896]
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</taxonomicName>
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,
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<taxonomicName id="4C27E3B77E7EFFF2FF3F70F4DE09FC0E" ID-CoL="95WVC" authorityName="Mabille" authorityYear="1878" box="[192,279,990,1013]" class="Insecta" family="Hesperiidae" genus="Antigonus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="incisus">
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<emphasis id="B95344267E7EFFF2FF3F70F4DE09FC0E" box="[192,279,990,1013]" italics="true" pageId="12" pageNumber="11">incisus</emphasis>
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</taxonomicName>
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is not monophyletic with the
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<taxonomicName id="4C27E3B77E7EFFF2FD7970F4DC1AFC0D" ID-CoL="92N65" authorityName="Godman and Salvin" authorityYear="1896" box="[646,772,990,1014]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B95344267E7EFFF2FD7970F4DC1AFC0D" box="[646,772,990,1014]" italics="true" pageId="12" pageNumber="11">Staphylus</emphasis>
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</taxonomicName>
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<typeStatus id="549C26967E7EFFF2FCF470F4DC5FFC0D" box="[779,833,990,1014]" pageId="12" pageNumber="11">type</typeStatus>
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species
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<taxonomicName id="4C27E3B77E7EFFF2FC5770F4DA74FC0D" authority="Staudinger, 1876" authorityName="Staudinger" authorityYear="1876" box="[936,1386,990,1014]" class="Insecta" family="Hesperiidae" genus="Helias" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="ascalaphus">
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<emphasis id="B95344267E7EFFF2FC5770F4DB95FC0D" box="[936,1163,990,1014]" italics="true" pageId="12" pageNumber="11">Helias ascalaphus</emphasis>
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Staudinger, 1876
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</taxonomicName>
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, nor with
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<taxonomicName id="4C27E3B77E7EFFF2FEFD70D4DC37FBED" authority="Godman and Salvin, [1896]" authorityName="Godman and Salvin" authorityYear="1896" box="[258,809,1022,1046]" class="Insecta" family="Hesperiidae" genus="Scantilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="opites">
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<emphasis id="B95344267E7EFFF2FEFD70D4DEDAFBEE" box="[258,452,1022,1046]" italics="true" pageId="12" pageNumber="11">Scantilla opites</emphasis>
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Godman and Salvin, [1896]
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</taxonomicName>
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(a junior subjective synonym of
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<taxonomicName id="4C27E3B77E7EFFF2FB3670D4DE55FBCD" ID-CoL="949WH" authority="Plotz, 1886" authorityName="Plotz" authorityYear="1886" class="Insecta" family="Hesperiidae" genus="Tagiades" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="vincula">
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<emphasis id="B95344267E7EFFF2FB3670D4DA81FBED" box="[1225,1439,1022,1046]" italics="true" pageId="12" pageNumber="11">Tagiades vincula</emphasis>
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Plötz, 1886
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</taxonomicName>
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) the
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<typeStatus id="549C26967E7EFFF2FE757734DEDEFBCD" box="[394,448,1054,1078]" pageId="12" pageNumber="11">type</typeStatus>
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species of
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<taxonomicName id="4C27E3B77E7EFFF2FDBB7734DB0CFBCD" authority="Godman and Salvin, [1896]" authorityName="Godman and Salvin" authorityYear="1896" box="[580,1042,1054,1078]" class="Insecta" family="Hesperiidae" genus="Scantilla" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B95344267E7EFFF2FDBB7734DDA8FBCD" box="[580,694,1054,1078]" italics="true" pageId="12" pageNumber="11">Scantilla</emphasis>
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Godman and Salvin, [1896]
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</taxonomicName>
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, a genus-group name treated as a subjective synonym of
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<taxonomicName id="4C27E3B77E7EFFF2FE0E7714DD71FBAD" ID-CoL="92N65" authorityName="Godman and Salvin" authorityYear="1896" box="[497,623,1086,1110]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B95344267E7EFFF2FE0E7714DD71FBAD" box="[497,623,1086,1110]" italics="true" pageId="12" pageNumber="11">Staphylus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="131C84B17E7EFFF2FD7F7714DDD5FBAD" box="[640,715,1086,1110]" captionStart="Figure 4" captionStartId="11.[192,275,1906,1928]" captionTargetBox="[193,1440,780,1883]" captionTargetId="figure@11.[192,1440,779,1883]" captionTargetPageId="11" captionText="Figure 4. Genomic trees of Carcharodini. See Fig. 2 legend for notations." figureDoi="http://doi.org/10.5281/zenodo.3677243" httpUri="https://zenodo.org/record/3677243/files/figure.png" pageId="12" pageNumber="11">Fig. 4</figureCitation>
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). Keys to E.32.35b in
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<bibRefCitation id="EFB6E5C57E7EFFF2FC237715DB9AFBAD" author="Evans" box="[988,1156,1086,1110]" firstAuthor="Evans" pageId="12" pageNumber="11" pagination="26 - 53" refId="ref27534" refString="Evans, W. H. 1953. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part III. Pyrginae. Section 2. British Museum (Natural History); London. v + 246 p., pl. 26 - 53" type="book chapter" year="1953">Evans (1953)</bibRefCitation>
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. Readily distinguished from its relatives by the wing shape: forewing inner margin concave, hindwing outer margin excavate; forewing with hyaline apical spots;
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<materialsCitation id="3B4F92697E7EFFF2FD8A7755DB89FB0D" ID-GBIF-Occurrence="2569665573" collectionCode="T" pageId="12" pageNumber="11" specimenCode="T843" specimenCount="1">
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uncus long and narrow, pointed at the tip, without side processes, no defined gnathos, valva less than twice longer than wide, penis shorter that in
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<taxonomicName id="4C27E3B77E7EFFF2FB3E77B4DA21FB4D" ID-CoL="92N65" authorityName="Godman and Salvin" authorityYear="1896" box="[1217,1343,1182,1206]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B95344267E7EFFF2FB3E77B4DA21FB4D" box="[1217,1343,1182,1206]" italics="true" pageId="12" pageNumber="11">Staphylus</emphasis>
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</taxonomicName>
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, with a spined cornutus. In DNA, a combination of the following base pairs is diagnostic: aly23605.8.3:C247
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<collectionCode id="ED3600F17E7EFFF2FA767794DA85FB2D" box="[1417,1435,1214,1238]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="11">T</collectionCode>
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, aly5294.23.1:A538C, aly9588.6.1:
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<specimenCode id="DB81304F7E7EFFF2FDA777F4DD89FB0D" box="[600,663,1246,1270]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="11">T843</specimenCode>
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C, aly
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<date id="FF99BEF47E7EFFF2FD2477F4DC58FB0D" box="[731,838,1246,1270]" pageId="12" pageNumber="11" value="1370-07-02">1370.7.2</date>
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:A1760G, aly345.13.8:A62C
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33DCBBF7E7EFFF2FF3F7624DC0DFA5D" pageId="12" pageNumber="11" type="etymology">
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<paragraph id="8B9898347E7EFFF2FF3F7624DA92FADD" blockId="12.[192,1420,1294,1319]" box="[192,1420,1294,1319]" pageId="12" pageNumber="11">
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<emphasis id="B95344267E7EFFF2FF3F7624DE45FADD" bold="true" box="[192,347,1294,1318]" pageId="12" pageNumber="11">Etymology.</emphasis>
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The name, a masculine noun in the nominative singular, echoes the
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<typeStatus id="549C26967E7EFFF2FB597625DBC5FADC" box="[1190,1243,1295,1319]" pageId="12" pageNumber="11">type</typeStatus>
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species name.
