154 lines
18 KiB
XML
154 lines
18 KiB
XML
<document id="84127B1C0D58F1DDF23BD353BACEACAD" ID-DOI="10.11646/zootaxa.4748.1.3" ID-GBIF-Dataset="f267c3f7-48d8-45f7-a480-b1660dc6b391" ID-ISSN="1175-5326" ID-Zenodo-Dep="3697578" ID-ZooBank="6F6367D1-5F1B-439D-B737-FEF761155D5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583398626205" checkinUser="plazi" docAuthor="Evans, Arthur V. & Smith, Andrew B. T." docDate="2020" docId="B63C87C2F375FFB72FA8FBE3FBA5FE58" docLanguage="en" docName="zootaxa.4748.1.3.pdf" docOrigin="Zootaxa 4748 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acoma eusexfoliata Evans & Smith 2020, new species" docType="treatment" docUuid="3E6AE0B3-F1E1-4B43-B817-93408AC25611" docUuidSource="ZooBank" docVersion="8" lastPageNumber="59" masterDocId="4A05FFBAF372FFBF2F3FFFFEFFD0FFF6" masterDocTitle="On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889" masterLastPageNumber="77" masterPageNumber="51" pageNumber="58" updateTime="1698724959701" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="948216877D98AB90ABA92AE9B0866792">
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<mods:title id="2FB8CF1B960C88A1F0F6549E43F01660">On the tribal classification of the Nearctic Melolonthinae (Coleoptera: Scarabaeidae), with descriptions of new species of Acoma Casey, 1889</mods:title>
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<mods:namePart id="503216E82D09C94C515F9E4C4586A580">Evans, Arthur V.</mods:namePart>
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<mods:name id="52C9EBE36D55429CBA2687B43CEB809B" type="personal">
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<mods:namePart id="AAA3F5DBBC048AEBA110A48F9D4E497D">Smith, Andrew B. T.</mods:namePart>
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<mods:title id="1715E4D2EF018BD96169C3BEB8A6EFDC">Zootaxa</mods:title>
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<mods:part id="C07040A66CFCEB094602F11CDA39A1A2">
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<mods:date id="20E3A397EE89ED5C5975FB004FAAEA7E">2020</mods:date>
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<mods:number id="8D8734C9A1A0E69E215F709BE35850E8">2020-03-05</mods:number>
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<mods:number id="94B8C870707E3CE921F610C83F8275F5">4748</mods:number>
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<mods:classification id="5D2A3B95059C96E23891309D6B298704">journal article</mods:classification>
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<mods:identifier id="76D68FA181F7B9F663F186F8A9D7F08A" type="DOI">10.11646/zootaxa.4748.1.3</mods:identifier>
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<treatment id="B63C87C2F375FFB72FA8FBE3FBA5FE58" ID-DOI="http://doi.org/10.5281/zenodo.3704218" ID-GBIF-Taxon="162443304" ID-Zenodo-Dep="3704218" LSID="urn:lsid:zoobank.org:act:3E6AE0B3-F1E1-4B43-B817-93408AC25611" httpUri="http://treatment.plazi.org/id/B63C87C2F375FFB72FA8FBE3FBA5FE58" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
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<subSubSection id="768F655FF375FFB82FA8FBE3FD3AFBC1" box="[151,746,1053,1080]" pageId="7" pageNumber="58" type="nomenclature">
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<paragraph id="3E2A36D4F375FFB82FA8FBE3FD3AFBC1" blockId="7.[151,919,1053,1144]" box="[151,746,1053,1080]" pageId="7" pageNumber="58">
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<heading id="656281B8F375FFB82FA8FBE3FD3AFBC1" bold="true" box="[151,746,1053,1080]" fontSize="11" level="1" pageId="7" pageNumber="58" reason="1">
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<emphasis id="0CE1EAC6F375FFB82FA8FBE3FD3AFBC1" bold="true" box="[151,746,1053,1080]" pageId="7" pageNumber="58">
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<taxonomicName id="F9954D57F375FFB82FA8FBE3FD9EFBC1" authority="Evans & Smith" authorityName="Evans & Smith" authorityYear="2020" box="[151,590,1053,1080]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="eusexfoliata" status="sp. nov.">
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<emphasis id="0CE1EAC6F375FFB82FA8FBE3FE56FBCE" bold="true" box="[151,390,1053,1080]" italics="true" pageId="7" pageNumber="58">Acoma eusexfoliata</emphasis>
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Evans & Smith
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</taxonomicName>
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,
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<taxonomicNameLabel id="17D257BDF375FFB82D64FBE3FD3AFBC1" box="[603,746,1053,1079]" pageId="7" pageNumber="58" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="768F655FF375FFB82FA8FBBFFC47FB8E" pageId="7" pageNumber="58" type="description">
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<paragraph id="3E2A36D4F375FFB82FA8FBBFFECBFBAD" blockId="7.[151,919,1053,1144]" box="[151,283,1089,1115]" pageId="7" pageNumber="58">
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<figureCitation id="A6AE2A51F375FFB82FA8FBBFFECBFBAD" box="[151,283,1089,1115]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1786,1811]" captionTargetBox="[151,1435,452,1760]" captionTargetId="figure@8.[151,1435,452,1760]" captionTargetPageId="8" captionText="FIGURES 11–16. Acoma eusexfoliata Evans & Smith, new species, male holotype. 11, Dorsal habitus; 12, oblique habitus; 13, ventral habitus; 14, head; 15, dorsal parameres; 16, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697584" httpUri="https://zenodo.org/record/3697584/files/figure.png" pageId="7" pageNumber="58">Figs. 11–16</figureCitation>
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</paragraph>
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<paragraph id="3E2A36D4F375FFB82FA8FB9EFC47FB8E" blockId="7.[151,919,1053,1144]" box="[151,919,1120,1144]" pageId="7" pageNumber="58">
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<uri id="4A043AD6F375FFB82FA8FB9EFC47FB8E" box="[151,919,1120,1144]" pageId="7" pageNumber="58">
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urn:lsid:zoobank.org:act:
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<uuid id="4A330C01F375FFB82E9DFB9EFC47FB8E" box="[418,919,1120,1144]" pageId="7" pageNumber="58">3E6AE0B3-F1E1-4B43-B817-93408AC25611</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="768F655FF375FFB82FA8FB58FAFDFAFE" pageId="7" pageNumber="58" type="materials_examined">
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<paragraph id="3E2A36D4F375FFB82FA8FB58FAFDFAFE" blockId="7.[151,1437,1189,2008]" pageId="7" pageNumber="58">
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<emphasis id="0CE1EAC6F375FFB82FA8FB58FEEEFB36" bold="true" box="[151,318,1189,1216]" pageId="7" pageNumber="58">Type material</emphasis>
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.
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<materialsCitation id="8EFD3C89F375FFB82E72FB58FAFDFAFE" ID-GBIF-Occurrence="2573655799" collectorName="R. L. Aalbu & Dr. Alan Hardy" country="Mexico" location="San Francisquito" pageId="7" pageNumber="58" specimenCount="2" typeStatus="holotype">
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<typeStatus id="E12E8876F375FFB82E72FB58FE65FB36" box="[333,437,1190,1216]" pageId="7" pageNumber="58" type="holotype">Holotype</typeStatus>
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male (CASTYPE 19860), labeled “
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<collectingCountry id="46827644F375FFB82C79FB58FC6AFB36" box="[838,954,1190,1216]" name="Mexico" pageId="7" pageNumber="58">MEXICO</collectingCountry>
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, Baja Calif. Norte /
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<quantity id="F96D9B31F375FFB82BA2FB58FB07FB36" box="[1181,1239,1190,1216]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="7" pageNumber="58" unit="mi" value="2.0">2 mi.</quantity>
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SW
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<location id="3B4A600FF375FFB82A30FB58FF01FB12" LSID="urn:lsid:plazi:treatment:B63C87C2F375FFB72FA8FBE3FBA5FE58:3B4A600FF375FFB82A30FB58FF01FB12" country="Mexico" name="San Francisquito" pageId="7" pageNumber="58">San Francisquito</location>
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/ box canyon with caves BL / VII-26/27-86
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<collectorName id="93605302F375FFB82D80FB35FC91FB12" box="[703,833,1226,1253]" pageId="7" pageNumber="58">R. L. Aalbu</collectorName>
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//
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<collectorName id="93605302F375FFB82C61FB35FBD9FB12" box="[862,1033,1226,1253]" pageId="7" pageNumber="58">Dr. Alan Hardy</collectorName>
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/ Donation to the / California Academy / of Sciences // CASENT / 8424551 //
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<typeStatus id="E12E8876F375FFB82D42FB10FCC1FAFE" box="[637,785,1262,1288]" pageId="7" pageNumber="58" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="F9954D57F375FFB82C1AFB10FCA6FAFE" authorityName="Casey" authorityYear="1889" box="[805,886,1262,1288]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">Acoma</taxonomicName>
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/ eusexfoliata / Evans & Smith, 2019”.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="768F655FF375FFB82FF8FAEFFA4CFA83" pageId="7" pageNumber="58" type="diagnosis">
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<paragraph id="3E2A36D4F375FFB82FF8FAEFFA4CFA83" blockId="7.[151,1437,1189,2008]" pageId="7" pageNumber="58">
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<emphasis id="0CE1EAC6F375FFB82FF8FAEFFE92FADA" bold="true" box="[199,322,1297,1324]" pageId="7" pageNumber="58">Diagnosis.</emphasis>
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Light reddish testaceous dorsally with head and pronotum darker; anterior clypeal margin weakly emarginate and strongly reflexed; antennae composed of 9 antennomeres, antennomere 3 much longer than 1+2 with anterior projection at base, 4–9 lamellate; protibia apparently bidentate; elytral disc with scattered erect setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="768F655FF375FFB72FF8FA83FC9CFF44" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58" type="description">
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<paragraph id="3E2A36D4F375FFB82FF8FA83FD96F82E" blockId="7.[151,1437,1189,2008]" pageId="7" pageNumber="58">
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<emphasis id="0CE1EAC6F375FFB82FF8FA83FE89FA6E" bold="true" box="[199,345,1405,1432]" pageId="7" pageNumber="58">Description.</emphasis>
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<typeStatus id="E12E8876F375FFB82E5DFA80FE1AFA6E" box="[354,458,1406,1432]" pageId="7" pageNumber="58" type="holotype">Holotype</typeStatus>
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male (
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<figureCitation id="A6AE2A51F375FFB82D23FA80FD76FA6F" box="[540,678,1406,1433]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1786,1811]" captionTargetBox="[151,1435,452,1760]" captionTargetId="figure@8.[151,1435,452,1760]" captionTargetPageId="8" captionText="FIGURES 11–16. Acoma eusexfoliata Evans & Smith, new species, male holotype. 11, Dorsal habitus; 12, oblique habitus; 13, ventral habitus; 14, head; 15, dorsal parameres; 16, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697584" httpUri="https://zenodo.org/record/3697584/files/figure.png" pageId="7" pageNumber="58">Figs. 11–16</figureCitation>
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), length: 6.0 mm, width: 4.0 mm. BODY somewhat robust, testaceous, with head, pronotum and elytral apices slightly darker. HEAD (
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<figureCitation id="A6AE2A51F375FFB82CF4FA5CFBCFFA4A" box="[971,1055,1442,1468]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1786,1811]" captionTargetBox="[151,1435,452,1760]" captionTargetId="figure@8.[151,1435,452,1760]" captionTargetPageId="8" captionText="FIGURES 11–16. Acoma eusexfoliata Evans & Smith, new species, male holotype. 11, Dorsal habitus; 12, oblique habitus; 13, ventral habitus; 14, head; 15, dorsal parameres; 16, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697584" httpUri="https://zenodo.org/record/3697584/files/figure.png" pageId="7" pageNumber="58">Fig. 14</figureCitation>
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) with vertex rugosely punctuate, clypeus shagreened with large shallow punctures separated by their own diameters or more; supraorbital carinae becoming obsolete at bases of head and clypeus. Clypeus subtrapezoidal with lateral margins broadly arcuate and weakly reflexed; anterior margin shallowly concave, not quite truncate, strongly reflexed, and angulate. Ocular canthus about one-quarter of eye length when viewed from side, with scattered pale setae each longer than canthus. PRONOTUM wider than long, broadest before middle, slightly more than one-third length of elytron; disc strongly convex, sparsely and shallowly punctate, with broad impunctate area medially; lateral and posterior margins with bead; anterior margin broadly concave between obtuse anterior angles; anterolateral margins straight and convergent anteriorly, posterolateral margins slightly concave and convergent posteriorly, with posterior angles broadly rounded. SCUTELLUM broadly parabolic, about one-quarter width of elytron, setigerously punctate at base, finely shagreened medially and apically. ELYTRON convex, shiny, finely shagreened, disc punctostriate with punctures becoming confused on apical and lateral declivities, setigerous punctures with short, suberect setae. LEGS with protibial apparently bidentate, basal third tooth barely indicated by weak angulation. PYGIDIUM parabolic, finely shagreened, with scattered setigerous punctures, setae long. GENITALIA with parameres symmetrical, moderately slender and more or less straight when viewed laterally, with membranous flange extending to apical 3/5, and shorter than phallobasis (
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<figureCitation id="A6AE2A51F375FFB82E8FF840FDEAF82E" box="[432,570,1981,2008]" captionStart="FIGURES 11–16" captionStartId="8.[151,264,1786,1811]" captionTargetBox="[151,1435,452,1760]" captionTargetId="figure@8.[151,1435,452,1760]" captionTargetPageId="8" captionText="FIGURES 11–16. Acoma eusexfoliata Evans & Smith, new species, male holotype. 11, Dorsal habitus; 12, oblique habitus; 13, ventral habitus; 14, head; 15, dorsal parameres; 16, lateral genitalia." figureDoi="http://doi.org/10.5281/zenodo.3697584" httpUri="https://zenodo.org/record/3697584/files/figure.png" pageId="7" pageNumber="58">Figs. 15–16</figureCitation>
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).
