Fifty new genera of Hesperiidae (Lepidoptera) Author Cong, Qian Author Zhang, Jing Author Shen, Jinhui Author Grishin, Nick V. text Insecta Mundi 2019 2019-10-11 731 731 1 56 journal article 10.5281/zenodo.3677235 1ee19d5d-27cf-4c93-989c-3fe3eb467203 1942-1354 3677235 BA35690A-FC73-4E5A-A805-FE9550275FEC Incisus Grishin , new genus http://zoobank.org/ 2ED74AC8-4CCF-4796-952B-0292620F0855 Type species. Antigonus incisus Mabille, 1878 . Definition. A genus without close relatives. Formerly placed in Staphylus Godman and Salvin, [1896] , incisus is not monophyletic with the Staphylus type species Helias ascalaphus Staudinger, 1876 , nor with Scantilla opites Godman and Salvin, [1896] (a junior subjective synonym of Tagiades vincula Plötz, 1886 ) the type species of Scantilla Godman and Salvin, [1896] , a genus-group name treated as a subjective synonym of Staphylus ( Fig. 4 ). Keys to E.32.35b in Evans (1953) . Readily distinguished from its relatives by the wing shape: forewing inner margin concave, hindwing outer margin excavate; forewing with hyaline apical spots; uncus long and narrow, pointed at the tip, without side processes, no defined gnathos, valva less than twice longer than wide, penis shorter that in Staphylus , with a spined cornutus. In DNA, a combination of the following base pairs is diagnostic: aly23605.8.3:C247 T , aly5294.23.1:A538C, aly9588.6.1: T843 C, aly 1370.7.2 :A1760G, aly345.13.8:A62C . Etymology. The name, a masculine noun in the nominative singular, echoes the type species name. Species included. The type species, Staphylus fasciatus Hayward, 1933 and Pholisora (?) angulata Bell, 1937. Parent taxon. Tribe Carcharodini Verity, 1940 . Comments. Steinhauser doubted the placement of some of these species in Staphylus on the basis of morphological analysis ( Steinhauser 1989 ), and he was correct.