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</paragraph>
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<paragraph id="8B9898347E7EFFF2FF3F7614DE5CFA8D" blockId="12.[192,1439,1342,1398]" pageId="12" pageNumber="11">
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<emphasis id="B95344267E7EFFF2FF3F7614DEAFFAAD" bold="true" box="[192,433,1342,1366]" pageId="12" pageNumber="11">Species included.</emphasis>
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The
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<typeStatus id="549C26967E7EFFF2FE0D7615DD39FAAC" box="[498,551,1343,1367]" pageId="12" pageNumber="11">type</typeStatus>
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species,
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<taxonomicName id="4C27E3B77E7EFFF2FD697614DB53FAAD" authority="Hayward, 1933" authorityName="Hayward" authorityYear="1933" box="[662,1101,1342,1366]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciatus">
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<emphasis id="B95344267E7EFFF2FD697614DC96FAAD" box="[662,904,1342,1366]" italics="true" pageId="12" pageNumber="11">Staphylus fasciatus</emphasis>
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Hayward, 1933
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</taxonomicName>
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and
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<taxonomicName id="4C27E3B77E7EFFF2FB727614DA1CFAAD" authorityName="Scudder" authorityYear="1872" box="[1165,1282,1342,1366]" class="Insecta" family="Hesperiidae" genus="Pholisora" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B95344267E7EFFF2FB727614DA1CFAAD" box="[1165,1282,1342,1366]" italics="true" pageId="12" pageNumber="11">Pholisora</emphasis>
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</taxonomicName>
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(?)
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<emphasis id="B95344267E7EFFF2FACE7614DA81FAAD" box="[1329,1439,1342,1366]" italics="true" pageId="12" pageNumber="11">angulata</emphasis>
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Bell, 1937.
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</paragraph>
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<paragraph id="8B9898347E7EFFF2FF3F76A4DC0DFA5D" blockId="12.[192,787,1422,1446]" box="[192,787,1422,1446]" pageId="12" pageNumber="11">
|
|
<emphasis id="B95344267E7EFFF2FF3F76A4DE62FA5D" bold="true" box="[192,380,1422,1446]" pageId="12" pageNumber="11">Parent taxon.</emphasis>
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|
Tribe
|
|
<taxonomicName id="4C27E3B77E7EFFF2FE3376A4DC11FA5D" authority="Verity, 1940" authorityName="Verity" authorityYear="1940" box="[460,783,1422,1446]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="tribe" tribe="Carcharodini">Carcharodini Verity, 1940</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C33DCBBF7E7EFFF2FF3F7694DCDBFA0D" pageId="12" pageNumber="11" type="discussion">
|
|
<paragraph id="8B9898347E7EFFF2FF3F7694DCDBFA0D" blockId="12.[192,1439,1470,1526]" pageId="12" pageNumber="11">
|
|
<emphasis id="B95344267E7EFFF2FF3F7694DE46FA2D" bold="true" box="[192,344,1470,1494]" pageId="12" pageNumber="11">Comments.</emphasis>
|
|
Steinhauser doubted the placement of some of these species in
|
|
<taxonomicName id="4C27E3B77E7EFFF2FB957694DBF8FA2D" authorityName="Godman and Salvin" authorityYear="1896" box="[1130,1254,1470,1494]" class="Insecta" family="Hesperiidae" genus="Staphylus" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B95344267E7EFFF2FB957694DBF8FA2D" box="[1130,1254,1470,1494]" italics="true" pageId="12" pageNumber="11">Staphylus</emphasis>
|
|
</taxonomicName>
|
|
on the basis of morphological analysis (
|
|
<bibRefCitation id="EFB6E5C57E7EFFF2FE1776F4DDDDFA0D" author="Steinhauser" box="[488,707,1502,1526]" firstAuthor="Steinhauser" pageId="12" pageNumber="11" pagination="1 - 70" refId="ref28360" refString="Steinhauser, S. R. 1989. Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part I. Pyrginae. Bulletin of the Allyn Museum 127: 1 - 70." type="journal article" year="1989">Steinhauser 1989</bibRefCitation>
|
|
), and he was correct.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |