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<document id="B048C19E16BB9611DE2C486064380CFF" ID-CLB-Dataset="298861" ID-DOI="10.3897/zookeys.1205.117807" ID-GBIF-Dataset="6af25ad8-6ada-4c5f-a06a-fbe27e26526c" ID-ZooBank="31CA30DA-E778-4046-B479-5F1705D4FC9A" ID-publisher-id="117807" URI-arpha="A212B4BB-598D-5DE6-8284-E0206AEF6042" URI-zoobank="http://zoobank.org/31CA30DA-E778-4046-B479-5F1705D4FC9A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1719235456447" checkinUser="pensoft" docAuthor="Lv, Hanhuiying, Xu, Yuanyuan, Liu, Shulin, Li, Linghui, Lin, Kejian &amp; Zhang, Xiao" docDate="2024" docId="4D62C3A043335E45BB6CE25A7BA17E3A" docLanguage="en" docName="ZooKeys 1205: 191-204" docOrigin="ZooKeys 1205" docSource="https://zookeys.pensoft.net/article/117807/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Toxorhina (Ceratocheilus) pianmica Xu, Lv &amp; Zhang 2024, sp. nov." docType="treatment" docUuid="C4EF4120-18B3-44FF-8398-0745D97927D3" docUuidSource="ZooBank" docVersion="5" dtd-version="3.0" lastPageNumber="204" masterDocId="A212B4BB598D5DE68284E0206AEF6042" masterDocTitle="An updated review of the genus Toxorhina Loew, 1850 (Diptera, Limoniidae) from Yunnan, China with a description of a new species" masterLastPageNumber="204" masterPageNumber="191" pageNumber="191" updateTime="1719327796742" updateUser="admin">
<mods:mods id="967FCCE32619D32B80A16BEA41176F88" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="32B73E3C2B5DF06E4664CC0FB6191DE6">
<mods:title id="53A61B3AA7F41A0B6704E3F0D455EDE5">An updated review of the genus Toxorhina Loew, 1850 (Diptera, Limoniidae) from Yunnan, China with a description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="313ADA25D2553341E531B592A0825C25" type="personal">
<mods:role id="5196967AEE3918A015942527A29EF021">
<mods:roleTerm id="F921DD01A25146941039151902670362">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CAA2B69266BACB4DCE8710777E0DEA92">Lv, Hanhuiying</mods:namePart>
<mods:nameIdentifier id="289AAFC68DB7745928339CB612C3DC69" type="ORCID">https://orcid.org/0009-0007-2912-5893</mods:nameIdentifier>
<mods:affiliation id="1D455B315670199719A8BEE6C10AB7B8">Key Laboratory of Biohazard Monitoring and Green Prevention and Control in Artificial Grassland, Ministry of Agriculture and Rural Affairs, Institute of Grassland Research, Chinese Academy of Agricultural Sciences, Hohhot 010010, China &amp; Shandong Engineering Research Center for Environment-Friendly Agricultural Pest Management, College of Plant Health and Medicine, Qingdao Agricultural University, Qingdao 266109, China</mods:affiliation>
</mods:name>
<mods:name id="0B0232D1EFB4C051BEC5DBCF70DBF6F7" type="personal">
<mods:role id="2B0CCCE04F07129A34F9DD9F9C1BB52D">
<mods:roleTerm id="ECB60F344678504F281D75040FE6A512">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B996BCA5206704F258D5EF5C7A740AF">Xu, Yuanyuan</mods:namePart>
<mods:nameIdentifier id="4F73383B2B91A1C8F15AD3819F1A44FC" type="ORCID">https://orcid.org/0009-0005-2118-0391</mods:nameIdentifier>
<mods:affiliation id="0C96E132529DBEAE01B0A54F5A8FC231">Shandong Engineering Research Center for Environment-Friendly Agricultural Pest Management, College of Plant Health and Medicine, Qingdao Agricultural University, Qingdao 266109, China &amp; Guangxi Key Laboratory of Agric-Environment and Agric-Products Safety, National Demonstration Center for Experimental Plant Science Education, Agricultural College, Guangxi University, Nanning 530004, China</mods:affiliation>
</mods:name>
<mods:name id="282E121C5B6989BCF4AE24CB4F21A6B7" type="personal">
<mods:role id="77B8E8AB0716C3984D528D0EC6B5B011">
<mods:roleTerm id="8F79F2EADCEC3BEF04499C0415EB1A19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57C405F93AEE04A481ADFA6A3C603A58">Liu, Shulin</mods:namePart>
<mods:nameIdentifier id="8D68F2AD9A51D678650B4397D1E5727B" type="ORCID">https://orcid.org/0009-0008-6858-6972</mods:nameIdentifier>
<mods:affiliation id="5593BCA8128C87B0193CD039F5898A0C">Shandong Engineering Research Center for Environment-Friendly Agricultural Pest Management, College of Plant Health and Medicine, Qingdao Agricultural University, Qingdao 266109, China</mods:affiliation>
</mods:name>
<mods:name id="E9FA3FB4FC042EA591F1E265ED9B8CF9" type="personal">
<mods:role id="86201E5CF1A02B52B8EC74B72A09C19E">
<mods:roleTerm id="2B056C2256B7DC320152B95D64B52646">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8297C4CDF52734194F954938ADAD2B67">Li, Linghui</mods:namePart>
<mods:nameIdentifier id="28AB69A03D95BFD1E4D41E8C4B5E0D10" type="ORCID">https://orcid.org/0009-0003-7432-9482</mods:nameIdentifier>
<mods:affiliation id="E3688C7E320FEBA044B7B9D0792C4338">Shandong Engineering Research Center for Environment-Friendly Agricultural Pest Management, College of Plant Health and Medicine, Qingdao Agricultural University, Qingdao 266109, China</mods:affiliation>
</mods:name>
<mods:name id="E14E2850099D820C1D2C3AA2390CFE00" type="personal">
<mods:role id="E75FA52629806E780F7CC9F39F048802">
<mods:roleTerm id="76070DEF271AC777DAA56DE737B0B648">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FC1CCC823CD05E5CEF943066D37428D">Lin, Kejian</mods:namePart>
<mods:nameIdentifier id="78A8BA2BFA461734A35F21AA184209C9" type="ORCID">0000-0002-8531-8506</mods:nameIdentifier>
<mods:affiliation id="BA69CA8A6305AA6DBCBBE9E1BCEDAE4A">Key Laboratory of Biohazard Monitoring and Green Prevention and Control in Artificial Grassland, Ministry of Agriculture and Rural Affairs, Institute of Grassland Research, Chinese Academy of Agricultural Sciences, Hohhot 010010, China</mods:affiliation>
</mods:name>
<mods:name id="F43CCD2B6C0EB0D3B85F0B194DBA09C6" type="personal">
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<mods:namePart id="73C3196875FFBE6A3C95977064638FB7">Zhang, Xiao</mods:namePart>
<mods:nameIdentifier id="B51F492B143010786F55E29CC3B28B9E" type="ORCID">0000-0002-8054-0968</mods:nameIdentifier>
<mods:affiliation id="D0745794D4A361EC7096F82B497049DD">Shandong Engineering Research Center for Environment-Friendly Agricultural Pest Management, College of Plant Health and Medicine, Qingdao Agricultural University, Qingdao 266109, China</mods:affiliation>
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<mods:typeOfResource id="E263F297ECBA0EFD29DC68B574316B88">text</mods:typeOfResource>
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<mods:title id="6CE289E22C876FA816C88499811A706E">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="661FD190F11C199FC9617266A2CB74E8">
<mods:date id="3EB2EEEC2CCBE68D747FDB8847BB4EEF">2024</mods:date>
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<mods:number id="33F10F84FB580D04B6C2CCF44CE796DA">2024-06-24</mods:number>
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<mods:detail id="C186C718C8F383C77FD87286D4CA88A1" type="volume">
<mods:number id="A7852F738386376B0FE0F484C2D45126">1205</mods:number>
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<mods:start id="EFBA689A6E5B1EC0CE5DDCE5A9B0272D">191</mods:start>
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<mods:identifier id="8D0841DE06FA293261BD5BADD339BF89" type="CLB-Dataset">298861</mods:identifier>
<mods:identifier id="20ED4445DB196A2580FDB84872E6F281" type="DOI">10.3897/zookeys.1205.117807</mods:identifier>
<mods:identifier id="35A4A5ED265225100EACD1653479B845" type="GBIF-Dataset">6af25ad8-6ada-4c5f-a06a-fbe27e26526c</mods:identifier>
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</mods:mods>
<treatment id="4D62C3A043335E45BB6CE25A7BA17E3A" ID-DOI="http://doi.org/10.5281/zenodo.12517552" ID-Zenodo-Dep="12517552" ID-arpha="4D62C3A0-4333-5E45-BB6C-E25A7BA17E3A" ID-zoobank="https://zoobank.org/C4EF4120-18B3-44FF-8398-0745D97927D3" LSID="urn:lsid:zoobank.org:act:C4EF4120-18B3-44FF-8398-0745D97927D3" LSID-ZBK="urn:lsid:zoobank.org:act:C4EF4120-18B3-44FF-8398-0745D97927D3" httpUri="http://treatment.plazi.org/id/4D62C3A043335E45BB6CE25A7BA17E3A">
<subSubSection id="7CA212C5E93ECE219EF6619E5E8252BC" type="nomenclature">
<paragraph id="5B1EB5A0EACE9567FFB277B111865D8B">
<taxonomicName id="072EB216E39357386F9377A2592264D2" authority="Xu, Lv &amp; Zhang" authorityName="Xu, Lv &amp; Zhang" authorityYear="2024" class="Insecta" family="Limoniidae" genus="Toxorhina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pianmica" status="sp. nov." subGenus="Ceratocheilus">
<emphasis id="16725B5ABB677B54E394B277B48336E2" italics="true">Toxorhina</emphasis>
(
<emphasis id="CDE117FC5380DAA4D9A3D18E66A34BD3" italics="true">Ceratocheilus</emphasis>
)
<emphasis id="0D6CEE500A90AAE6CCD46EEF3901E2BD" italics="true">pianmica</emphasis>
Xu, Lv &amp; Zhang
</taxonomicName>
<taxonomicNameLabel id="888ACB37F70606CC6C562875D12D71F2" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3D9DA850D28D99E37DF7A050737413E1" type="description">
<paragraph id="6BAAF04C2AFF15F660B70F5108CF00A0">
<figureCitation id="830C94BFA625CFB602C016E11034FF3A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">Figs 4</figureCitation>
,
<figureCitation id="9D6C0827798D43548FD24D195041089D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECUAE" type="type material">
<paragraph id="83E459DA7FAF63ECF81F27B587C1C6F9">
<heading id="7E165D4528DE196D26855E3C7F0D304E" reason="title">Type material.</heading>
</paragraph>
<paragraph id="0C23A4E91ECBE2062CCD0F3834677E38">
<materialsCitation id="12083B3D8E6685BE430F3DF8382ADFEC" collectingDate="2013-07-06" collectionCode="CAU" collectorName="Xuankun Li" country="China" county="Lushui County" elevation="2123" latitude="26.0142" location="Pianma Town" longLatPrecision="6" longitude="98.6272" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="C13B4888C3406E1FB88F15C587D30D77" bold="true">
<emphasis id="330D42AC569B26445688BC4CC9701AD0" bold="true" italics="true">
<typeStatus id="F45FA807E28C562C56A70A5A64ED1460" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="5895672FC16BA70692C79200AADA71F2" name="China">China</collectingCountry>
•
<specimenCount id="DD23981B1F4A1EAA00015618C9608B7F" count="1" type="male">♂</specimenCount>
;
<collectingRegion id="0F17717E640919D75C825D58036E6160" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCounty id="C9C6D79F361D08C7E0C171343FEF7DBB">Lushui County</collectingCounty>
,
<location id="50E4E59699CCFA47B178D05FC6DACDFA" LSID="urn:lsid:plazi:treatment:4D62C3A043335E45BB6CE25A7BA17E3A:50E4E59699CCFA47B178D05FC6DACDFA" country="China" county="Lushui County" latitude="26.0142" longLatPrecision="6" longitude="98.6272" name="Pianma Town" stateProvince="Yunnan">Pianma Town</location>
;
<elevation id="66EC9EF3B21BF0544EF75E4C1945E807" metricMagnitude="3" metricUnit="m" metricValue="2.123" unit="m" value="2123.0">
<quantity id="329410510FC5800DB6576CDA6A3F612B" metricMagnitude="3" metricUnit="m" metricValue="2.123" unit="m" value="2123.0">2123 m</quantity>
a. s. l.
</elevation>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQUAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[98.627200,26.014200]}">
<geoCoordinate id="C6E953D57FE4B02FAC2B956B606B754D" degrees="26.0142" direction="north" orientation="latitude" precision="5" value="26.0142">26.0142 ° N</geoCoordinate>
,
<geoCoordinate id="859EFE6A04485ABBB8D6C90B7821954E" degrees="98.6272" direction="east" orientation="longitude" precision="5" value="98.6272">98.6272 ° E</geoCoordinate>
</named-content>
;
<date id="0757F3D6D668222214537B374A5CF309" value="2013-07-06">
<collectingDate id="D079584A0EADB58BB0AF8498DE02226E" value="2013-07-06">6 July 2013</collectingDate>
</date>
;
<collectorName id="48C7F9C637FF8D5BAEF00E987BB2C620">Xuankun Li</collectorName>
leg.;
<collectingMethod id="7A78C045B47650167DB12475EBB29EE2">light trap</collectingMethod>
;
<collectionCode id="83D9BEE2B3A58CA57CFE0C1284DCEAEE" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" type="">CAU</collectionCode>
</materialsCitation>
.
<materialsCitation id="F57472E01DD2279DABB2910C29FDA548" collectingDate="2013-07-06" collectionCode="CAU" collectorName="Xuankun Li" country="China" county="Lushui County" elevation="2123" latitude="26.0142" location="Pianma Town" longLatPrecision="6" longitude="98.6272" specimenCount="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="372E7D4E00F04A22FE4BDD622513C61E" bold="true" italics="true">
<typeStatus id="F92FA9C17D66868325C4299EB4711B94" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="CBF382C93A485F26C3F754C277956B0B" name="China">China</collectingCountry>
•
<specimenCount id="41D1C7CEC7F23C9CE241DF479615E7FF" count="3" type="male">3 ♂♂</specimenCount>
; same data as for holotype;
<collectionCode id="DEF14AEF001AD0FA971D39DA4DE72B6B" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" type="">CAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYUAE" type="diagnosis">
<paragraph id="91BCF6992B2F23C1D5FA8534B41DABB4">
<heading id="C3463CD2939EDEEB46E0968BE04D0389" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="AB945ECC005E469D0BC6BEA85DA46CA4">
Prescutum and presutural scutum dark brown with edges of prescutal suture and both sides of caudal edge darker (Fig.
<figureCitation id="3297668D5D7AB0CD9BA33700CA40F194" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 C</figureCitation>
). Pleuron dark brown. Wing with cell d open by absence of m-m. R
<subScript id="54E7709FA65FD7DEA183542C7D602ECA">4</subScript>
ending slightly before end of basal section of R
<subScript id="55CE6F8053760F090DE666E110513D83">5</subScript>
, m-cu before fork of M by about 1 / 3 of its own length. Gonocoxite with 1 gonostylus. Gonocoxite dorsally with three brownish-black, stout preapical setae, and a blunt lobe at inside of base; the lobe with nine stout setae. Interbase elongated, rod-shaped. Tip of aedeagus bifid with arms short and divergent.
</paragraph>
<caption id="181BB4CCB932EA04079296937DADA70C" ID-DOI="10.3897/zookeys.1205.117807.figure4" ID-arpha="72EDAF3E-6A6C-52F6-AFD9-4E0AFB0AF214" httpUri="https://binary.pensoft.net/fig/1078387" startId="F4">
<paragraph id="1783A663288D3A62327648001EC4A08C">
<label id="72CF35171D637E199BE3B385A6ED3921">Figure 4.</label>
</paragraph>
<paragraph id="1EE3C2B99CD0840143B59016E76B76C9">
<taxonomicName id="AB9D78967FC22C214684A442BA28F00D" authorityName="Lv &amp; Xu &amp; Liu &amp; Li &amp; Lin &amp; Zhang" authorityYear="2024" class="Insecta" family="Limoniidae" genus="Toxorhina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pianmica" status="sp. nov." subGenus="Ceratocheilus">
<emphasis id="EBD185D410277D2D6C98C2ACAF946EA1" italics="true">Toxorhina</emphasis>
(
<emphasis id="403E32626E33418E798E82020AE7CC01" italics="true">Ceratocheilus</emphasis>
)
<emphasis id="84A9F1915873FF4B2C9D4AFBB0965385" italics="true">pianmica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE2ECD16E636DEDEAE84D1850911B7A8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="A6F6E2E7AE29ED0306C383E59A8F7F94" bold="true">A</emphasis>
habitus of male, lateral view
<emphasis id="E079968E32573A9FAF726B755E080C4E" bold="true">B</emphasis>
head, lateral view
<emphasis id="2D2E56D9CBD0FE4D032A0D5C460EF35A" bold="true">C</emphasis>
thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum)
<emphasis id="8A8C6456ECCA1C04B077E1ADBD76DE08" bold="true">D</emphasis>
wing. Scale bars: 1.0 mm (
<emphasis id="07CA696DB1B1A146FDCF04010C294A74" bold="true">A, D</emphasis>
); 0.5 mm (
<emphasis id="7FE999512B2EA3917018450F952D1BE1" bold="true">B, C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ENWAE" type="description">
<paragraph id="71A0DF9675B9A9CD2C14A12FC47F603C">
<heading id="7F5DBA426955B1E746CDEFB777970A21" reason="title">Description.</heading>
</paragraph>
<paragraph id="2A1CFC138A547022D2830DD880EAA114">
<emphasis id="092113A071C526B370BEF160BC6149BC" bold="true">Male</emphasis>
(Fig.
<figureCitation id="15794D12693B002FBB92E12EFA3DE694" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 A</figureCitation>
). Body length
<quantity id="EB06101FA39666EDD67F14F3DA86603D" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.9" metricValueMin="4.4" unit="mm" value="4.65" valueMax="4.9" valueMin="4.4">4.4–4.9 mm</quantity>
(excluding rostrum), wing length 4.5–5.0 mm, rostrum length 2.7–3.0 mm, halter length
<quantity id="9CEA92C07A24A057AAE75BBFD887861D" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
.
</paragraph>
<paragraph id="CC5C6724B683DA3F114AE62858FCF53E">
Head (Fig.
<figureCitation id="DC1572F80218AE7E169EA71FACE70C17" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 B</figureCitation>
). Dark brown. Setae on head dark brown. Antenna with scape pale brown, pedicel and first flagellomere brown, remaining flagellomeres pale brown. Scape short cylindrical, with dark brown setae. Pedicel globular, with dark brown setae. First flagellomere oval, remaining flagellomeres cylindrical; terminal two flagellomeres longest with pale brown verticils. Rostrum about 3 / 5 of length of wing, brown with dark brown setae.
</paragraph>
<paragraph id="2257EED262ED52ADB1A995FC8AC7F596">
Thorax (Fig.
<figureCitation id="426DD45159DA4201E90631FE6F0C90B9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 C</figureCitation>
). Pronotum brown. Prescutum and presutural scutum dark brown, with edges of prescutal suture and both sides of caudal edge darker. Postsutural scutum dark brown, paler in middle area. Scutellum dark brown with side and caudal edges black, paler in middle area. Mediotergite brown with side edges brownish-black. Pleuron dark brown (Fig.
<figureCitation id="3FB43F9D8C2B2FF9B0B2CEB27350C72C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 A</figureCitation>
). Coxae yellow with bases brown; trochanters yellow; femora brown with bases paler; tibiae brown; base of fore tarsus brown, remaining tarsi missing. Setae on legs brown. Wing (Fig.
<figureCitation id="A2E6E70B2AA790E50C83460BEAF239B1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 D</figureCitation>
) pale brown, without stigma. Veins brown. Venation: Sc ending slightly beyond origin of Rs; sc-r a great distance before tip of Sc; R
<subScript id="6859CBD5BD98EF166FBAD9ED3C649281">4</subScript>
ending slightly before end of basal section of R
<subScript id="12EFDF7915AE2A7969679EC5B28FB1DE">5</subScript>
; distal section of R
<subScript id="E2B92EA637F7F2FFABE91B356380F5D3">5</subScript>
approach M
<subScript id="A20E8B1A3DC03E655A0A31E2BC89EBCE">1 + 2</subScript>
toward tip; CuP curved suddenly at middle; cell d open by absence of m-m; m-cu before fork of M by about 1 / 3 its own length. Halter yellow.
</paragraph>
<paragraph id="A21F871ADCD9B488DBB00D72757BC3C0">
Abdomen (Fig.
<figureCitation id="882C4AAA8E34F78638267F8BCA35D9DC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 A</figureCitation>
). Segments 1–6 brown with caudal edges darker, segments 7–8 dark brown.
</paragraph>
<paragraph id="02B6D789DB1B6F42CEA2F21BB6DF4972">
Hypopygium (Fig.
<figureCitation id="898B054CF8E1BCD8926807AF872BFE1F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5</figureCitation>
). Tergite 9 dark brown with dark brown setae, posterior margin with two large translucent areas on both sides (Fig.
<figureCitation id="9B786B5F0F142F6E74B6F391AF6B1445" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A, E</figureCitation>
). Gonocoxite brown, conical with tip round (Fig.
<figureCitation id="0D97F070A9A03E8A6766DF75C99C555B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A, B</figureCitation>
); dorsal face with a few sparse brown setae at outside, three brownish-black, stout preapical setae (Fig.
<figureCitation id="C7EB6C43D92503B65EC36D6AA930280B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A, F</figureCitation>
), and a blunt lobe at inside of base, this lobe with nine stout setae (Fig.
<figureCitation id="0A0268CFECDA9FCA9B8ACA102F3DE30C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A, G</figureCitation>
); ventral face with numerous brown setae at inside (Fig.
<figureCitation id="331996347A4779767DF4E99FCBC26255" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 B</figureCitation>
). Gonostylus brownish-yellow, curved inwards at basal 1 / 3; the basal 1 / 3 stout with a longitudinal groove, distal 2 / 3 slender with tip round; outer side of basal 1 / 3 with outwardly curved spine (Fig.
<figureCitation id="FCC28C1CF256250B8B1F114E409731A0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A, B, H</figureCitation>
). Interbase pale brownish-yellow, elongated, rod-shaped, about 10 times as long as wide; tip inflated and round (Fig.
<figureCitation id="CCBB9793D37199DB28C707BEC56F84BD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A – D</figureCitation>
). Parameres dark brown, medially arched fused, distally connecting to base of interbase (Fig.
<figureCitation id="A4E1E3579D5AFFB41370E77B248622B8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 C, D</figureCitation>
). Ejaculatory apodeme dark brown except pale base, rod-shaped (Fig.
<figureCitation id="FA91872A7A8715D1DAD675859D5E3918" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 C, D</figureCitation>
). Aedeagus dark brown, stout at base, tip bifid with arms short and divergent (Fig.
<figureCitation id="B7DCE55681E026FAF80797A08DE3E3D4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A – D, I</figureCitation>
).
</paragraph>
<caption id="FF530BD3D4AFB3A088092B01E76F3731" ID-DOI="10.3897/zookeys.1205.117807.figure5" ID-arpha="07F85419-C069-5BA8-839A-456FF05DB21E" httpUri="https://binary.pensoft.net/fig/1078388" startId="F5">
<paragraph id="D9826882D60DFAFA15328ADA1F326700">
<label id="D3FD371A1B83EA561824C8AC35168209">Figure 5.</label>
</paragraph>
<paragraph id="4C300AD7DBF9EF31D8C7E77699620E80">
<taxonomicName id="7D0AF14F424448BD067FD0EDEB1D58F2" authorityName="Lv &amp; Xu &amp; Liu &amp; Li &amp; Lin &amp; Zhang" authorityYear="2024" class="Insecta" family="Limoniidae" genus="Toxorhina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="pianmica" status="sp. nov." subGenus="Ceratocheilus">
<emphasis id="D2ADD884C06E94AFAF71C9355ADAB97C" italics="true">Toxorhina</emphasis>
(
<emphasis id="BF09C4B57B8D83916CD1E1239374945D" italics="true">Ceratocheilus</emphasis>
)
<emphasis id="F802596BF16C0B6AC09C6C756710F86F" italics="true">pianmica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4213B9C53D19403EFE0C6585EBFEC055" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="597EEC3D1F6DB91E6D3558F7A31C62A8" bold="true">A</emphasis>
male hypopygium, dorsal view
<emphasis id="BB6BD756F29AAD56C7F5F189D683A1CF" bold="true">B</emphasis>
male hypopygium, ventral view
<emphasis id="A927B1D6BD4BFE77BC70A95E10EC319E" bold="true">C</emphasis>
aedeagal complex, dorsal view
<emphasis id="59E998EBC5FD8D6B18E62A770033B4D8" bold="true">D</emphasis>
aedeagal complex, ventral view
<emphasis id="41279817062488F4A1CDA6F0AC1DB10A" bold="true">E</emphasis>
posterior margin of tergite 9, dorsal view
<emphasis id="5B53CD87A388EF7B80E7752B409FF416" bold="true">F</emphasis>
preapical setae gonocoxite, dorsal view
<emphasis id="E6A0ECA73D0CC0C2E1883D472CBB3A34" bold="true">G</emphasis>
setae at base of gonocoxite, dorsal view
<emphasis id="5F42C85FD6E5E1804944306E9A5A76F8" bold="true">H</emphasis>
gonostylus, dorsal view
<emphasis id="98502D9A04A4A52CB3FB269E6F20912B" bold="true">I</emphasis>
tip of aedeagus, ventral view. Scale bars: 0.1 mm (
<emphasis id="CB5FA65B9146F62D58CAEC21CDB4E85A" bold="true">A – E, H</emphasis>
); 0.02 mm (
<emphasis id="2A7BF96265F1DFE4621A2A0B2DC1012B" bold="true">F, G, I</emphasis>
).
</paragraph>
</caption>
<paragraph id="5EF7CF5704B2B7451D58F664355FF918">
<emphasis id="1C7BB86FB60C3723C3142F63EC7B9DDC" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E51AE" type="etymology">
<paragraph id="8DC7C54A83854B9579EF22B92E7C67E8">
<heading id="765C2BA55F9F2542EF5431051C6E0BDE" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="CB043E9ADC36BD96F872A8FE3F2481F3">The species is named after the type locality, Pianma Town.</paragraph>
</subSubSection>
<subSubSection id="SECID0ED2AE" type="distribution">
<paragraph id="03DB0EA269DB6D994847CD0B51B1F28C">
<heading id="C101E8ECA1F3DF85564811DB6B2AA1D3" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="93D9FB635776B6A50A6EE965DE8C817C">
<collectingCountry id="9EB5AF75B01B0290BD1EA49CD453148D" name="China">China</collectingCountry>
(
<collectingRegion id="2A4B69D4D983527996E51A5E38C4C08F" country="China" name="Yunnan">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EI2AE" type="Behavior">
<paragraph id="D720655294C9D1B64536CABC65EBAE41">
<heading id="FE0990968C4C6D0CC9A3F3C28246200B" reason="title">Behavior.</heading>
</paragraph>
<paragraph id="F43851B853C30D77254C7042E64CC231">The species can be attracted by light.</paragraph>
</subSubSection>
<subSubSection id="SECID0EN2AE" type="remarks">
<paragraph id="5627610DE380A4C975B75E257645BDF1">
<heading id="A471576561556C70442067B5B5F9EA15" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="376E4C6AA58D2462B60ACE30A63CC8C9">
The new species is similar to
<taxonomicName id="9B73C7DBA2DF18D306C749AFA5717D20" authority="Alexander, 1967" authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Toxorhina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="simplicistyla" subGenus="Ceratocheilus">
<emphasis id="804F08FF88D6C4C01758A7A0FB8758B4" italics="true">T.</emphasis>
(
<emphasis id="D98BB8F643B567FB786E5936CC16B26B" italics="true">C.</emphasis>
)
<emphasis id="21B0FE17304332721AD8073473345197" italics="true">simplicistyla</emphasis>
<bibRefCitation id="A9F002C0DEB8CF83A5FBFD2610BA2F3B" author="Alexander CP" journalOrPublisher="Journal of the New York Entomological Society" pagination="183-187" refId="B5" refString="Alexander CP (1967) Undescribed species of crane flies from the Himalaya mountains (Diptera: Tipulidae). XV. Journal of the New York Entomological Society 75: 183 - 187." title="Undescribed species of crane flies from the Himalaya mountains (Diptera: Tipulidae). XV." volume="75" year="1967">Alexander, 1967</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="92C461F094546C40BF31ABF9AA574C01" name="India">India</collectingCountry>
in having similar wing venation, but can be separated by the brown abdomen with segments 7–8 darker (Fig.
<figureCitation id="A5579652A673D4EA3331A3B627F5478D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Toxorhina (Ceratocheilus) pianmica sp. nov. A habitus of male, lateral view B head, lateral view C thorax, dorsal view (the arrows refer to the dark areas on prescutum and presutural scutum) D wing. Scale bars: 1.0 mm (A, D); 0.5 mm (B, C)." figureDoi="10.3897/zookeys.1205.117807.figure4" httpUri="https://binary.pensoft.net/fig/1078387">4 A</figureCitation>
), the gonostylus conspicuously curved with a spine at the basal 1 / 3 (Fig.
<figureCitation id="DE5C27A7E5BE0076F6A88D68F25612B4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A, B, H</figureCitation>
), and the interbase with the tip inflated (Fig.
<figureCitation id="76761D126D86DC8BF67EBF008D7A92C0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Toxorhina (Ceratocheilus) pianmica sp. nov. A male hypopygium, dorsal view B male hypopygium, ventral view C aedeagal complex, dorsal view D aedeagal complex, ventral view E posterior margin of tergite 9, dorsal view F preapical setae gonocoxite, dorsal view G setae at base of gonocoxite, dorsal view H gonostylus, dorsal view I tip of aedeagus, ventral view. Scale bars: 0.1 mm (A – E, H); 0.02 mm (F, G, I)." figureDoi="10.3897/zookeys.1205.117807.figure5" httpUri="https://binary.pensoft.net/fig/1078388">5 A – D</figureCitation>
). In
<taxonomicName id="1A751D7EC32DF7B5DC80A8C7D77BB2AD" authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Toxorhina" kingdom="Animalia" order="Diptera" phylum="Arthropoda" rank="species" species="simplicistyla" subGenus="Ceratocheilus">
<emphasis id="2DC29A54C369355F4320AA91550AAA18" italics="true">T.</emphasis>
(
<emphasis id="98A4180EB2F26613266EE3CD6AC4D9F6" italics="true">C.</emphasis>
)
<emphasis id="6CC9F6064F144963834B4BA58B1F50FD" italics="true">simplicistyla</emphasis>
</taxonomicName>
, the abdomen is brownish-black, the gonostylus is very gently curved with a spine near the middle, and the tip of the interbase is not inflated (
<bibRefCitation id="B7A73572C80366C470ED11E06E84AF9A" author="Alexander" firstAuthor="Alexander" journalOrPublisher="Journal of the New York Entomological Society" pagination="183-187" refId="B5" refString="Alexander CP (1967) Undescribed species of crane flies from the Himalaya mountains (Diptera: Tipulidae). XV. Journal of the New York Entomological Society 75: 183–187." title="Undescribed species of crane flies from the Himalaya mountains (Diptera: Tipulidae). XV." volume="75" year="1967">Alexander 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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