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</paragraph>
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<paragraph id="3E2A36D4F37AFFB72FF8FF66FC9CFF44" blockId="8.[151,1436,152,431]" box="[199,844,152,179]" pageId="8" pageNumber="59">
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<emphasis id="0CE1EAC6F37AFFB72FF8FF66FEEFFF45" bold="true" box="[199,319,152,179]" pageId="8" pageNumber="59">Variation.</emphasis>
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This species is known only from the
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<typeStatus id="E12E8876F37AFFB72DDDFF66FC98FF44" box="[738,840,152,178]" pageId="8" pageNumber="59" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="768F655FF37AFFB72FF8FF43FDB3FF0D" pageId="8" pageNumber="59" type="etymology">
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<paragraph id="3E2A36D4F37AFFB72FF8FF43FDB3FF0D" blockId="8.[151,1436,152,431]" pageId="8" pageNumber="59">
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<emphasis id="0CE1EAC6F37AFFB72FF8FF43FE9EFF21" bold="true" box="[199,334,189,215]" pageId="8" pageNumber="59">Etymology.</emphasis>
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This species is named for actually having 6 antennomeres in its antennal club, unlike
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<taxonomicName id="F9954D57F37AFFB72A2FFF43FEF8FF0D" authority="Saylor, 1948" authorityName="Saylor" authorityYear="1948" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="sexfoliata">
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<emphasis id="0CE1EAC6F37AFFB72A2FFF43FAF1FF21" box="[1296,1313,189,215]" italics="true" pageId="8" pageNumber="59">A</emphasis>
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.
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<emphasis id="0CE1EAC6F37AFFB72A10FF42FA4CFF20" box="[1327,1436,188,214]" italics="true" pageId="8" pageNumber="59">sexfoliata</emphasis>
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Saylor, 1948
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</taxonomicName>
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, which has 7 antennomeres.
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</paragraph>
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</subSubSection>
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<subSubSection id="768F655FF37AFFB72FF8FEFBFBA5FE58" pageId="8" pageNumber="59" type="discussion">
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<paragraph id="3E2A36D4F37AFFB72FF8FEFBFBA5FE58" blockId="8.[151,1436,152,431]" pageId="8" pageNumber="59">
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<emphasis id="0CE1EAC6F37AFFB72FF8FEFBFEEBFEE9" bold="true" box="[199,315,261,287]" pageId="8" pageNumber="59">Remarks.</emphasis>
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<taxonomicName id="F9954D57F37AFFB72E7DFEFBFDCEFEE8" authority="Evans & Smith, 2020" authorityName="Evans & Smith" authorityYear="2020" box="[322,542,260,287]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="eusexfoliata" status="sp. nov.">
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<emphasis id="0CE1EAC6F37AFFB72E7DFEFBFDCEFEE8" box="[322,542,260,287]" italics="true" pageId="8" pageNumber="59">Acoma eusexfoliata</emphasis>
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</taxonomicName>
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<emphasis id="0CE1EAC6F37AFFB72D19FEFBFD60FEE9" bold="true" box="[550,688,260,287]" pageId="8" pageNumber="59">
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<taxonomicNameLabel id="17D257BDF37AFFB72D19FEFBFD60FEE9" box="[550,688,260,287]" pageId="8" pageNumber="59" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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is distinguished from all other known
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<taxonomicName id="F9954D57F37AFFB72B5BFEFBFB62FEE9" authorityName="Casey" authorityYear="1889" box="[1124,1202,261,287]" class="Insecta" family="Pleocomidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="0CE1EAC6F37AFFB72B5BFEFBFB62FEE9" box="[1124,1202,261,287]" italics="true" pageId="8" pageNumber="59">Acoma</emphasis>
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</taxonomicName>
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species by its antennal configuration. The unique
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<typeStatus id="E12E8876F37AFFB72ED6FED6FD9BFEB4" box="[489,587,296,322]" pageId="8" pageNumber="59" type="holotype">holotype</typeStatus>
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was originally glued by its abdomen to a point. The specimen was placed in hot water and removed from the point and pinned. While removing the abdomen for genitalic dissection, the right protibia was accidentally detached the body at the base of the femur. Rather than gluing the detached appendage back in place, it was affixed intact onto a separate point and pinned below the specimen.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |