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455 lines
57 KiB
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<document ID-CLB-Dataset="4348" ID-DOI="http://dx.doi.org/10.3897/zookeys.1059.68385" ID-GBIF-Dataset="a0c40684-8d8e-4aa2-ab8e-461105f5ea96" ID-PMC="PMC8426314" ID-Pensoft-Pub="1313-2970-1059-1" ID-Pensoft-UUID="ABC7C4F5491951E59A72F1FFC1BC0A5F" ID-PubMed="34566444" ID-ZooBank="650D7F4C990E4CBF88E3F0CD38EEC377" ModsDocID="1313-2970-1059-1" checkinTime="1630544413484" checkinUser="pensoft" docAuthor="Chen, Zhe-Yu, Lyu, Zhi-Tong & Wu, Min" docDate="2021" docId="B0722042814951EBA94DD218446E36F0" docLanguage="en" docName="ZooKeys 1059: 1-21" docOrigin="ZooKeys 1059" docPubDate="2021-09-01" docSource="http://dx.doi.org/10.3897/zookeys.1059.68385" docTitle="Stegodera angusticollis" docType="treatment" docVersion="6" id="ABC7C4F5491951E59A72F1FFC1BC0A5F" lastPageNumber="1" masterDocId="ABC7C4F5491951E59A72F1FFC1BC0A5F" masterDocTitle="Systematic revision of Stegodera Martens, 1876 (Gastropoda, Stylommatophora, Camaenidae), with description of a new genus" masterLastPageNumber="21" masterPageNumber="1" pageNumber="1" updateTime="1732803461375" updateUser="ExternalLinkService">
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<mods:title id="D003BD31F3503067D72A0A0C4B46DCD9">Systematic revision of Stegodera Martens, 1876 (Gastropoda, Stylommatophora, Camaenidae), with description of a new genus</mods:title>
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<mods:namePart id="83E10C40A4A4D3136B0B829036E9E773">Chen, Zhe-Yu</mods:namePart>
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<mods:affiliation id="D002D21CA1C60E4FD8E2CA5C1DD07892">College of food science and engineering, Wuhan Polytechnic University, Wuhan 430023, China & School of Life Sciences, Nanjing University, Nanjing 210023, China</mods:affiliation>
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<mods:namePart id="0B4F6D2EC7766479DBAC07C783427B92">Lyu, Zhi-Tong</mods:namePart>
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<mods:affiliation id="5C6021F2DDECD8CB34E81157FB6D1B7D">The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
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<mods:namePart id="C6CA1C25838D3B5C188D92DAF124A148">Wu, Min</mods:namePart>
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<mods:affiliation id="6885817A829D7D02DE9A9B690FC6564B">School of Life Sciences, Nanjing University, Nanjing 210023, China</mods:affiliation>
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<mods:date id="9F6F6E7EF9E707D1EC7224358011356E">2021</mods:date>
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<mods:number id="03D27DD0AC2062DFF6CD9E143DFF4CA4">1059</mods:number>
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<treatment id="B0722042814951EBA94DD218446E36F0" ID-GBIF-Taxon="185459886" LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0" httpUri="http://treatment.plazi.org/id/B0722042814951EBA94DD218446E36F0" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Camaenidae" scope_order="Stylommatophora">
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<subSubSection id="8EC4B9B8F8076AB3FEF6C19DDC5F6BC7" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="E329B494CF839DD75CB3888803D35920" pageId="0" pageNumber="1">
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<taxonomicName id="9F1428C5F6A376CBC274E6D50847FE50" ID-CoL="7T7K6" LSID="B0722042-8149-51EB-A94D-D218446E36F0" authority="(Martens, 1875)" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">Stegodera angusticollis (Martens, 1875)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="369942AA8E982EC6F55EC028783B717D" pageId="0" pageNumber="1" type="description">
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<paragraph id="667917CE5C9534AE2E101DCE575A5BEB" pageId="0" pageNumber="1">
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<figureCitation id="4D68045903D2D3BD658AB483873A2DCF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Bayesian Inferences of camaenids (representatives of genera in Table 1) based on partial mitochondrial 16 S and CO 1 sequences. Numbers near nodes indicate Bayesian posterior probabilities. The squares on the right of each taxon name indicate its character states. Abbreviations: PS: penial sheath; EpP: epiphallic papilla; PC: penial caecum; Fl: flagellum; DS: dart sac apparatus A phylogram without Australian camaenids B the results of ancestral states (tagged on branch in red, blue, or black) reconstruction mapping on the phylogram with some Australian camaenid genera added. Orange part: the topologies supported by the results respectively by using both Bayesian Inference and Maximum Likelihood methods. Character states are from Schileyko (2003) and Wu (2019). Scale bars for substitutions per site." figureDoi="10.3897/zookeys.1059.68385.figure1" httpUri="https://binary.pensoft.net/fig/584114" pageId="0" pageNumber="1">Figures 1</figureCitation>
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<figureCitation id="CEFCC38BF1F3D5297FFDF61FF8A4E1BB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Stegodera and Pseudostegodera qiului gen. et sp. nov. Triangle: new locality of Stegodera angusticollis (Martens, 1875); dashed range: area of Stegodera angusticollis; circle: type locality of Pseudostegodera qiului gen. et sp. nov." figureDoi="10.3897/zookeys.1059.68385.figure2" httpUri="https://binary.pensoft.net/fig/584115" pageId="0" pageNumber="1">, 2</figureCitation>
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<figureCitation id="987732E731D3E5F2B4D844119CA97797" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells A Stegodera angusticollis Martens, 1876, HBUMM 08435 B-D Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 C p rotoconch, magnified D shell surface, magnified. Scale bar: 5 mm (A, B)." figureDoi="10.3897/zookeys.1059.68385.figure3" httpUri="https://binary.pensoft.net/fig/584116" pageId="0" pageNumber="1">, 3A</figureCitation>
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<figureCitation id="7443B688BE555CCB96BA12805561726A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">, 4A-D</figureCitation>
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<figureCitation id="B8053666002206F712E3B16B0D223A5A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">, 5A-C</figureCitation>
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<figureCitation id="F05873CBB5863641DE5B78F7F2B3B83B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">, 6A, B</figureCitation>
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<figureCitation id="9233790EF62254DF6C088C1884482DB3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitat and living specimens of Stegodera angusticollis (Martens, 1875) A habitat (photograph Kai-Chen Ouyang) B the living specimen of HBUMM 08435 C a living specimen from Chibi City (photograph Lu Qiu)." figureDoi="10.3897/zookeys.1059.68385.figure8" httpUri="https://binary.pensoft.net/fig/584121" pageId="0" pageNumber="1">, 8A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0421A80E565419667DFF43AF7A59BAA0" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="B3A9412AFAA3EBF9AA8ABD75F38BABA4" pageId="0" pageNumber="1">
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<taxonomicName id="558EF9A77F802D57FBB75B525B6680FE" authorityName="Martens" authorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">Helix angusticollis</taxonomicName>
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Martens, 1875a: 2.
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<bibRefCitation id="CD89ACA2DACCD391DFF782BF8AFA7915" author="Martens, E von" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="1" pagination="185 - 188" refId="B22" refString="Martens, E von, 1875b. Binnen-Mollusken aus dem mittlern China. Malakozoologische Blaetter 22: 185 - 188" title="Binnen-Mollusken aus dem mittlern China." volume="22" year="1875 b">Martens 1875b</bibRefCitation>
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: 185; Pfeiffer 1875: 449, 1876: 149, pl. 134, figs 7-10;
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<bibRefCitation id="02B953905F33FC00DB8098076D5E7492" author="Gredler, V" journalOrPublisher="Malakozoologische Blaetter, Neue Folge" pageId="0" pageNumber="1" pagination="165 - 187" refId="B10" refString="Gredler, V, 1882. Uebersicht der Binnenschnecken von China. Malakozoologische Blaetter, Neue Folge 5: 165 - 187" title="Uebersicht der Binnenschnecken von China." volume="5" year="1882">Gredler 1882</bibRefCitation>
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: 175;
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<bibRefCitation id="E8D6E5BE7A60952397AB498AFAD8C3B4" DOI="https://doi.org/10.5962/bhl.title.50365" author="Heude, PM" journalOrPublisher="Memoires concernant l'histoire naturelle de l'Empire chinois" pageId="0" pageNumber="1" pagination="1 - 84" refId="B13" refString="Heude, PM, 1882. Notes sur les mollusques terrestres de la vallee du Fleuve Bleu. Memoires concernant l'histoire naturelle de l'Empire chinois 1: 1 - 84, DOI: https://doi.org/10.5962/bhl.title.50365" title="Notes sur les mollusques terrestres de la vallee du Fleuve Bleu." url="https://doi.org/10.5962/bhl.title.50365" volume="1" year="1882">Heude 1882</bibRefCitation>
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: 36, pl. 15, fig. 8;
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<bibRefCitation id="9DFED818614EE4BB40D5FA2ED513D829" author="Moellendorff, OF" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="307 - 391" refId="B23" refString="Moellendorff, OF, 1884. Materialien zur Fauna von China. Jahrbuecher der Deutschen Malakozoologischen Gesellschaft 11: 307 - 391" title="Materialien zur Fauna von China." volume="11" year="1884">
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<normalizedToken id="F47D79800A2218777E858E7BF818D986" originalValue="Möllendorff">Moellendorff</normalizedToken>
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1884
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</bibRefCitation>
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: 387.
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</paragraph>
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<paragraph id="9DFB5A9CD4768D7A119690EE5DB5A480" pageId="0" pageNumber="1">
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<taxonomicName id="C5E66B8C18D9EE2537756725B9469D80" authorityName="Martens" authorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Stegodera) angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis" subGenus="Stegodera">Helix (Stegodera) angusticollis</taxonomicName>
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. - Martens, 1876 (in
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<bibRefCitation id="45F479731445E7D75AA449CC0DAC38BB" author="Pfeiffer, L" journalOrPublisher="Jahrbuecher der Deutschen Malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B26" refString="Pfeiffer, L, 1870-1876. Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles extramarines nouvelles ou peu connues. Tome IV, 1-8, pls 109-111 [1870], 9-40, pls 112-117 [1871], 41-88, pls 118-125 [1872], 89-112, pls 126-130 [1874], 113-144, pls 131-134 [1875], 145-171, pls 135-137." title="Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles extramarines nouvelles ou peu connues. Tome IV, 1 - 8, pls 109 - 111 [1870], 9 - 40, pls 112 - 117 [1871], 41 - 88, pls 118 - 125 [1872], 89 - 112, pls 126 - 130 [1874], 113 - 144, pls 131 - 134 [1875], 145 - 171, pls 135 - 137." year="1870 - 1876">Pfeiffer 1870-1876</bibRefCitation>
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): 149-150;
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<bibRefCitation id="D9701F63634066501D629B77791A03D8" author="Pilsbry, HA" journalOrPublisher="Academy of Natural Science Philadelphia, PA" pageId="0" pageNumber="1" refId="B27" refString="Pilsbry, HA, 1890. [1890-1891] Manual of conchology, Structural and systematic, with illustrations of the species, Second series, Volume 6 (Helicidae, Vol. IV). Academy of Natural Science Philadelphia, PA" title="[1890 - 1891] Manual of conchology, Structural and systematic, with illustrations of the species, Second series, Volume 6 (Helicidae, Vol. IV)." year="1890">Pilsbry 1890</bibRefCitation>
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: 7, pl. 1, figs 15-17.
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</paragraph>
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<paragraph id="B93C81AF756A917F34D87FA91D4AB7A4" pageId="0" pageNumber="1">
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<taxonomicName id="2053B93E714E661B5F7606E3DD680C9C" class="Gastropoda" family="Camaenidae" genus="Plectopylis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectopylis (Stegodera) angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis" subGenus="Stegodera">Plectopylis (Stegodera) angusticollis</taxonomicName>
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. - Pilsbry 1894: 147.
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</paragraph>
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<paragraph id="5935EC60851C9AD0A730DA13B8DE2080" pageId="0" pageNumber="1">
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<taxonomicName id="DC962BDB80E0088B3A8B35B8657FE351" class="Gastropoda" family="Camaenidae" genus="Planispira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Planispira (Stegodera) angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis" subGenus="Stegodera">Planispira (Stegodera) angusticollis</taxonomicName>
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. -
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<bibRefCitation id="60F43F2DE5F3627755047F46CFCDA249" author="Thiele, J" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" refId="B40" refString="Thiele, J, 1931. Handbuch der systematischen Weichtierkunde. Vol. 1, Teil 2: 377-778. Gustav Fischer, Jena." title="Handbuch der systematischen Weichtierkunde. Vol. 1, Teil 2: 377 - 778. Gustav Fischer, Jena." year="1931">Thiele 1931</bibRefCitation>
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: 681.
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</paragraph>
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<paragraph id="489F95F5ACD7FADF8A120D28AAAA36EC" pageId="0" pageNumber="1">
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<taxonomicName id="11821BFBDB28D88AA6F8AE5FB9B4AA70" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">Stegodera angusticollis</taxonomicName>
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. -
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<bibRefCitation id="9ADD52B6B3CFBC0813C1755B41F8DAF2" author="Gude, GK" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="1 - 11" refId="B11" refString="Gude, GK, 1902. A classified list of the helicoid land shells of Asia. Proceedings of the Malacological Society of London 9 (1): 1 - 11" title="A classified list of the helicoid land shells of Asia." volume="9" year="1902">Gude 1902</bibRefCitation>
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: 4; Kobelt 1905: 78;
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<bibRefCitation id="CF41CCDF986D0C2C83F550C839013E8C" author="Pilsbry, HA" journalOrPublisher="The Nautilus" pageId="0" pageNumber="1" pagination="63 - 67" refId="B29" refString="Pilsbry, HA, 1905. Notes on Moellendorffia and Stegodera. The Nautilus 19: 63 - 67" title="Notes on Moellendorffia and Stegodera." volume="19" year="1905">Pilsbry 1905</bibRefCitation>
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: 64, 66, pl. 2, figs 1-3;
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<bibRefCitation id="B8539D10F0442E56C92750C87E80C433" author="Gude, GK" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="52 - 73" refId="B12" refString="Gude, GK, 1920. The armature of land Mollusca. Proceedings of the Malacological Society of London 14 (1): 52 - 73" title="The armature of land Mollusca." volume="14" year="1920">Gude 1920</bibRefCitation>
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: 60;
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|
<bibRefCitation id="C694C42F9326B4C022F64E1CE7660E6D" author="Yen, TC" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 234" refId="B50" refString="Yen, TC, 1939. Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 444: 1 - 234" title="Die chinesischen Land- und Suesswasser-Gastropoden des Natur-Museums Senckenberg." volume="444" year="1939">Yen 1939</bibRefCitation>
|
|
: 126, pl. 13, fig. 9;
|
|
<bibRefCitation id="A7350A762E41A366B24AAB0C00B35B2C" author="Zilch, A" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="1" refId="B51" refString="Zilch, A, 1960. Teil 2, Euthyneura. In: Wenz W (Ed.) Gastropoda, Handbuch der Palaeozoologie (ed. Schindewolf OH), Band 6, Gastropoda. Berlin: Gebrueder Borntraeger, 835 pp. [(4): 601-835 (1960)] [Abb. 2112-2515, I-XII]" title="Teil 2, Euthyneura. In: Wenz W (Ed.) Gastropoda, Handbuch der Palaeozoologie (ed. Schindewolf OH), Band 6, Gastropoda. Berlin: Gebrueder Borntraeger, 835 pp. [(4): 601 - 835 (1960)] [Abb. 2112 - 2515, I-XII]" year="1960">Zilch 1960</bibRefCitation>
|
|
: fig. 2138;
|
|
<bibRefCitation id="AFF09F624771A3188B49AFAC51BF5129" author="Richardson, CL" journalOrPublisher="Tryonia" pageId="0" pageNumber="1" pagination="1 - 253" refId="B34" refString="Richardson, CL, 1983. Bradybaenidae: Catalog of species. Tryonia 12: 1 - 253" title="Bradybaenidae: Catalog of species." volume="12" year="1983">Richardson 1983</bibRefCitation>
|
|
: 291;
|
|
<bibRefCitation id="EF5DB8B8760830B58FD55311F5BF27CC" author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="1" pagination="1467 - 1626" refId="B36" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonate molluscs, Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae. Ruthenica, Supplement 2 (11): 1467 - 1626" title="Treatise on recent terrestrial pulmonate molluscs, Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." volume="2" year="2003">Schileyko 2003</bibRefCitation>
|
|
: 1512, fig. 1948;
|
|
<bibRefCitation id="C163F4B7DE84EE290A493B1BBF00DDCA" author="Chen, DN" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B4" refString="Chen, DN, Gao, JX, 1987. Economic Fauna of China (Terrestrial Molluscs). Science Press, Beijing" title="Economic Fauna of China (Terrestrial Molluscs)." year="1987">Chen and Gao 1987</bibRefCitation>
|
|
: 107, fig. 137; Qian et al. 2008: 289, fig. 169;
|
|
<bibRefCitation id="59A608927AE9A1F3768395AA16B2CDF3" author="Qian, ZX" journalOrPublisher="Zhejiang People's Fine Arts Publishing House, Hangzhou" pageId="0" pageNumber="1" refId="B30" refString="Qian, ZX, Zhou, WC, 2014. Illustrated Handbook of Common Terrestrial Mollusks in China. Zhejiang People's Fine Arts Publishing House, Hangzhou" title="Illustrated Handbook of Common Terrestrial Mollusks in China." year="2014">Qian and Zhou 2014</bibRefCitation>
|
|
: 120, figure in text.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="501971C17DAB8130E56E2F569460B4BC" pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph id="DE0875DF295A1793ED9DD50DD8532494" pageId="0" pageNumber="1">Museum material examined.</paragraph>
|
|
<paragraph id="244EBA04CB877FB329B19F1F8C7E7A0B" pageId="0" pageNumber="1">
|
|
<materialsCitation id="209B158604D75B01C105F30548B1DD17" accessionNumber="SMF27113, SMF27114, SMF42579, TM108733, TM158903, TM159272" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="v. Richthofen, O. v. Moellendorff, K. Hashagen" country="China" location="Hubei" specimenCount="1" stateProvince="Hubei" typeStatus="syntype">
|
|
• ZMB. Moll. 31044,
|
|
<typeStatus id="943BC763717BFCDA83E6318AABBED9BC">syntype</typeStatus>
|
|
, Poyangsee,
|
|
<collectingCountry id="D447A8BBDA02E14706DD214FEA52D807" name="China">China</collectingCountry>
|
|
, slg.
|
|
<collectorName id="C75F2869F2C34EC4103E0DF2EEC8AC1C">v. Richthofen</collectorName>
|
|
; ZMB. • ZMB. Moll. 3710/1
|
|
<collectingCountry id="56D68F5ABD59298178A1210C88BE1CC6" name="China">China</collectingCountry>
|
|
, slg. Paetel; •
|
|
<accessionNumber id="5B4943FD761E855ED876B7CE6896CC82" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF27113">SMF27113</accessionNumber>
|
|
/3,
|
|
<collectingCountry id="7E2AD7413E5F52BEE265BD6FA20EE5C8" name="China">China</collectingCountry>
|
|
: Prov. Hupei, slg.
|
|
<collectorName id="67C4FF6B9036A8DBD2D74BF727C78EB3">O. v. Moellendorff</collectorName>
|
|
; •
|
|
<accessionNumber id="945DF184AFB3344F52C6559568CAC467" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF27114">SMF27114</accessionNumber>
|
|
/2,
|
|
<collectingCountry id="06340324DD53885B97765DA984F412FC" name="China">China</collectingCountry>
|
|
:
|
|
<normalizedToken id="D56B7916878B4E90C59421984ABCE143" originalValue="Wǔ-chang-fú">Wǔ-chang-fu</normalizedToken>
|
|
, slg.
|
|
<collectorName id="C6D889A40B52DCDBF9BCB82649C7DA1C">K. Hashagen</collectorName>
|
|
(ex. Schmacker); •
|
|
<accessionNumber id="A78FE84ECA5825DA326FC6A184FCE276" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF42579">SMF42579</accessionNumber>
|
|
/3,
|
|
<collectingCountry id="1998A3298525E48C8F32F321ABEC9E64" name="China">China</collectingCountry>
|
|
:
|
|
<normalizedToken id="B0F9473F716AF020F6B3ED3D1AF39DD4" originalValue="Húbei">
|
|
<collectingRegion id="DDD5D638403926644A4ACEFBC79AD366" country="China" name="Hubei">Hubei</collectingRegion>
|
|
</normalizedToken>
|
|
(Hupe), slg. Ehrmann; • IZCAS
|
|
<accessionNumber id="390D28E5ED32EA308D35C165D883F8F4" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TM108733">TM108733</accessionNumber>
|
|
/1, Kieou-Kiang; • IZCAS
|
|
<accessionNumber id="A52B4EC72F381153E5C8505BE42D067B" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TM158903">TM158903</accessionNumber>
|
|
-158920/18, Ou-tchang h[ien], ex.
|
|
<normalizedToken id="9FE0A0E9936EDC8599C79704FC319F0A" originalValue="Musée">Musee</normalizedToken>
|
|
Heude; • IZCAS
|
|
<accessionNumber id="8B3BD4F76AD9889E747C55DC80FC0E2A" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TM159272">TM159272</accessionNumber>
|
|
-159397/126, Kieou-Kiang, ex.
|
|
<normalizedToken id="7DC9CE0D3E72F465426B5A06D35BB24B" originalValue="Musée">Musee</normalizedToken>
|
|
Heude
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="858F1FE3258FA58EDB8E3DAA8FCA9C4F" pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph id="D99564217241FC44699EBF6937C13999" pageId="0" pageNumber="1">New material examined.</paragraph>
|
|
<paragraph id="013E2E9D6394675611AA954F4503BA7B" pageId="0" pageNumber="1">
|
|
<materialsCitation id="FAF9AD11C3F6F2CC3B513AECB16ECE1C" collectingDate="2020-10-15" collectorName="Xiao-Long Wang, Zi-Hao Shen" country="China" county="Yangxin County" elevation="140" latitude="30.00106" location="One" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" specimenCount="1" stateProvince="Hubei Province">
|
|
•
|
|
<location id="DB5BE5FCBEEEBDAAC388CC85D09E8027" LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:DB5BE5FCBEEEBDAAC388CC85D09E8027" country="China" county="Yangxin County" latitude="30.00106" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" name="One" stateProvince="Hubei Province">One</location>
|
|
shell of HBUMM08435 (dissected),
|
|
<location id="0662AF0339E1A57D283FB9415CC034FC" LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:0662AF0339E1A57D283FB9415CC034FC" country="China" county="Yangxin County" latitude="30.00106" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" name="Guanyin Cave" stateProvince="Hubei Province">Guanyin Cave</location>
|
|
[观音洞],
|
|
<location id="2FC890E59434868C23A203998F2BB7C1" LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:2FC890E59434868C23A203998F2BB7C1" country="China" county="Yangxin County" latitude="30.00106" longLatPrecision="1" longitude="115.19012" municipality="Huangshi City" name="Taizi Town" stateProvince="Hubei Province">Taizi Town</location>
|
|
[太子镇],
|
|
<collectingCounty id="D8438FB2526CC968BFF4C15112AF19C8">Yangxin County</collectingCounty>
|
|
[阳新县],
|
|
<collectingMunicipality id="CA21839FE0C9DF1C3D1C7412DCF37C62">Huangshi City</collectingMunicipality>
|
|
[黄石市],
|
|
<collectingRegion id="4C758820A6A996B4916A89A79DDD2F4F" country="China" name="Hubei">Hubei Province</collectingRegion>
|
|
,
|
|
<collectingCountry id="CF71D2515BC98195AECBE598FC9889C2" name="China">China</collectingCountry>
|
|
,
|
|
<geoCoordinate id="DAC4D4F2F3A8D8389BB12414103FF633" degrees="30" direction="north" minutes="0" orientation="latitude" precision="1" seconds="3.816" value="30.00106">30°0'3.816"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="8A2AD808A2A7DC582CE77DDE48D91DB9" degrees="115" direction="east" minutes="11" orientation="longitude" precision="1" seconds="24.428" value="115.19012">115°11'24.428"E</geoCoordinate>
|
|
,
|
|
<elevation id="E5EDE05FCB88CD00F5BAA44B705BA164" metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">
|
|
<quantity id="93C3A7ACAD760168C899119EF0401133" metricMagnitude="2" metricUnit="m" metricValue="1.4" unit="m" value="140.0">140 m</quantity>
|
|
a.s.l.
|
|
</elevation>
|
|
,
|
|
<collectingDate id="7ED340F52776BC5A386C96C8A0CF150A" value="2020-10-15">2020-X-15</collectingDate>
|
|
, leg.
|
|
<collectorName id="67F3239832C09E2C4071886466488FFB">Xiao-Long Wang</collectorName>
|
|
&
|
|
<collectorName id="7763C6C471B8F2C450A8DC8649C2B932">Zi-Hao Shen</collectorName>
|
|
(SDmaj =
|
|
<quantity id="A58E1DB246E757431835E1E6A7E344A7" metricMagnitude="-2" metricUnit="m" metricValue="2.67" unit="mm" value="26.7">26.7 mm</quantity>
|
|
, SDmin =
|
|
<quantity id="EB3C93B105CF6E050BECCFB8938F604A" metricMagnitude="-2" metricUnit="m" metricValue="2.13" unit="mm" value="21.3">21.3 mm</quantity>
|
|
, SH =
|
|
<quantity id="B59D9163FBFD035C2E270124E962EC4A" metricMagnitude="-2" metricUnit="m" metricValue="1.13" unit="mm" value="11.3">11.3 mm</quantity>
|
|
)
|
|
</materialsCitation>
|
|
. •
|
|
<materialsCitation id="F02D5459C9C09E79E898EC32B35F35F5" collectingDate="2020-10-03" collectorName="Di Yu, Kai-Chen Ouyang." country="China" county="Xianning City" latitude="29.67" location="Three" longLatPrecision="20" longitude="113.97639" municipality="Chibi City" specimenCount="1" stateProvince="Hubei Province">
|
|
<location id="B1C6973FC7A51022A15687C0556A357C" LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:B1C6973FC7A51022A15687C0556A357C" country="China" county="Xianning City" latitude="29.67" longLatPrecision="20" longitude="113.97639" municipality="Chibi City" name="Three" stateProvince="Hubei Province">Three</location>
|
|
shells of OYKC, one shell of QL,
|
|
<location id="DEC5B63CA9E0187F0D34A8261F45FB1E" LSID="urn:lsid:plazi:treatment:B0722042814951EBA94DD218446E36F0:DEC5B63CA9E0187F0D34A8261F45FB1E" country="China" county="Xianning City" latitude="29.67" longLatPrecision="20" longitude="113.97639" municipality="Chibi City" name="Lushui Lake" stateProvince="Hubei Province">Lushui Lake</location>
|
|
[陆水湖],
|
|
<collectingMunicipality id="DA711906D10D0375F79139CD9292FA98">Chibi City</collectingMunicipality>
|
|
[赤壁市],
|
|
<collectingCounty id="F578A23B60BFB9D3E15DEB092D0D28A4">Xianning City</collectingCounty>
|
|
[咸宁市],
|
|
<collectingRegion id="386B51AA77D54AC47F5CAB79AFC0FF3B" country="China" name="Hubei">Hubei Province</collectingRegion>
|
|
,
|
|
<collectingCountry id="CEAB8FD4D623AACDFFD90648D2DCCB66" name="China">China</collectingCountry>
|
|
,
|
|
<geoCoordinate id="BBB27D019FA0940B283531A628FD0941" degrees="29" direction="north" minutes="40" orientation="latitude" precision="15" seconds="12" value="29.67">29°40'12"N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="3800E41DFB3A2A095A40B28E7C640D6D" degrees="113" direction="east" minutes="58" orientation="longitude" precision="15" seconds="35" value="113.97639">113°58'35"E</geoCoordinate>
|
|
,
|
|
<collectingDate id="7B71779F549BB4F010AAD5E682C3081A" value="2020-10-03">2020-X-03</collectingDate>
|
|
, leg.
|
|
<collectorName id="EC81F6AEA3081B81768393783D306925">Di Yu</collectorName>
|
|
&
|
|
<collectorName id="A91361EBB0420A654D0D33FD7831855C">Kai-Chen Ouyang.</collectorName>
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="56F36543053CD85C855339342176E4E2" pageId="0" pageNumber="1" type="type locality">
|
|
<paragraph id="4460599ECFE4ADB59AB3E2F8FD0D4081" pageId="0" pageNumber="1">Type locality.</paragraph>
|
|
<paragraph id="DB1DFFF335DE5EAD7185C26BE6E7848F" pageId="0" pageNumber="1">Poyang-See (= Poyang Lake [鄱阳湖], Jiangxi).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BBF2DCE327D58F1CD880A068A7E6B058" pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph id="5FD5A44147B631030CCCF3137F20E46E" pageId="0" pageNumber="1">Measurements of new material.</paragraph>
|
|
<paragraph id="17E4B91C07BF8A4735B7825215BA2BDD" pageId="0" pageNumber="1">
|
|
SDmaj =
|
|
<quantity id="B6354DF7A04E6C299C70DDD0068DFCAD" metricMagnitude="-2" metricUnit="m" metricValue="2.8649999999999998" metricValueMax="3.06" metricValueMin="2.67" unit="mm" value="28.65" valueMax="30.6" valueMin="26.7">26.7-30.6 mm</quantity>
|
|
, SDmin =
|
|
<quantity id="AA87EF5FB4BF1410FCD1DD5E1D03E4BB" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.37" metricValueMin="2.13" unit="mm" value="22.5" valueMax="23.7" valueMin="21.3">21.3-23.7 mm</quantity>
|
|
, SH =
|
|
<quantity id="1CAC793D52D2715C94C1B824566D1ED3" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.22" metricValueMin="1.13" unit="mm" value="11.75" valueMax="12.2" valueMin="11.3">11.3-12.2 mm</quantity>
|
|
(n = 6).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="7D9EBA7925B4CEC52ECE867F87A850BB" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="29403A070FC08F9BB25BB81FF81EFF7D" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="9E5667480511CF90748C08C470C0BFB4" pageId="0" pageNumber="1">Body whorl completely covering partial penultimate whorl. Penial sheath weakly present. Epiphallic papilla absent. Dart sac apparatus and flagellum present. Mucous glands with numerous tubes.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="80F68EDD5057C405C5C7E667DAEAB427" pageId="0" pageNumber="1" type="redescription">
|
|
<paragraph id="503DCAFBCE9D28E2E0E9FBE3BAA2A76F" pageId="0" pageNumber="1">Redescription.</paragraph>
|
|
<paragraph id="14F1273D111F955620715A5EF2F9A57D" pageId="0" pageNumber="1">
|
|
Shell (Fig.
|
|
<figureCitation id="AD68C74FCD266E72F3DED8C3440545B4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Shells A Stegodera angusticollis Martens, 1876, HBUMM 08435 B-D Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 C p rotoconch, magnified D shell surface, magnified. Scale bar: 5 mm (A, B)." figureDoi="10.3897/zookeys.1059.68385.figure3" httpUri="https://binary.pensoft.net/fig/584116" pageId="0" pageNumber="1">3A</figureCitation>
|
|
). Sinistral, large, solid, rather flat, five whorls, in chestnut and darker near aperture. Suture impressed. Protoconch 11/4 whorls. After ~ 31/2 whorls, growth lines broken into regularly arranged tubercles. Body whorl compressed from the last 1/4 to the last 1/8 whorls, completely covering corresponding part of penultimate whorl including the suture. After the last 1/8 whorls, body whorl becoming as broad as normal again. On the above compressed region, one shorter ventral depression and one longer apical depression present. Aperture semilunar, slightly descending. Peristome chestnut, thickened, expanded, and slightly reflexed. Umbilicus moderately broad; approximately 1/5 of shell major diameter. Protoconch visible through umbilicus.
|
|
</paragraph>
|
|
<paragraph id="A21C5526F909EBDE94C23F9F62ADD485" pageId="0" pageNumber="1">
|
|
General anatomy (Fig.
|
|
<figureCitation id="B88C18CFE9CF22462945890582E9BEB8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4A-D</figureCitation>
|
|
). Head wart absent. On internal body wall of head region between ommatophore insertions with tiny pits rather than glands (Fig.
|
|
<figureCitation id="CC44A0FD01202E3538211545ED40FBC4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4C</figureCitation>
|
|
). Each side of mantle edge with a leaf-shaped appendage (Fig.
|
|
<figureCitation id="D039D0D5FF796975105536A4C0A0B7E2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4A, B</figureCitation>
|
|
). Body reddish brown, central dorsa with light longitudinal stripes. Sole dirty white. Jaw arcuate; with 12 more or less projecting ribs (Fig.
|
|
<figureCitation id="3CDA7CBADAD00C390D17DF5851CF29D3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge." figureDoi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1">4D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="996D7D0D0E25BA9D6CBB8E75B220536C" pageId="0" pageNumber="1">
|
|
Genitalia (Figs
|
|
<figureCitation id="41A74316B2A94641C79ED195C7296767" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">5A-C</figureCitation>
|
|
,
|
|
<figureCitation id="0296899A57675B121253D26C0596B210" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6A, B</figureCitation>
|
|
). Penis sheath present but thin and very short. Penis medially slightly thickened, moderately long, ~ 8.8 mm, externally simple. Inside penis with several thin longitudinal pilasters. Epiphallus slightly longer than penis. A very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle (Fig.
|
|
<figureCitation id="AC50AFD578B6E5827C8D7080516C2DCC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6B</figureCitation>
|
|
). Flagellum cylindrical, tapering. Vas deferens thin throughout. Dart sac apparatus distally inserting on vagina. Dart sac small. Dart not observed. Membranous sac surrounding terminal genitalia absent. Accessory sac relatively large. Tubes of mucous glands more than twenty, neatly inserting in a single row on distal and same side of dart sac on accessory sac (Figs
|
|
<figureCitation id="3FEFAAC9ACCFBFBF6C9158C3BE445F94" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">5C</figureCitation>
|
|
,
|
|
<figureCitation id="F3D43E2A17489AAD4D3065CFEA5EB2D3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6B</figureCitation>
|
|
). From proximal to distal accessory sac, mucous gland tubes increased gradually in length (Figs
|
|
<figureCitation id="E2485B58795F26B7E28454EB77944F58" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands." figureDoi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1">5C</figureCitation>
|
|
,
|
|
<figureCitation id="D8DD1A22414B37B65FFE852CB5864F37" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6B</figureCitation>
|
|
). Papilla distally leading to mucous glands on inner wall of accessory sac integrated into one long thick spongious pilaster on the side of mucous gland tube insertion (Fig.
|
|
<figureCitation id="6930BC514F03FE80887D0757E8834DD3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle." figureDoi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1">6A</figureCitation>
|
|
). Bursa copulatrix ball-shaped.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0DF9922DC12679E0B775BE47B9178472" pageId="0" pageNumber="1" type="ecology">
|
|
<paragraph id="F17C637C010BF00FD064B117403D0C04" pageId="0" pageNumber="1">Ecology.</paragraph>
|
|
<paragraph id="A4C135B6711198EAB81CF93BD490670D" pageId="0" pageNumber="1">This species was observed under litter layer, on nearby rocks and crevices.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E141E56194AFA07629A3E84B350B29D7" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="40D50DD545FB507F6EA41A47EE2A7259" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="B22027E2908B7F4DC1C12A67DE7EE47D" pageId="0" pageNumber="1">
|
|
Central China: Hubei, Jiangxi (Fig.
|
|
<figureCitation id="51C39D0D15EE58D3E734A9E490050AA7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Stegodera and Pseudostegodera qiului gen. et sp. nov. Triangle: new locality of Stegodera angusticollis (Martens, 1875); dashed range: area of Stegodera angusticollis; circle: type locality of Pseudostegodera qiului gen. et sp. nov." figureDoi="10.3897/zookeys.1059.68385.figure2" httpUri="https://binary.pensoft.net/fig/584115" pageId="0" pageNumber="1">2</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="D4D9932002FC837726D0DE206E1C32EF" doi="10.3897/zookeys.1059.68385.figure2" httpUri="https://binary.pensoft.net/fig/584115" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
|
|
<paragraph id="3B07CA64742D36C6D69AC949B3DE4DBB" pageId="0" pageNumber="1">
|
|
<emphasis id="1A48308F609F92794B3D902947E1312D" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
|
|
Distribution of
|
|
<taxonomicName id="211EFCA63AF4F1EA706EBD5C9AA10C78" authorityName="Martens" authorityYear="1876" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis id="5FCD61FFB3A5F15997EF025A8DF93AE7" italics="true" pageId="0" pageNumber="1">Stegodera</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="649169C0094084142D12DD62A3AAF05D" authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
|
|
<emphasis id="E18F066AD1DF9BB531CD2DE21ADDBF73" italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov. Triangle: new locality of
|
|
<taxonomicName id="5C2A4DD57785E4E33A443BEAF111047A" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
|
|
<emphasis id="277916F67EEB6C8AD541F0A0622AA783" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
|
|
</taxonomicName>
|
|
(Martens, 1875); dashed range: area of
|
|
<taxonomicName id="F2C14210886BB16D85CD50A92D78F7B3" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
|
|
<emphasis id="F2D7FD0C4F9C4E3303C0415D69125F57" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
|
|
</taxonomicName>
|
|
; circle: type locality of
|
|
<taxonomicName id="472A7F891787F5AD135A60B6A50814AE" authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
|
|
<emphasis id="0A23AE970D3606E2B64B145220C18A98" italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="A4EB0F49F79D4C113645AD54969D0783" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="A68AB49C9B80DD80B7BAF91F675D7069" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="F88DA1B4BCC454427BED9048FBC5A693" pageId="0" pageNumber="1">
|
|
The peristome of
|
|
<taxonomicName id="762A685DDB8321C82BEB992EFB90B8DB" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
|
|
<emphasis id="810454899161279CFD41DF28CF380BA8" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
|
|
</taxonomicName>
|
|
is in chestnut in fresh specimens, but in most museum specimens this color gets faint to white. In
|
|
<taxonomicName id="8FB14549299BA6E415E05BFCBC618CCA" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
|
|
<emphasis id="2EF8D86149CD34B78A4AD9F276245F11" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
|
|
</taxonomicName>
|
|
, the compressed part of body whorl usually completely cover the penultimate whorl, while in
|
|
<taxonomicName id="9E85357B8ECECE6AA9A7547F2B0040D5" authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
|
|
<emphasis id="D1083624156E26610C9922E81334F5C7" italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov. the corresponding part is not so compressed and the compressed part of body whorl can hardly cover the penultimate whorl.
|
|
</paragraph>
|
|
<caption id="0042F3168D6A85C69D2A548138DD62BC" doi="10.3897/zookeys.1059.68385.figure3" httpUri="https://binary.pensoft.net/fig/584116" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
|
<paragraph id="71FCFF91CB7B90CEE58DE353174367FF" pageId="0" pageNumber="1">
|
|
<emphasis id="ACE4FE6D999FC350AC912892912D2E5B" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
|
Shells
|
|
<emphasis id="94891AE2378F30A4975DC83654D3AB2D" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
<taxonomicName id="317D6EA69754C81E593A6E27C2390E6C" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
|
|
<emphasis id="A8A4596989230FE227F2A18A8E8F8FE6" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
|
|
</taxonomicName>
|
|
Martens, 1876, HBUMM08435
|
|
<emphasis id="1CFA36F1A07770340D477A0ECCCAF94B" bold="true" pageId="0" pageNumber="1">B-D</emphasis>
|
|
<taxonomicName id="85C55C36EB7D38FEA0FA2D7FF9BBEC45" authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
|
|
<emphasis id="15283FE1293B1E971513764CEA74E8E3" italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., holotype IZCAS TM206978
|
|
<emphasis id="94EEFB5B66CA0DA1DAD9BBE00966D81F" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
p rotoconch, magnified
|
|
<emphasis id="1CF2C2A2CB67F4293FB3EA9F3FDAA949" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
shell surface, magnified. Scale bar: 5 mm (
|
|
<emphasis id="55D6F555DA5ADBA2270932637F9AB160" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="1A9EED42B7DBEF40F520743CA9A8F804" doi="10.3897/zookeys.1059.68385.figure4" httpUri="https://binary.pensoft.net/fig/584117" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
|
|
<paragraph id="E23B1A76ED928BBF26B8C467F19820EC" pageId="0" pageNumber="1">
|
|
<emphasis id="995FFFC21B7027CCE21469FAC828B76A" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
|
General anatomy
|
|
<emphasis id="78A94EAEDA8541EF5253F3B224F71D44" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
|
|
<taxonomicName id="6286B91AE6A5BA58456C5C8117F04C4A" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
|
|
<emphasis id="A524ECC384797B11AC9785F51F7FF1E5" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
|
|
</taxonomicName>
|
|
(Martens, 1875), HBUMM08435
|
|
<emphasis id="A2F7B5523C32A0E5626E06739223542A" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
lobe (arrowed) on the left side of mantle edge
|
|
<emphasis id="26D55E743EF236F203AF0FEC035BAF62" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
lobe (arrowed) on the right side of mantle edge
|
|
<emphasis id="DD4CBC74FDBC56880C8A7A1088ABB837" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
internal surface of head region
|
|
<emphasis id="FD3164BD4E3FE8CA388D79F509B91455" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
jaw
|
|
<emphasis id="F182DB0B32762BFA1D0F88B70A924DA5" bold="true" pageId="0" pageNumber="1">E-H</emphasis>
|
|
<taxonomicName id="E530063426B1E2DD07106439C57F5424" authorityName="Chen" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Pseudostegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudostegodera qiului" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="qiului">
|
|
<emphasis id="240422A92E521F8E66D19336AEC27E63" italics="true" pageId="0" pageNumber="1">Pseudostegodera qiului</emphasis>
|
|
</taxonomicName>
|
|
gen. et sp. nov., holotype IZCAS TM206978
|
|
<emphasis id="F62EDF97E547AF64B63094ED330E81F7" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
jaw
|
|
<emphasis id="29600DFC770C983260A5D1AC11E36F00" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
internal surface of head region
|
|
<emphasis id="7611E7FB834733E59E92CB6701921BB9" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
|
lobe (arrowed) on the left side of mantle edge
|
|
<emphasis id="3CED45E2B954C895D85FA6CFBC7DCBA6" bold="true" pageId="0" pageNumber="1">H</emphasis>
|
|
lobe (arrowed) on the right side of mantle edge.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="4F7A3754797E1DB54420E2C95287943D" pageId="0" pageNumber="1">
|
|
Some authors (
|
|
<bibRefCitation id="92A2274C07E777C391FD17EB0F2B3E4F" author="Chen, DN" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="1" refId="B4" refString="Chen, DN, Gao, JX, 1987. Economic Fauna of China (Terrestrial Molluscs). Science Press, Beijing" title="Economic Fauna of China (Terrestrial Molluscs)." year="1987">Chen and Gao 1987</bibRefCitation>
|
|
; Qian et al. 2008;
|
|
<bibRefCitation id="E845635D56D449585ACD61BB14C154DC" author="Qian, ZX" journalOrPublisher="Zhejiang People's Fine Arts Publishing House, Hangzhou" pageId="0" pageNumber="1" refId="B30" refString="Qian, ZX, Zhou, WC, 2014. Illustrated Handbook of Common Terrestrial Mollusks in China. Zhejiang People's Fine Arts Publishing House, Hangzhou" title="Illustrated Handbook of Common Terrestrial Mollusks in China." year="2014">Qian and Zhou 2014</bibRefCitation>
|
|
) recorded this species in Jiangsu, Zhejiang and Anhui, but voucher specimens from these provinces were neither recorded therein nor being found in the mollusk collection deposited in IZCAS where some of them once worked. Therefore, the accuracy of these records needs to be further confirmed.
|
|
</paragraph>
|
|
<caption id="077A2CF108FB8613D16D940C5D2583BB" doi="10.3897/zookeys.1059.68385.figure5" httpUri="https://binary.pensoft.net/fig/584118" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
|
|
<paragraph id="3FC165841001EC750CE84A8DDDC3D6BA" pageId="0" pageNumber="1">
|
|
<emphasis id="9BE847961AACAD372A8E9FDCC7176939" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
|
|
Genitalia of
|
|
<taxonomicName id="727AA34B8044095DF1715716AD8B5277" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
|
|
<emphasis id="BAA4EB5A68918E23D98A8CE423B62536" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
|
|
</taxonomicName>
|
|
(Martens, 1875), HBUMM08435
|
|
<emphasis id="AF668EBFEA399D4A3C8F6CC7473F8FC9" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
general view
|
|
<emphasis id="3F0D96AF9F225026AF8E971BDDFF247E" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
male part, showing the vas deferens connecting epiphallus (arrowed)
|
|
<emphasis id="F8413A2F95CC958649C5DE8E0923F6E2" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
female part, showing the dart sac, accessory sac, and mucous glands.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="D2055D001A3C5FBB8043B492F3095621" doi="10.3897/zookeys.1059.68385.figure6" httpUri="https://binary.pensoft.net/fig/584119" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
|
|
<paragraph id="3DED034D08D064B46A8BF2101A7636BB" pageId="0" pageNumber="1">
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<emphasis id="8B0CFDE57CEABFCE735C4994EF812A5D" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
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Genitalia of
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<taxonomicName id="EC1CDF05518AEBFC407FCD89C2B6BD5D" baseAuthorityName="Martens" baseAuthorityYear="1875" class="Gastropoda" family="Camaenidae" genus="Stegodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegodera angusticollis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="angusticollis">
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<emphasis id="03B5214743AD09E91C5C791F03D22DC1" italics="true" pageId="0" pageNumber="1">Stegodera angusticollis</emphasis>
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</taxonomicName>
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(Martens, 1875), HBUMM08435
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<emphasis id="20FA39A09B54DE89636DC171DBEA1918" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
exposed dart sac apparatus
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<emphasis id="E772D13A0CCCFDB6BD4017574F589F02" bold="true" pageId="0" pageNumber="1">B</emphasis>
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magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |