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<mods:title id="156961AF5EA59B868990D59540679815">Further notes on the Sri Lankan uropeltid snakes Rhinophis saffragamus (Kelaart, 1853) and Uropeltis ruhunae Deraniyagala, 1954</mods:title>
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<treatment id="9239F147FFE15F77FF5754709A3909D9" ID-DOI="http://doi.org/10.5281/zenodo.5921261" ID-GBIF-Taxon="154533353" ID-Zenodo-Dep="5921261" LSID="urn:lsid:plazi:treatment:9239F147FFE15F77FF5754709A3909D9" httpUri="http://treatment.plazi.org/id/9239F147FFE15F77FF5754709A3909D9" lastPageNumber="592" pageId="0" pageNumber="592">
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<paragraph id="1A2F4051FFE15F77FF5754709C610F8F" blockId="0.[151,1436,884,1497]" box="[151,621,884,910]" pageId="0" pageNumber="592">
<heading id="4167F73DFFE15F77FF5754709C610F8F" bold="true" box="[151,621,884,910]" fontSize="11" level="1" pageId="0" pageNumber="592" reason="1">
<taxonomicName id="DD903BD2FFE15F77FF5754709C610F8F" authority="Deraniyagala, 1954" authorityName="Deraniyagala" authorityYear="1954" box="[151,621,884,910]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="ruhunae">
<emphasis id="28E49C43FFE15F77FF5754709C610F8F" bold="true" box="[151,621,884,910]" pageId="0" pageNumber="592">
<emphasis id="28E49C43FFE15F77FF5754709F790F8F" bold="true" box="[151,373,884,910]" italics="true" pageId="0" pageNumber="592">Uropeltis ruhunae</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FEBC54709C610F8F" author="Deraniyagala, P." box="[380,621,884,910]" pageId="0" pageNumber="592" pagination="27 - 28" refId="ref2973" refString="Deraniyagala, P. (1954) Two new snakes from Ceylon. Proceedings of the Tenth Annual Session of the Ceylon Association for the Advancement of Science (25 th- 27 th Nov. 1954), 1, 27 - 28 [24]." type="journal article" year="1954">Deraniyagala, 1954</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="1A2F4051FFE15F77FF5754939A3909D9" blockId="0.[151,1436,884,1497]" pageId="0" pageNumber="592">
As detailed by
<bibRefCitation id="7E013DA0FFE15F77FE8254939C160FAF" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[322,538,919,942]" pageId="0" pageNumber="592" pagination="453 - 506" refId="ref3447" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V. &amp; Somaweera, R. (2016) A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae). Zoosystema, 38, 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2" type="journal article" year="2016">
Pyron
<emphasis id="28E49C43FFE15F77FE4D54939FB30FAF" box="[397,447,919,942]" italics="true" pageId="0" pageNumber="592">et al</emphasis>
. (2016)
</bibRefCitation>
,
<bibRefCitation id="7E013DA0FFE15F77FDE854939D040FAF" author="Deraniyagala, P." box="[552,776,919,942]" pageId="0" pageNumber="592" pagination="27 - 28" refId="ref2973" refString="Deraniyagala, P. (1954) Two new snakes from Ceylon. Proceedings of the Tenth Annual Session of the Ceylon Association for the Advancement of Science (25 th- 27 th Nov. 1954), 1, 27 - 28 [24]." type="journal article" year="1954">Deraniyagala (1954)</bibRefCitation>
described
<emphasis id="28E49C43FFE15F77FC4354939A250FAF" box="[899,1065,919,942]" italics="true" pageId="0" pageNumber="592">Platyplectrurus</emphasis>
<taxonomicName id="DD903BD2FFE15F77FBF354939B670FAF" authority="Deraniyagala, 1954" authorityName="Deraniyagala" authorityYear="1954" box="[1075,1387,919,942]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="ruhunae">
<emphasis id="28E49C43FFE15F77FBF354939A800FAF" box="[1075,1164,919,942]" italics="true" pageId="0" pageNumber="592">ruhunae</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FB5654939B670FAF" author="Deraniyagala, P." box="[1174,1387,919,942]" pageId="0" pageNumber="592" pagination="27 - 28" refId="ref2973" refString="Deraniyagala, P. (1954) Two new snakes from Ceylon. Proceedings of the Tenth Annual Session of the Ceylon Association for the Advancement of Science (25 th- 27 th Nov. 1954), 1, 27 - 28 [24]." type="journal article" year="1954">Deraniyagala, 1954</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="DD903BD2FFE15F77FF5754B39C3C0FCF" authority="Deraniyagala, 1954" authorityName="Deraniyagala" authorityYear="1954" box="[151,560,951,974]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="ruhunae">
<emphasis id="28E49C43FFE15F77FF5754B39F560FCF" box="[151,346,951,974]" italics="true" pageId="0" pageNumber="592">Uropeltis ruhunae</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FEA054B39C3C0FCF" author="Deraniyagala, P." box="[352,560,951,974]" pageId="0" pageNumber="592" pagination="27 - 28" refId="ref2973" refString="Deraniyagala, P. (1954) Two new snakes from Ceylon. Proceedings of the Tenth Annual Session of the Ceylon Association for the Advancement of Science (25 th- 27 th Nov. 1954), 1, 27 - 28 [24]." type="journal article" year="1954">Deraniyagala, 1954</bibRefCitation>
</taxonomicName>
(both with type locality “Galle”) from specimens found in a jar containing snakes from both the Galle district of
<collectingCountry id="628700C1FFE15F77FE1D54D39C4A0FEF" box="[477,582,983,1006]" name="Sri Lanka" pageId="0" pageNumber="592">Sri Lanka</collectingCountry>
and the Madurai district of
<collectingCountry id="628700C1FFE15F77FCB054D39DAB0FEF" box="[880,935,983,1006]" name="India" pageId="0" pageNumber="592">India</collectingCountry>
(
<bibRefCitation id="7E013DA0FFE15F77FC7654D39A580FEF" author="De Silva, P. H. D. H." box="[950,1108,983,1006]" pageId="0" pageNumber="592" refId="ref2929" refString="De Silva, P. H. D. H. (1980) Snake fauna of Sri Lanka: with special reference to skull, dentition, and venom in snakes. National Museums of Sri Lanka, Colombo, 472 pp." type="book" year="1980">De Silva 1980</bibRefCitation>
). Thus, origin of the
<typeStatus id="C52BFEF3FFE15F77FAF454D39B900FEF" box="[1332,1436,983,1006]" pageId="0" pageNumber="592" type="holotype">holotypes</typeStatus>
from
<collectingCountry id="628700C1FFE15F77FF1354F39F32080F" box="[211,318,1015,1038]" name="Sri Lanka" pageId="0" pageNumber="592">Sri Lanka</collectingCountry>
is in serious doubt. While recent works (e.g.,
<bibRefCitation id="7E013DA0FFE15F77FCF954F39A3A080F" author="McDiarmid, R. W. &amp; Campbell J. A. &amp; Toure, T." box="[825,1078,1015,1038]" pageId="0" pageNumber="592" refId="ref3278" refString="McDiarmid, R. W., Campbell J. A. &amp; Toure, T. (1999) Snake species of the world: a taxonomic and geographic reference. Herpetologists' League, Washington, 511 pp." type="book" year="1999">
McDiarmid
<emphasis id="28E49C43FFE15F77FC7E54F39DE3080F" box="[958,1007,1015,1038]" italics="true" pageId="0" pageNumber="592">et al</emphasis>
. 1999
</bibRefCitation>
;
<bibRefCitation id="7E013DA0FFE15F77FB8354F39B15080F" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[1091,1305,1015,1038]" pageId="0" pageNumber="592" refId="ref3877" refString="Wallach, V., Williams, K. L. &amp; Boundy, J. (2014) Snakes of the world: a catalogue of living and extinct species. CRC Press, Boca Raton, 1209 pp. https: // doi. org / 10.1201 / b 16901" type="book" year="2014">
Wallach
<emphasis id="28E49C43FFE15F77FB6254F39ADF080F" box="[1186,1235,1015,1038]" italics="true" pageId="0" pageNumber="592">et al</emphasis>
. 2014
</bibRefCitation>
) considered
<emphasis id="28E49C43FFE15F77FF5753139F31082F" box="[151,317,1047,1070]" italics="true" pageId="0" pageNumber="592">Platyplectrurus</emphasis>
<taxonomicName id="DD903BD2FFE15F77FE8453139C79082F" authority="Deraniyagala, 1954" authorityName="Deraniyagala" authorityYear="1954" box="[324,629,1047,1070]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="ruhunae">
<emphasis id="28E49C43FFE15F77FE8453139F91082F" box="[324,413,1047,1070]" italics="true" pageId="0" pageNumber="592">ruhunae</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FE6453139C79082F" author="Deraniyagala, P." box="[420,629,1047,1070]" pageId="0" pageNumber="592" pagination="27 - 28" refId="ref2973" refString="Deraniyagala, P. (1954) Two new snakes from Ceylon. Proceedings of the Tenth Annual Session of the Ceylon Association for the Advancement of Science (25 th- 27 th Nov. 1954), 1, 27 - 28 [24]." type="journal article" year="1954">Deraniyagala, 1954</bibRefCitation>
</taxonomicName>
to be a synonym of
<emphasis id="28E49C43FFE15F77FC9253139A75082F" box="[850,1145,1047,1070]" italics="true" pageId="0" pageNumber="592">Platyplectrurus madurensis</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FB4053139B20082F" author="Beddome, R. H." box="[1152,1324,1047,1070]" pageId="0" pageNumber="592" pagination="167 - 168" refId="ref2605" refString="Beddome, R. H. (1877) Descriptions of three new snakes of the family Uropeltidae from southern India. Proceedings of the Zoological Society of London, 45, 167 - 168." type="journal article" year="1877">Beddome, 1877</bibRefCitation>
;
<taxonomicName id="DD903BD2FFE15F77FAF853139FC5084F" authority="Deraniyagala, 1954" authorityName="Deraniyagala" authorityYear="1954" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="ruhunae">
<emphasis id="28E49C43FFE15F77FAF853139EFC084F" italics="true" pageId="0" pageNumber="592">Uropeltis ruhunae</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FF3753339FC5084F" author="Deraniyagala, P." box="[247,457,1079,1102]" pageId="0" pageNumber="592" pagination="27 - 28" refId="ref2973" refString="Deraniyagala, P. (1954) Two new snakes from Ceylon. Proceedings of the Tenth Annual Session of the Ceylon Association for the Advancement of Science (25 th- 27 th Nov. 1954), 1, 27 - 28 [24]." type="journal article" year="1954">Deraniyagala, 1954</bibRefCitation>
</taxonomicName>
was still considered valid (see
<bibRefCitation id="7E013DA0FFE15F77FCDD53339DF8084F" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[797,1012,1079,1102]" pageId="0" pageNumber="592" refId="ref3877" refString="Wallach, V., Williams, K. L. &amp; Boundy, J. (2014) Snakes of the world: a catalogue of living and extinct species. CRC Press, Boca Raton, 1209 pp. https: // doi. org / 10.1201 / b 16901" type="book" year="2014">
Wallach
<emphasis id="28E49C43FFE15F77FCBC53339DBF084F" box="[892,947,1079,1102]" italics="true" pageId="0" pageNumber="592">et al.</emphasis>
2014
</bibRefCitation>
).
<bibRefCitation id="7E013DA0FFE15F77FBC653339AC1084F" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[1030,1229,1079,1102]" pageId="0" pageNumber="592" pagination="453 - 506" refId="ref3447" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V. &amp; Somaweera, R. (2016) A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae). Zoosystema, 38, 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2" type="journal article" year="2016">
Pyron
<emphasis id="28E49C43FFE15F77FB8E53339A72084F" box="[1102,1150,1079,1102]" italics="true" pageId="0" pageNumber="592">et al</emphasis>
. (2016
</bibRefCitation>
:464) compared the scalation of the
<typeStatus id="C52BFEF3FFE15F77FE8453539FAD086F" box="[324,417,1111,1134]" pageId="0" pageNumber="592" type="holotype">holotype</typeStatus>
(NMSL R.S. 52) of
<taxonomicName id="DD903BD2FFE15F77FD4353539A2D086F" authority="Deraniyagala, 1954" authorityName="Deraniyagala" authorityYear="1954" box="[643,1057,1111,1134]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="ruhunae">
<emphasis id="28E49C43FFE15F77FD4353539D4B086F" box="[643,839,1111,1134]" italics="true" pageId="0" pageNumber="592">Uropeltis ruhunae</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FC8F53539A2D086F" author="Deraniyagala, P." box="[847,1057,1111,1134]" pageId="0" pageNumber="592" pagination="27 - 28" refId="ref2973" refString="Deraniyagala, P. (1954) Two new snakes from Ceylon. Proceedings of the Tenth Annual Session of the Ceylon Association for the Advancement of Science (25 th- 27 th Nov. 1954), 1, 27 - 28 [24]." type="journal article" year="1954">Deraniyagala, 1954</bibRefCitation>
</taxonomicName>
to known populations of
<taxonomicName id="DD903BD2FFE15F77FAF853539FD3088F" authority="(Theobald, 1876)" authorityName="Theobald" authorityYear="1876" baseAuthorityName="Theobald" baseAuthorityYear="1876" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="woodmasoni">
<emphasis id="28E49C43FFE15F77FAF853539F12088F" italics="true" pageId="0" pageNumber="592">Uropeltis woodmasoni</emphasis>
(
<bibRefCitation id="7E013DA0FFE15F77FEEB53739FD4088F" author="Theobald, W." box="[299,472,1143,1166]" pageId="0" pageNumber="592" refId="ref3765" refString="Theobald, W. (1876) Descriptive catalogue of the reptiles of British India. Thacker, Spink &amp; Co., Calcutta, 238 pp. https: // doi. org / 10.5962 / bhl. title. 54001" type="book" year="1876">Theobald, 1876</bibRefCitation>
)
</taxonomicName>
, concluding that it was a member of that species. Here, we present photographs of NMSL R.S. 52 (
<figureCitation id="82AB5CD4FFE15F77FF3A53939F3608AF" box="[250,314,1175,1198]" captionStart="FIGURE 1" captionStartId="2.[151,244,1974,1996]" captionTargetBox="[164,1428,226,1912]" captionTargetId="figure@2.[164,1428,226,1912]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal (a) and ventral (b) views of the holotype (NMSL R. S. 52) of Uropeltis ruhunae Deraniyagala, 1954. Major ticks are 1 cm." httpUri="https://zenodo.org/record/2627779/files/figure.png" pageId="0" pageNumber="592">Fig. 1</figureCitation>
), as well as photographs of two
<typeStatus id="C52BFEF3FFE15F77FD6653939D0F08AF" box="[678,771,1175,1198]" pageId="0" pageNumber="592" type="syntype">syntypes</typeStatus>
of
<emphasis id="28E49C43FFE15F77FCEC53939DC508AF" box="[812,969,1175,1198]" italics="true" pageId="0" pageNumber="592">Silybura nigra</emphasis>
<bibRefCitation id="7E013DA0FFE15F77FC1253939A7108AF" author="Beddome, R. H." box="[978,1149,1175,1198]" pageId="0" pageNumber="592" pagination="154 - 155" refId="ref2641" refString="Beddome, R. H. (1878) Descriptions of new Uropeltidae from Southern India, with remarks on some previously? described species. Proceedings of the Zoological Society of London, 46, 154 - 155. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07946. x" type="journal article" year="1878">Beddome, 1878</bibRefCitation>
(MNHN-RA-1895.85a–b;
<figureCitation id="82AB5CD4FFE15F77FF5753B39EEC08CF" box="[151,224,1207,1230]" captionStart="FIGURE 2" captionStartId="3.[151,244,1930,1952]" captionTargetBox="[215,1378,194,1903]" captionTargetId="figure@3.[210,1378,191,1909]" captionTargetPageId="3" captionText="FIGURE 2. Syntypes (MNHN-RA-1895.95a–b, the latter re-catalogued as MNHN-RA-2012.421) of Silybura nigra Beddome, 1878, a junior synonym of Uropeltis woodmasoni (Theobald, 1876). From top to bottom: a) &amp; b) dorsal and ventral views of MNHN-RA- 1895.95, c) &amp; d) dorsal and ventral views of MNHN-RA-2012.421. Red scale bar is 1 cm." httpUri="https://zenodo.org/record/2627781/files/figure.png" pageId="0" pageNumber="592">Figs. 2</figureCitation>
a–d), a junior synonym of
<taxonomicName id="DD903BD2FFE15F77FDC153B39DB408CF" authority="(Theobald, 1876)" authorityName="Theobald" authorityYear="1876" baseAuthorityName="Theobald" baseAuthorityYear="1876" box="[513,952,1207,1230]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="woodmasoni">
<emphasis id="28E49C43FFE15F77FDC153B39CFF08CF" box="[513,755,1207,1230]" italics="true" pageId="0" pageNumber="592">Uropeltis woodmasoni</emphasis>
(
<bibRefCitation id="7E013DA0FFE15F77FCC253B39DBC08CF" author="Theobald, W." box="[770,944,1207,1230]" pageId="0" pageNumber="592" refId="ref3765" refString="Theobald, W. (1876) Descriptive catalogue of the reptiles of British India. Thacker, Spink &amp; Co., Calcutta, 238 pp. https: // doi. org / 10.5962 / bhl. title. 54001" type="book" year="1876">Theobald, 1876</bibRefCitation>
)
</taxonomicName>
. In addition to the lepidosis data reported by
<bibRefCitation id="7E013DA0FFE15F77FF5753DC9F6308EE" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[151,367,1240,1263]" pageId="0" pageNumber="592" pagination="453 - 506" refId="ref3447" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V. &amp; Somaweera, R. (2016) A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae). Zoosystema, 38, 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2" type="journal article" year="2016">
Pyron
<emphasis id="28E49C43FFE15F77FF2153DC9F1808EE" box="[225,276,1240,1263]" italics="true" pageId="0" pageNumber="592">et al</emphasis>
. (2016)
</bibRefCitation>
, visual comparisons of the dorsal and ventral color-pattern further cement this association. The
<typeStatus id="C52BFEF3FFE15F77FF5753FC9EF50910" box="[151,249,1272,1297]" pageId="0" pageNumber="592" type="holotype">holotype</typeStatus>
(ZSI 8760) of
<taxonomicName id="DD903BD2FFE15F77FE6953FD9D7B0910" authority="(Theobald, 1876)" authorityName="Theobald" authorityYear="1876" baseAuthorityName="Theobald" baseAuthorityYear="1876" box="[425,887,1272,1297]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="woodmasoni">
<emphasis id="28E49C43FFE15F77FE6953FD9CA50910" box="[425,681,1273,1297]" italics="true" pageId="0" pageNumber="592">Uropeltis woodmasoni</emphasis>
(
<bibRefCitation id="7E013DA0FFE15F77FD7B53FC9D630910" author="Theobald, W." box="[699,879,1272,1297]" pageId="0" pageNumber="592" refId="ref3765" refString="Theobald, W. (1876) Descriptive catalogue of the reptiles of British India. Thacker, Spink &amp; Co., Calcutta, 238 pp. https: // doi. org / 10.5962 / bhl. title. 54001" type="book" year="1876">Theobald, 1876</bibRefCitation>
)
</taxonomicName>
is in relatively poor condition, but photographs provided by I. Das are essentially identical to the
<emphasis id="28E49C43FFE15F77FD0D521F9D050933" box="[717,777,1307,1330]" italics="true" pageId="0" pageNumber="592">nigra</emphasis>
and
<taxonomicName id="DD903BD2FFE15F77FC82521E9D930933" authorityName="Deraniyagala" authorityYear="1954" box="[834,927,1306,1330]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="0" pageNumber="592" phylum="Chordata" rank="species" species="ruhunae">
<emphasis id="28E49C43FFE15F77FC82521E9D930933" box="[834,927,1306,1330]" italics="true" pageId="0" pageNumber="592">ruhunae</emphasis>
</taxonomicName>
types in overall shape and remaining visible color-pattern. Specifically, hypertrophied anterior trunk musculature and “swollen” aspect of the first ~1/4 of body; irregular yellow stripe on the first several dorsal scale rows, beginning on posteriormost labials and extending ~1/4 the length of the body, dark brown dorsum with broken rings of yellow ocellations or speckles on the posteriormost ¾ of body, spaced 3-4 scale rows apart; and irregular yellow and brown zigzags or blotches on the venter, with yellow occasionally and irregularly extending on to the first few dorsal scale rows.
</paragraph>
</subSubSection>
</treatment>
</document>
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