<documentID-CLB-Dataset="4348"ID-DOI="http://dx.doi.org/10.3897/zookeys.1059.68385"ID-GBIF-Dataset="a0c40684-8d8e-4aa2-ab8e-461105f5ea96"ID-PMC="PMC8426314"ID-Pensoft-Pub="1313-2970-1059-1"ID-Pensoft-UUID="ABC7C4F5491951E59A72F1FFC1BC0A5F"ID-PubMed="34566444"ID-ZooBank="650D7F4C990E4CBF88E3F0CD38EEC377"ModsDocID="1313-2970-1059-1"checkinTime="1630544413484"checkinUser="pensoft"docAuthor="Chen, Zhe-Yu, Lyu, Zhi-Tong & Wu, Min"docDate="2021"docId="B0722042814951EBA94DD218446E36F0"docLanguage="en"docName="ZooKeys 1059: 1-21"docOrigin="ZooKeys 1059"docPubDate="2021-09-01"docSource="http://dx.doi.org/10.3897/zookeys.1059.68385"docTitle="Stegodera angusticollis"docType="treatment"docVersion="6"id="ABC7C4F5491951E59A72F1FFC1BC0A5F"lastPageNumber="1"masterDocId="ABC7C4F5491951E59A72F1FFC1BC0A5F"masterDocTitle="Systematic revision of Stegodera Martens, 1876 (Gastropoda, Stylommatophora, Camaenidae), with description of a new genus"masterLastPageNumber="21"masterPageNumber="1"pageNumber="1"updateTime="1732803461375"updateUser="ExternalLinkService">
<mods:titleid="D003BD31F3503067D72A0A0C4B46DCD9">Systematic revision of Stegodera Martens, 1876 (Gastropoda, Stylommatophora, Camaenidae), with description of a new genus</mods:title>
<mods:affiliationid="D002D21CA1C60E4FD8E2CA5C1DD07892">College of food science and engineering, Wuhan Polytechnic University, Wuhan 430023, China & School of Life Sciences, Nanjing University, Nanjing 210023, China</mods:affiliation>
<mods:affiliationid="5C6021F2DDECD8CB34E81157FB6D1B7D">The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
<figureCitationid="4D68045903D2D3BD658AB483873A2DCF"captionStart="Figure 1"captionStartId="F1"captionText="Figure 1. Bayesian Inferences of camaenids (representatives of genera in Table 1) based on partial mitochondrial 16 S and CO 1 sequences. Numbers near nodes indicate Bayesian posterior probabilities. The squares on the right of each taxon name indicate its character states. Abbreviations: PS: penial sheath; EpP: epiphallic papilla; PC: penial caecum; Fl: flagellum; DS: dart sac apparatus A phylogram without Australian camaenids B the results of ancestral states (tagged on branch in red, blue, or black) reconstruction mapping on the phylogram with some Australian camaenid genera added. Orange part: the topologies supported by the results respectively by using both Bayesian Inference and Maximum Likelihood methods. Character states are from Schileyko (2003) and Wu (2019). Scale bars for substitutions per site."figureDoi="10.3897/zookeys.1059.68385.figure1"httpUri="https://binary.pensoft.net/fig/584114"pageId="0"pageNumber="1">Figures 1</figureCitation>
<figureCitationid="CEFCC38BF1F3D5297FFDF61FF8A4E1BB"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Distribution of Stegodera and Pseudostegodera qiului gen. et sp. nov. Triangle: new locality of Stegodera angusticollis (Martens, 1875); dashed range: area of Stegodera angusticollis; circle: type locality of Pseudostegodera qiului gen. et sp. nov."figureDoi="10.3897/zookeys.1059.68385.figure2"httpUri="https://binary.pensoft.net/fig/584115"pageId="0"pageNumber="1">, 2</figureCitation>
<figureCitationid="987732E731D3E5F2B4D844119CA97797"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Shells A Stegodera angusticollis Martens, 1876, HBUMM 08435 B-D Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 C p rotoconch, magnified D shell surface, magnified. Scale bar: 5 mm (A, B)."figureDoi="10.3897/zookeys.1059.68385.figure3"httpUri="https://binary.pensoft.net/fig/584116"pageId="0"pageNumber="1">, 3A</figureCitation>
<figureCitationid="7443B688BE555CCB96BA12805561726A"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge."figureDoi="10.3897/zookeys.1059.68385.figure4"httpUri="https://binary.pensoft.net/fig/584117"pageId="0"pageNumber="1">, 4A-D</figureCitation>
<figureCitationid="B8053666002206F712E3B16B0D223A5A"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands."figureDoi="10.3897/zookeys.1059.68385.figure5"httpUri="https://binary.pensoft.net/fig/584118"pageId="0"pageNumber="1">, 5A-C</figureCitation>
<figureCitationid="F05873CBB5863641DE5B78F7F2B3B83B"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle."figureDoi="10.3897/zookeys.1059.68385.figure6"httpUri="https://binary.pensoft.net/fig/584119"pageId="0"pageNumber="1">, 6A, B</figureCitation>
<figureCitationid="9233790EF62254DF6C088C1884482DB3"captionStart="Figure 8"captionStartId="F8"captionText="Figure 8. Habitat and living specimens of Stegodera angusticollis (Martens, 1875) A habitat (photograph Kai-Chen Ouyang) B the living specimen of HBUMM 08435 C a living specimen from Chibi City (photograph Lu Qiu)."figureDoi="10.3897/zookeys.1059.68385.figure8"httpUri="https://binary.pensoft.net/fig/584121"pageId="0"pageNumber="1">, 8A-C</figureCitation>
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<materialsCitationid="209B158604D75B01C105F30548B1DD17"accessionNumber="SMF27113, SMF27114, SMF42579, TM108733, TM158903, TM159272"collectingDate="2021-01-01"collectingDateMax="2021-12-31"collectingDateMin="2021-01-01"collectorName="v. Richthofen, O. v. Moellendorff, K. Hashagen"country="China"location="Hubei"specimenCount="1"stateProvince="Hubei"typeStatus="syntype">
<figureCitationid="AD68C74FCD266E72F3DED8C3440545B4"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Shells A Stegodera angusticollis Martens, 1876, HBUMM 08435 B-D Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 C p rotoconch, magnified D shell surface, magnified. Scale bar: 5 mm (A, B)."figureDoi="10.3897/zookeys.1059.68385.figure3"httpUri="https://binary.pensoft.net/fig/584116"pageId="0"pageNumber="1">3A</figureCitation>
). Sinistral, large, solid, rather flat, five whorls, in chestnut and darker near aperture. Suture impressed. Protoconch 11/4 whorls. After ~ 31/2 whorls, growth lines broken into regularly arranged tubercles. Body whorl compressed from the last 1/4 to the last 1/8 whorls, completely covering corresponding part of penultimate whorl including the suture. After the last 1/8 whorls, body whorl becoming as broad as normal again. On the above compressed region, one shorter ventral depression and one longer apical depression present. Aperture semilunar, slightly descending. Peristome chestnut, thickened, expanded, and slightly reflexed. Umbilicus moderately broad; approximately 1/5 of shell major diameter. Protoconch visible through umbilicus.
<figureCitationid="B88C18CFE9CF22462945890582E9BEB8"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge."figureDoi="10.3897/zookeys.1059.68385.figure4"httpUri="https://binary.pensoft.net/fig/584117"pageId="0"pageNumber="1">4A-D</figureCitation>
<figureCitationid="CC44A0FD01202E3538211545ED40FBC4"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge."figureDoi="10.3897/zookeys.1059.68385.figure4"httpUri="https://binary.pensoft.net/fig/584117"pageId="0"pageNumber="1">4C</figureCitation>
<figureCitationid="D039D0D5FF796975105536A4C0A0B7E2"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge."figureDoi="10.3897/zookeys.1059.68385.figure4"httpUri="https://binary.pensoft.net/fig/584117"pageId="0"pageNumber="1">4A, B</figureCitation>
<figureCitationid="3CDA7CBADAD00C390D17DF5851CF29D3"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. General anatomy A-D Stegodera angusticollis (Martens, 1875), HBUMM 08435 A lobe (arrowed) on the left side of mantle edge B lobe (arrowed) on the right side of mantle edge C internal surface of head region D jaw E-H Pseudostegodera qiului gen. et sp. nov., holotype IZCAS TM 206978 E jaw F internal surface of head region G lobe (arrowed) on the left side of mantle edge H lobe (arrowed) on the right side of mantle edge."figureDoi="10.3897/zookeys.1059.68385.figure4"httpUri="https://binary.pensoft.net/fig/584117"pageId="0"pageNumber="1">4D</figureCitation>
<figureCitationid="41A74316B2A94641C79ED195C7296767"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands."figureDoi="10.3897/zookeys.1059.68385.figure5"httpUri="https://binary.pensoft.net/fig/584118"pageId="0"pageNumber="1">5A-C</figureCitation>
<figureCitationid="0296899A57675B121253D26C0596B210"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle."figureDoi="10.3897/zookeys.1059.68385.figure6"httpUri="https://binary.pensoft.net/fig/584119"pageId="0"pageNumber="1">6A, B</figureCitation>
). Penis sheath present but thin and very short. Penis medially slightly thickened, moderately long, ~ 8.8 mm, externally simple. Inside penis with several thin longitudinal pilasters. Epiphallus slightly longer than penis. A very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle (Fig.
<figureCitationid="AC50AFD578B6E5827C8D7080516C2DCC"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle."figureDoi="10.3897/zookeys.1059.68385.figure6"httpUri="https://binary.pensoft.net/fig/584119"pageId="0"pageNumber="1">6B</figureCitation>
). Flagellum cylindrical, tapering. Vas deferens thin throughout. Dart sac apparatus distally inserting on vagina. Dart sac small. Dart not observed. Membranous sac surrounding terminal genitalia absent. Accessory sac relatively large. Tubes of mucous glands more than twenty, neatly inserting in a single row on distal and same side of dart sac on accessory sac (Figs
<figureCitationid="3FEFAAC9ACCFBFBF6C9158C3BE445F94"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands."figureDoi="10.3897/zookeys.1059.68385.figure5"httpUri="https://binary.pensoft.net/fig/584118"pageId="0"pageNumber="1">5C</figureCitation>
<figureCitationid="F3D43E2A17489AAD4D3065CFEA5EB2D3"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle."figureDoi="10.3897/zookeys.1059.68385.figure6"httpUri="https://binary.pensoft.net/fig/584119"pageId="0"pageNumber="1">6B</figureCitation>
<figureCitationid="E2485B58795F26B7E28454EB77944F58"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A general view B male part, showing the vas deferens connecting epiphallus (arrowed) C female part, showing the dart sac, accessory sac, and mucous glands."figureDoi="10.3897/zookeys.1059.68385.figure5"httpUri="https://binary.pensoft.net/fig/584118"pageId="0"pageNumber="1">5C</figureCitation>
<figureCitationid="D8DD1A22414B37B65FFE852CB5864F37"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle."figureDoi="10.3897/zookeys.1059.68385.figure6"httpUri="https://binary.pensoft.net/fig/584119"pageId="0"pageNumber="1">6B</figureCitation>
). Papilla distally leading to mucous glands on inner wall of accessory sac integrated into one long thick spongious pilaster on the side of mucous gland tube insertion (Fig.
<figureCitationid="6930BC514F03FE80887D0757E8834DD3"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Genitalia of Stegodera angusticollis (Martens, 1875), HBUMM 08435 A exposed dart sac apparatus B magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle."figureDoi="10.3897/zookeys.1059.68385.figure6"httpUri="https://binary.pensoft.net/fig/584119"pageId="0"pageNumber="1">6A</figureCitation>
<paragraphid="A4C135B6711198EAB81CF93BD490670D"pageId="0"pageNumber="1">This species was observed under litter layer, on nearby rocks and crevices.</paragraph>
<figureCitationid="51C39D0D15EE58D3E734A9E490050AA7"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Distribution of Stegodera and Pseudostegodera qiului gen. et sp. nov. Triangle: new locality of Stegodera angusticollis (Martens, 1875); dashed range: area of Stegodera angusticollis; circle: type locality of Pseudostegodera qiului gen. et sp. nov."figureDoi="10.3897/zookeys.1059.68385.figure2"httpUri="https://binary.pensoft.net/fig/584115"pageId="0"pageNumber="1">2</figureCitation>
<bibRefCitationid="92A2274C07E777C391FD17EB0F2B3E4F"author="Chen, DN"journalOrPublisher="Science Press, Beijing"pageId="0"pageNumber="1"refId="B4"refString="Chen, DN, Gao, JX, 1987. Economic Fauna of China (Terrestrial Molluscs). Science Press, Beijing"title="Economic Fauna of China (Terrestrial Molluscs)."year="1987">Chen and Gao 1987</bibRefCitation>
<bibRefCitationid="E845635D56D449585ACD61BB14C154DC"author="Qian, ZX"journalOrPublisher="Zhejiang People's Fine Arts Publishing House, Hangzhou"pageId="0"pageNumber="1"refId="B30"refString="Qian, ZX, Zhou, WC, 2014. Illustrated Handbook of Common Terrestrial Mollusks in China. Zhejiang People's Fine Arts Publishing House, Hangzhou"title="Illustrated Handbook of Common Terrestrial Mollusks in China."year="2014">Qian and Zhou 2014</bibRefCitation>
) recorded this species in Jiangsu, Zhejiang and Anhui, but voucher specimens from these provinces were neither recorded therein nor being found in the mollusk collection deposited in IZCAS where some of them once worked. Therefore, the accuracy of these records needs to be further confirmed.
magnified partial penis and partial epiphallus. Arrow showing a very thin membranous sac wrapping distal half of penis with distal end connecting basal penial retractor muscle.
