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<document id="CF227F172701190FC45F5DADC862E64C" ID-CLB-Dataset="24411" ID-DOI="10.11646/zootaxa.4718.4.3" ID-GBIF-Dataset="c3fec010-6e0c-41e3-aece-3c6caccc951a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602674" ID-ZooBank="32C51811-97BA-40E4-B503-F25F8C4CF98A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578565448735" checkinUser="plazi" docAuthor="Lukashevich, Elena D. &amp; Blagoderov, Vladimir A." docDate="2020" docId="03C75260FF99FFA1FF26A0EB4D8DF837" docLanguage="en" docName="zootaxa.4718.4.3.pdf" docOrigin="Zootaxa 4718 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Collessomma Lukashevich &amp; Blagoderov 2020, gen. nov." docType="treatment" docVersion="9" lastPageNumber="484" masterDocId="FFFE2A18FF9AFFA2FFB1A1114926FF99" masterDocTitle="Review of Mesozoic Perissommatidae (Insecta: Diptera)" masterLastPageNumber="496" masterPageNumber="481" pageNumber="484" updateTime="1698776762553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D84A026BF9EA82928A75F554316F647E">Review of Mesozoic Perissommatidae (Insecta: Diptera)</mods:title>
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<treatment id="03C75260FF99FFA1FF26A0EB4D8DF837" ID-DOI="http://doi.org/10.5281/zenodo.5920770" ID-GBIF-Taxon="161671137" ID-Zenodo-Dep="5920770" LSID="urn:lsid:plazi:treatment:03C75260FF99FFA1FF26A0EB4D8DF837" httpUri="http://treatment.plazi.org/id/03C75260FF99FFA1FF26A0EB4D8DF837" lastPageNumber="484" pageId="3" pageNumber="484">
<subSubSection id="C374B0FDFF99FFA1FF26A0EB4888FD8D" box="[151,430,506,533]" pageId="3" pageNumber="484" type="nomenclature">
<paragraph id="8BD1E376FF99FFA1FF26A0EB4888FD8D" blockId="3.[151,983,506,598]" box="[151,430,506,533]" pageId="3" pageNumber="484">
<heading id="D099541AFF99FFA1FF26A0EB4888FD8D" bold="true" box="[151,430,506,533]" fontSize="11" level="1" pageId="3" pageNumber="484" reason="1">
<emphasis id="B91A3F64FF99FFA1FF26A0EB4888FD8D" bold="true" box="[151,430,506,533]" pageId="3" pageNumber="484">
<taxonomicName id="4C6E98F5FF99FFA1FF26A0EB481EFD8C" ID-CoL="89PXL" authority="Lukashevich &amp; Blagoderov, 2020" authorityName="Lukashevich &amp; Blagoderov" authorityYear="2020" box="[151,312,506,533]" class="Insecta" family="Perissommatidae" genus="Collessomma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91A3F64FF99FFA1FF26A0EB481EFD8C" bold="true" box="[151,312,506,533]" italics="true" pageId="3" pageNumber="484">Collessomma</emphasis>
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<taxonomicNameLabel id="A229821FFF99FFA1FEF1A0EB4888FD8D" box="[320,430,506,532]" pageId="3" pageNumber="484" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="C374B0FDFF99FFA1FF26A30F4AF1FDCF" pageId="3" pageNumber="484" type="description">
<paragraph id="8BD1E376FF99FFA1FF26A30F4809FDA0" blockId="3.[151,983,506,598]" box="[151,303,542,569]" pageId="3" pageNumber="484">
(
<figureCitation id="1355FFF3FF99FFA1FF2EA30F4801FDA0" box="[159,295,542,569]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="4.[151,250,1914,1939]" captionStartId-1="5.[151,250,1898,1923]" captionStartId-2="6.[151,250,1841,1866]" captionStartId-3="8.[151,250,1924,1949]" captionStartId-4="9.[151,250,1834,1859]" captionTargetBox-0="[151,1436,174,1875]" captionTargetBox-1="[214,1373,174,1878]" captionTargetBox-2="[201,1415,290,1703]" captionTargetBox-3="[203,1385,166,1904]" captionTargetBox-4="[151,1436,439,1812]" captionTargetId-0="figure@4.[151,1436,174,1875]" captionTargetId-1="figure@5.[214,1373,174,1878]" captionTargetId-2="figure@6.[153,1433,249,1731]" captionTargetId-3="figure@8.[203,1385,166,1904]" captionTargetId-4="figure@9.[151,1436,439,1815]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." captionText-1="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." captionText-2="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." captionText-3="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." captionText-4="FIGURE 6. Collessomma mongolica gen. et sp. nov. (Bon-Tsagan, K1). A, B. Holotype PIN No. 3559/10039, total view and head. C. Paratype PIN No. 3559/4154, wing. Scale bars 500 μm (A, C), 200 μm (B)." figureDoi-0="http://doi.org/10.5281/zenodo.3602678" figureDoi-1="http://doi.org/10.5281/zenodo.3602680" figureDoi-2="http://doi.org/10.5281/zenodo.3602682" figureDoi-3="http://doi.org/10.5281/zenodo.3602684" figureDoi-4="http://doi.org/10.5281/zenodo.3602686" httpUri-0="https://zenodo.org/record/3602678/files/figure.png" httpUri-1="https://zenodo.org/record/3602680/files/figure.png" httpUri-2="https://zenodo.org/record/3602682/files/figure.png" httpUri-3="https://zenodo.org/record/3602684/files/figure.png" httpUri-4="https://zenodo.org/record/3602686/files/figure.png" pageId="3" pageNumber="484">Figures 26</figureCitation>
)
</paragraph>
<paragraph id="8BD1E376FF99FFA1FF26A32A4AF1FDCF" blockId="3.[151,983,506,598]" box="[151,983,571,598]" pageId="3" pageNumber="484">
<uri id="FFFFEF74FF99FFA1FF26A32A4AF1FDCF" box="[151,983,571,598]" pageId="3" pageNumber="484">
urn:lsid:zoobank.org:act:
<uuid id="FFC8D9A3FF99FFA1FE16A32A4AF1FDCF" box="[423,983,571,598]" pageId="3" pageNumber="484">
E4
<collectingCountry id="F379A3E6FF99FFA1FE76A32A4B31FDCF" box="[455,535,571,598]" name="Chad" pageId="3" pageNumber="484">CEAD</collectingCountry>
3D-3FA8-4B7A-B54F-E661FEFB6995
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</subSubSection>
<subSubSection id="C374B0FDFF99FFA1FF26A3954BAEFD07" box="[151,648,643,670]" pageId="3" pageNumber="484" type="type_taxon">
<paragraph id="8BD1E376FF99FFA1FF26A3954BAEFD07" blockId="3.[151,1437,643,1966]" box="[151,648,643,670]" pageId="3" pageNumber="484">
<emphasis id="B91A3F64FF99FFA1FF26A395480DFD07" bold="true" box="[151,299,643,670]" pageId="3" pageNumber="484">
<typeStatus id="54D55DD4FF99FFA1FF26A39549F4FD07" box="[151,210,644,670]" pageId="3" pageNumber="484">Type</typeStatus>
species
</emphasis>
<taxonomicName id="4C6E98F5FF99FFA1FE83A3924B00FD04" ID-CoL="8BJRH" authority="Lukashevich &amp; Blagoderov, 2020" authorityName="Lukashevich &amp; Blagoderov" authorityYear="2020" box="[306,550,643,669]" class="Insecta" family="Perissommatidae" genus="Collessomma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="species" species="sibirica" status="sp. nov.">
<emphasis id="B91A3F64FF99FFA1FE83A3924B00FD04" box="[306,550,643,669]" italics="true" pageId="3" pageNumber="484">Collessomma sibirica</emphasis>
</taxonomicName>
<emphasis id="B91A3F64FF99FFA1FD9CA3954BAEFD07" bold="true" box="[557,648,644,670]" pageId="3" pageNumber="484">
<taxonomicNameLabel id="A229821FFF99FFA1FD9CA3954BAEFD07" box="[557,648,644,670]" pageId="3" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C374B0FDFF99FFA1FF76A3B94CA3FD7C" pageId="3" pageNumber="484" type="etymology">
<paragraph id="8BD1E376FF99FFA1FF76A3B94CA3FD7C" blockId="3.[151,1437,643,1966]" pageId="3" pageNumber="484">
<emphasis id="B91A3F64FF99FFA1FF76A3B94868FD5B" bold="true" box="[199,334,680,706]" pageId="3" pageNumber="484">Etymology.</emphasis>
The generic epithet refers to Donald Henry (Don) Colless, an outstanding expert on Australian
<taxonomicName id="4C6E98F5FF99FFA1FF26A3DA49C8FD7C" baseAuthorityName="Cumming &amp; Wood" baseAuthorityYear="2017" box="[151,238,715,741]" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="484" rank="order">Diptera</taxonomicName>
, who established the family
<taxonomicName id="4C6E98F5FF99FFA1FD9CA3DA4BC9FD7C" authorityName="Colless" authorityYear="1962" box="[557,751,715,741]" class="Insecta" family="Perissommatidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="family">Perissommatidae</taxonomicName>
, and to the Greek term
<emphasis id="B91A3F64FF99FFA1FC44A3DD4D1FFD7C" box="[1013,1081,716,741]" italics="true" pageId="3" pageNumber="484">omma</emphasis>
—eye. The name is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C374B0FDFF99FFA1FF76A3FE4DEEFCB7" pageId="3" pageNumber="484" type="diagnosis">
<paragraph id="8BD1E376FF99FFA1FF76A3FE4DEEFCB7" blockId="3.[151,1437,643,1966]" pageId="3" pageNumber="484">
<emphasis id="B91A3F64FF99FFA1FF76A3FE4864FC93" bold="true" box="[199,322,751,778]" pageId="3" pageNumber="484">Diagnosis.</emphasis>
Eyes in male holoptic, with facets of different size, veins with macrotrichia, dense on radial ones except for R
<subScript id="17EAE133FF99FFA1FE93A234480CFCAA" attach="left" box="[290,298,805,819]" fontSize="6" pageId="3" pageNumber="484">5</subScript>
, discal cell elongate, 1/4 wing length, CuP ending at level
<emphasis id="B91A3F64FF99FFA1FC0DA2054AD2FCB4" box="[956,1012,788,813]" italics="true" pageId="3" pageNumber="484">m-cu</emphasis>
, anal lobe present.
</paragraph>
</subSubSection>
<subSubSection id="C374B0FDFF99FFA1FF76A2264A95FA08" pageId="3" pageNumber="484" type="description">
<paragraph id="8BD1E376FF99FFA1FF76A226489FFAD3" blockId="3.[151,1437,643,1966]" pageId="3" pageNumber="484">
<emphasis id="B91A3F64FF99FFA1FF76A226487FFCCB" bold="true" box="[199,345,823,850]" pageId="3" pageNumber="484">Description.</emphasis>
Antenna moniliform, rather short and stout, with 11 segments, flagellum not clearly differentiated, segments 2 and 3 (pedicel and first flagellomere) enlarged, all segments with fine setae only (
<figureCitation id="1355FFF3FF99FFA1FAB9A24A4C43FCEF" box="[1288,1381,859,886]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2C</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FAC2A24A49EDFC03" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5A, B, F</figureCitation>
). Ocelli probably absent. Eyes in male holoptic, with facets of different size, without disconnected zone (
<figureCitation id="1355FFF3FF99FFA1FADAA26E49DAFC27" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2C, D, F</figureCitation>
;
<figureCitation id="1355FFF3FF99FFA1FEB9A2B2486BFC27" box="[264,333,931,958]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5A, B</figureCitation>
); in female dichopic, with facets of the same size (
<figureCitation id="1355FFF3FF99FFA1FC3BA2B24AC5FC27" box="[906,995,931,958]" captionStart="FIGURE 6" captionStartId="9.[151,250,1834,1859]" captionTargetBox="[151,1436,439,1812]" captionTargetId="figure@9.[151,1436,439,1815]" captionTargetPageId="9" captionText="FIGURE 6. Collessomma mongolica gen. et sp. nov. (Bon-Tsagan, K1). A, B. Holotype PIN No. 3559/10039, total view and head. C. Paratype PIN No. 3559/4154, wing. Scale bars 500 μm (A, C), 200 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602686" httpUri="https://zenodo.org/record/3602686/files/figure.png" pageId="3" pageNumber="484">Fig. 6B</figureCitation>
). Mouthparts very short (
<figureCitation id="1355FFF3FF99FFA1FAB1A2B24C78FC27" box="[1280,1374,931,958]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2D</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FADBA2B24CB6FC27" box="[1386,1424,931,958]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5A</figureCitation>
). Mesonotum arched anterodorsally, scutellum separated from postscutum by distinct suture, metanotum very large (
<figureCitation id="1355FFF3FF99FFA1FF2EA2FA49D9FB9F" box="[159,255,1003,1030]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2A</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FEBFA2FD4814FB9F" box="[270,306,1004,1030]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[214,1373,174,1878]" captionTargetId="figure@5.[214,1373,174,1878]" captionTargetPageId="5" captionText="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602680" httpUri="https://zenodo.org/record/3602680/files/figure.png" pageId="3" pageNumber="484">3A</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FEF1A2FA4841FB9F" box="[320,359,1003,1030]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5H</figureCitation>
). Legs slender, with small tibial spurs, femur slightly expanded, first tarsomere longest, forth tarsomere shortest, empodium and pulvilli well-developed, narrow; pubescence of empodium and pulvilli invisible (
<figureCitation id="1355FFF3FF99FFA1FF2EA522483AFBD7" box="[159,284,1075,1102]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[214,1373,174,1878]" captionTargetId="figure@5.[214,1373,174,1878]" captionTargetPageId="5" captionText="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602680" httpUri="https://zenodo.org/record/3602680/files/figure.png" pageId="3" pageNumber="484">Figs 3A, B</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FE96A525486CFBD7" box="[295,330,1076,1102]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4B</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FEE7A522485CFBD7" box="[342,378,1075,1102]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5C</figureCitation>
). Wing relatively broad, longer than abdomen, with anal lobe but without alula, with numerous macrotrichia on pterostigma and on veins, denser on radial veins (
<figureCitation id="1355FFF3FF99FFA1FC36A5464D20FBEB" box="[903,1030,1111,1138]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2A, E</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FBA2A5494D7CFBEB" box="[1043,1114,1112,1138]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[214,1373,174,1878]" captionTargetId="figure@5.[214,1373,174,1878]" captionTargetPageId="5" captionText="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602680" httpUri="https://zenodo.org/record/3602680/files/figure.png" pageId="3" pageNumber="484">3A, C</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FBD6A5494DA3FBEB" box="[1127,1157,1112,1138]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4F</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FB23A5464DFCFBEB" box="[1170,1242,1111,1138]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5D, H</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FB56A5464C09FBEB" box="[1255,1327,1111,1138]" captionStart="FIGURE 6" captionStartId="9.[151,250,1834,1859]" captionTargetBox="[151,1436,439,1812]" captionTargetId="figure@9.[151,1436,439,1815]" captionTargetPageId="9" captionText="FIGURE 6. Collessomma mongolica gen. et sp. nov. (Bon-Tsagan, K1). A, B. Holotype PIN No. 3559/10039, total view and head. C. Paratype PIN No. 3559/4154, wing. Scale bars 500 μm (A, C), 200 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602686" httpUri="https://zenodo.org/record/3602686/files/figure.png" pageId="3" pageNumber="484">6A, C</figureCitation>
). R
<subScript id="17EAE133FF99FFA1FAE6A5784C79FBEE" attach="left" box="[1367,1375,1129,1143]" fontSize="6" pageId="3" pageNumber="484">1</subScript>
apex expanded; fork R
<subScript id="17EAE133FF99FFA1FEEBA59C4852FB02" attach="left" box="[346,372,1165,1179]" fontSize="6" pageId="3" pageNumber="484">3+4</subScript>
forms relatively acute angle, about 45°; crossvein
<emphasis id="B91A3F64FF99FFA1FC18A56D4AF7FB0C" box="[937,977,1148,1173]" italics="true" pageId="3" pageNumber="484">r-m</emphasis>
proximal to
<emphasis id="B91A3F64FF99FFA1FBD0A56D4D5BFB0C" box="[1121,1149,1148,1173]" italics="true" pageId="3" pageNumber="484">im</emphasis>
and distal to
<emphasis id="B91A3F64FF99FFA1FAA5A56D4C6AFB0C" box="[1300,1356,1148,1173]" italics="true" pageId="3" pageNumber="484">m-cu</emphasis>
; discal cell elongate, 1/4 wing length, CuP ending at level
<emphasis id="B91A3F64FF99FFA1FD64A5B14A2BFB20" box="[725,781,1184,1209]" italics="true" pageId="3" pageNumber="484">m-cu</emphasis>
(
<figureCitation id="1355FFF3FF99FFA1FCADA58E4ABFFB23" box="[796,921,1183,1210]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2A, E</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FC14A5B14ACDFB23" box="[933,1003,1184,1210]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[214,1373,174,1878]" captionTargetId="figure@5.[214,1373,174,1878]" captionTargetPageId="5" captionText="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602680" httpUri="https://zenodo.org/record/3602680/files/figure.png" pageId="3" pageNumber="484">3A, C</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FC46A5B14D7CFB23" box="[1015,1114,1184,1210]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4A, D, F</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FBD4A58E4DACFB23" box="[1125,1162,1183,1210]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5H</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FB27A58E4D9CFB23" box="[1174,1210,1183,1210]" captionStart="FIGURE 6" captionStartId="9.[151,250,1834,1859]" captionTargetBox="[151,1436,439,1812]" captionTargetId="figure@9.[151,1436,439,1815]" captionTargetPageId="9" captionText="FIGURE 6. Collessomma mongolica gen. et sp. nov. (Bon-Tsagan, K1). A, B. Holotype PIN No. 3559/10039, total view and head. C. Paratype PIN No. 3559/4154, wing. Scale bars 500 μm (A, C), 200 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602686" httpUri="https://zenodo.org/record/3602686/files/figure.png" pageId="3" pageNumber="484">6C</figureCitation>
). Male hypopygium narrower than VIII abdominal segment and relatively simple, not rotated, gonocoxites elongate and massive, gonostyli simple, stout; ninth tergite prominent (
<figureCitation id="1355FFF3FF99FFA1FDC9A5F64BF2FA9B" box="[632,724,1255,1282]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2B</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FD6FA5F94A39FA9B" box="[734,799,1256,1282]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4C, E</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FC9AA5F64A49FA9B" box="[811,879,1255,1282]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5D, E</figureCitation>
). Female terminalia with large, prominent bilobed eighth sternite, cerci prominent, well chitinized but not horny, and with numerous setae; three sclerotized spermathecae (
<figureCitation id="1355FFF3FF99FFA1FF5FA43E486DFAD3" box="[238,331,1327,1354]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">Figs 4G</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FEE7A43E485DFAD3" box="[342,379,1327,1354]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5G</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FE36A43E488AFAD3" box="[391,428,1327,1354]" captionStart="FIGURE 6" captionStartId="9.[151,250,1834,1859]" captionTargetBox="[151,1436,439,1812]" captionTargetId="figure@9.[151,1436,439,1815]" captionTargetPageId="9" captionText="FIGURE 6. Collessomma mongolica gen. et sp. nov. (Bon-Tsagan, K1). A, B. Holotype PIN No. 3559/10039, total view and head. C. Paratype PIN No. 3559/4154, wing. Scale bars 500 μm (A, C), 200 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602686" httpUri="https://zenodo.org/record/3602686/files/figure.png" pageId="3" pageNumber="484">6A</figureCitation>
).
</paragraph>
<paragraph id="8BD1E376FF99FFA1FF76A4424A95FA08" blockId="3.[151,1437,643,1966]" pageId="3" pageNumber="484">
<emphasis id="B91A3F64FF99FFA1FF76A44248B3FAF7" bold="true" box="[199,405,1363,1390]" pageId="3" pageNumber="484">Species included.</emphasis>
Besides
<typeStatus id="54D55DD4FF99FFA1FE4CA4454B0BFAF7" box="[509,557,1364,1390]" pageId="3" pageNumber="484">type</typeStatus>
species,
<taxonomicName id="4C6E98F5FF99FFA1FD26A4454A23FAF4" authority="Lukashevich &amp; Blagoderov, 2020" authorityName="Lukashevich &amp; Blagoderov" authorityYear="2020" box="[663,773,1364,1390]" class="Insecta" family="Perissommatidae" genus="Collessomma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="species" species="gnoma" status="gen. et sp. nov.">
<emphasis id="B91A3F64FF99FFA1FD26A4454A23FAF4" box="[663,773,1364,1390]" italics="true" pageId="3" pageNumber="484">C. gnoma</emphasis>
</taxonomicName>
<emphasis id="B91A3F64FF99FFA1FCBFA4454AE7FAF7" bold="true" box="[782,961,1364,1390]" pageId="3" pageNumber="484">
<taxonomicNameLabel id="A229821FFF99FFA1FCBFA4454AE7FAF7" box="[782,961,1364,1390]" pageId="3" pageNumber="484" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
from the same locality and
<taxonomicName id="4C6E98F5FF99FFA1FAB5A4454CBAFAF4" authority="Lukashevich &amp; Blagoderov, 2020" authorityName="Lukashevich &amp; Blagoderov" authorityYear="2020" box="[1284,1436,1363,1390]" class="Insecta" family="Perissommatidae" genus="Collessomma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="species" species="mongolica" status="gen. et sp. nov.">
<emphasis id="B91A3F64FF99FFA1FAB5A4454CBAFAF4" box="[1284,1436,1363,1390]" italics="true" pageId="3" pageNumber="484">C. mongolica</emphasis>
</taxonomicName>
<emphasis id="B91A3F64FF99FFA1FF26A4694863FA0B" bold="true" box="[151,325,1400,1426]" pageId="3" pageNumber="484">
<taxonomicNameLabel id="A229821FFF99FFA1FF26A4694863FA0B" box="[151,325,1400,1426]" pageId="3" pageNumber="484" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
from Lower Cretaceous of
<collectingCountry id="F379A3E6FF99FFA1FDCDA4664BCAFA08" box="[636,748,1399,1425]" name="Mongolia" pageId="3" pageNumber="484">Mongolia</collectingCountry>
, described below.
</paragraph>
</subSubSection>
<subSubSection id="C374B0FDFF99FFA1FF76A48D4D8DF837" pageId="3" pageNumber="484" type="discussion">
<paragraph id="8BD1E376FF99FFA1FF76A48D48B1F92B" blockId="3.[151,1437,643,1966]" pageId="3" pageNumber="484">
<emphasis id="B91A3F64FF99FFA1FF76A48D481DFA2F" bold="true" box="[199,315,1436,1462]" pageId="3" pageNumber="484">Remarks.</emphasis>
Characters typical of the family are observed in the wing venation of the new genus: the costal vein extends along the entire wing, strongly thickened at the anterior and distal margins, weak but distinct along the posterior margin; the peculiar position of the pterostigma between R
<subScript id="17EAE133FF99FFA1FC2EA4E44A81F99A" attach="left" box="[927,935,1525,1539]" fontSize="6" pageId="3" pageNumber="484">1</subScript>
and R
<subScript id="17EAE133FF99FFA1FC43A4E44ADCF99A" attach="left" box="[1010,1018,1525,1539]" fontSize="6" pageId="3" pageNumber="484">3</subScript>
; a forked anterior branch of Rs, R
<subScript id="17EAE133FF99FFA1FA32A4E44CBBF99A" attach="left" box="[1411,1437,1525,1539]" fontSize="6" pageId="3" pageNumber="484">3+4</subScript>
combined with a simple posterior branch R
<subScript id="17EAE133FF99FFA1FDC4A7084B5BF9BE" attach="left" box="[629,637,1561,1575]" fontSize="6" pageId="3" pageNumber="484">5</subScript>
;
<emphasis id="B91A3F64FF99FFA1FD3AA7194B9EF9B8" box="[651,696,1544,1569]" italics="true" pageId="3" pageNumber="484">rm</emphasis>
connects R
<subScript id="17EAE133FF99FFA1FC88A7084A67F9BE" attach="left" box="[825,833,1561,1575]" fontSize="6" pageId="3" pageNumber="484">5</subScript>
and M
<subScript id="17EAE133FF99FFA1FC3EA7084A8EF9BE" attach="left" box="[911,936,1561,1575]" fontSize="6" pageId="3" pageNumber="484">1+2</subScript>
; M stem very weak, fold-like, M
<subScript id="17EAE133FF99FFA1FAA7A7084C16F9BE" attach="left" box="[1302,1328,1561,1575]" fontSize="6" pageId="3" pageNumber="484">1+2</subScript>
is forked, М
<subScript id="17EAE133FF99FFA1FF01A72C49ECF9D2" attach="left" box="[176,202,1597,1611]" fontSize="6" pageId="3" pageNumber="484">3+4</subScript>
is simple;
<emphasis id="B91A3F64FF99FFA1FEF2A73D48A6F9DC" box="[323,384,1580,1605]" italics="true" pageId="3" pageNumber="484">mcu</emphasis>
is positioned in the proximal part of the wing; CuP is distally reduced (
<figureCitation id="1355FFF3FF99FFA1FB2AA73A4C31F9DF" box="[1179,1303,1579,1606]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2A, E</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FA92A73D4C4EF9DF" box="[1315,1384,1580,1606]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[214,1373,174,1878]" captionTargetId="figure@5.[214,1373,174,1878]" captionTargetPageId="5" captionText="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602680" httpUri="https://zenodo.org/record/3602680/files/figure.png" pageId="3" pageNumber="484">3A, C</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FAC2A73D4CBEF9DF" box="[1395,1432,1580,1606]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4A</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FF26A75E499AF9F3" box="[151,188,1615,1642]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">5H</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FF76A75E49CDF9F3" box="[199,235,1615,1642]" captionStart="FIGURE 6" captionStartId="9.[151,250,1834,1859]" captionTargetBox="[151,1436,439,1812]" captionTargetId="figure@9.[151,1436,439,1815]" captionTargetPageId="9" captionText="FIGURE 6. Collessomma mongolica gen. et sp. nov. (Bon-Tsagan, K1). A, B. Holotype PIN No. 3559/10039, total view and head. C. Paratype PIN No. 3559/4154, wing. Scale bars 500 μm (A, C), 200 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602686" httpUri="https://zenodo.org/record/3602686/files/figure.png" pageId="3" pageNumber="484">6C</figureCitation>
). As all members of subfamily
<taxonomicName id="4C6E98F5FF99FFA1FDF8A75E4A2EF9F0" authorityName="Colless" authorityYear="1962" box="[585,776,1615,1641]" class="Insecta" family="Perissommatidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="subFamily" subFamily="Perissommatinae">Perissommatinae</taxonomicName>
<taxonomicName id="4C6E98F5FF99FFA1FCBEA75E4A80F9F0" authority="Lukashevich &amp; Blagoderov, 2020" authorityName="Lukashevich &amp; Blagoderov" authorityYear="2020" box="[783,934,1615,1641]" class="Insecta" family="Perissommatidae" genus="Collessomma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91A3F64FF99FFA1FCBEA75E4A80F9F0" box="[783,934,1615,1641]" italics="true" pageId="3" pageNumber="484">Collessomma</emphasis>
</taxonomicName>
<emphasis id="B91A3F64FF99FFA1FC1CA7414D30F9F3" bold="true" box="[941,1046,1616,1642]" pageId="3" pageNumber="484">
<taxonomicNameLabel id="A229821FFF99FFA1FC1CA7414D30F9F3" box="[941,1046,1616,1642]" pageId="3" pageNumber="484" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
has short Sc not ending at C, short R
<subScript id="17EAE133FF99FFA1FF1BA7944994F90A" attach="left" box="[170,178,1669,1683]" fontSize="6" pageId="3" pageNumber="484">1</subScript>
(about half of the wing length); R
<subScript id="17EAE133FF99FFA1FD9FA7944B10F90A" attach="left" box="[558,566,1669,1683]" fontSize="6" pageId="3" pageNumber="484">3</subScript>
distally closer to R
<subScript id="17EAE133FF99FFA1FCBFA7944A30F90A" attach="left" box="[782,790,1669,1683]" fontSize="6" pageId="3" pageNumber="484">4</subScript>
than to R
<subScript id="17EAE133FF99FFA1FC32A7944AADF90A" attach="left" box="[899,907,1669,1683]" fontSize="6" pageId="3" pageNumber="484">1</subScript>
;
<emphasis id="B91A3F64FF99FFA1FC28A7654AE7F914" box="[921,961,1652,1677]" italics="true" pageId="3" pageNumber="484">r-m</emphasis>
in proximal half of wing and
<emphasis id="B91A3F64FF99FFA1FABCA7654C6CF914" box="[1293,1354,1652,1677]" italics="true" pageId="3" pageNumber="484">mcu</emphasis>
ending on M
<subScript id="17EAE133FF99FFA1FF62A7B849CBF92E" attach="left" box="[211,237,1705,1719]" fontSize="6" pageId="3" pageNumber="484">3+4</subScript>
(
<figureCitation id="1355FFF3FF99FFA1FF4AA786487EF92B" box="[251,344,1687,1714]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[214,1373,174,1878]" captionTargetId="figure@5.[214,1373,174,1878]" captionTargetPageId="5" captionText="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602680" httpUri="https://zenodo.org/record/3602680/files/figure.png" pageId="3" pageNumber="484">Figs 3A</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FED5A78948ADF92B" box="[356,395,1688,1714]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4A</figureCitation>
).
</paragraph>
<paragraph id="8BD1E376FF99FFA1FF76A7AA4D8DF837" blockId="3.[151,1437,643,1966]" pageId="3" pageNumber="484">
Tibial spurs are found in the new genus but they are unknown in the
<typeStatus id="54D55DD4FF99FFA1FC03A7AD4AC4F94F" box="[946,994,1724,1750]" pageId="3" pageNumber="484">type</typeStatus>
genus
<taxonomicName id="4C6E98F5FF99FFA1FB9EA7AD4D9FF94C" authorityName="Colless" authorityYear="1962" box="[1071,1209,1724,1749]" class="Insecta" family="Perissommatidae" genus="Perissomma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B91A3F64FF99FFA1FB9EA7AD4D9FF94C" box="[1071,1209,1724,1749]" italics="true" pageId="3" pageNumber="484">Perissomma</emphasis>
</taxonomicName>
. In extant
<taxonomicName id="4C6E98F5FF99FFA1FA9DA7AA49D6F960" authorityName="Colless" authorityYear="1962" class="Insecta" family="Perissommatidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="family">Perissommatidae</taxonomicName>
“V-shaped suture of the tipuloid
<typeStatus id="54D55DD4FF99FFA1FDD6A7F14BBCF963" box="[615,666,1760,1786]" pageId="3" pageNumber="484">type</typeStatus>
” present, in the new genus transverse suture on scutum present with certainty, but its shape is unclear. However, some other familial characters (coinciding with that of
<taxonomicName id="4C6E98F5FF99FFA1FB51A6154C4CF884" authorityName="Colless" authorityYear="1962" box="[1248,1386,1796,1821]" class="Insecta" family="Perissommatidae" genus="Perissomma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="484" phylum="Arthropoda" rank="genus">
<emphasis id="B91A3F64FF99FFA1FB51A6154C4CF884" box="[1248,1386,1796,1821]" italics="true" pageId="3" pageNumber="484">Perissomma</emphasis>
</taxonomicName>
) are observed in body structures: antenna moniliform, with 11 segments, flagellum not clearly differentiated, pedicel and first flagellomere enlarged (
<figureCitation id="1355FFF3FF99FFA1FE7DA65A4B05F8FF" box="[460,547,1867,1894]" captionStart="FIGURE 5" captionStartId="8.[151,250,1924,1949]" captionTargetBox="[203,1385,166,1904]" captionTargetId="figure@8.[203,1385,166,1904]" captionTargetPageId="8" captionText="FIGURE 5. C.gnoma gen. et sp. nov. AC. Paratype PIN No. 5340/2982, male. A, B. Ventral and dorsal view of head (part and counterpart). C. Leg. D, E. Holotype PIN No. 5026/201, male, total view and terminalia. FH. Paratype PIN No. 5340/2981, female. F. Antenna. G. Cerci. H. Total view. Scale bars 200 μm (A, B, C, E, F, G), 1 mm (D, H)." figureDoi="http://doi.org/10.5281/zenodo.3602684" httpUri="https://zenodo.org/record/3602684/files/figure.png" pageId="3" pageNumber="484">Fig. 5B</figureCitation>
), thorax with large metanotum; halters long-stemmed; legs with well-developed empodium and pulvilli (
<figureCitation id="1355FFF3FF99FFA1FE1BA67E4B0DF813" box="[426,555,1903,1930]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[214,1373,174,1878]" captionTargetId="figure@5.[214,1373,174,1878]" captionTargetPageId="5" captionText="FIGURE 3. Collessomma sibirica gen. et sp. nov. Paratype PIN No. 5026/206,?male.A. Total view. B. Tarsus. C. Anterior part of wing with pterostigma.Abbreviations: CuA, anterior branch of cubital vein; CuP, posterior branch of cubital vein; im, medial crossvein; M1, M2, M3+4, branches of media; m-cu, mediacubital crossvein; R1, R3, R4, R5, branches of radius; r-m, radialmedial crossvein; Sc, subcostal vein. Scale bars 1 mm (A), 500 μm (C), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602680" httpUri="https://zenodo.org/record/3602680/files/figure.png" pageId="3" pageNumber="484">Figs 3A, B</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FD89A6614B7AF813" box="[568,604,1904,1930]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4B</figureCitation>
); well-developed eighth abdominal segment; male hypopygium narrower than VIII abdominal segment and relatively simple, ninth tergite prominent (
<figureCitation id="1355FFF3FF99FFA1FC41A6824D48F837" box="[1008,1134,1939,1966]" captionStart="FIGURE 2" captionStartId="4.[151,250,1914,1939]" captionTargetBox="[151,1436,174,1875]" captionTargetId="figure@4.[151,1436,174,1875]" captionTargetPageId="4" captionText="FIGURE 2. Male of Collessomma sibirica gen. et sp. nov. AE. holotype PIN No. 5340/1681. A. Total view. B. Terminalia. C, D. Head, dorsal and ventral view. E.Anterior part of wing with pterostigma. F. PIN No. 5026/1558, head with facets of different size. Scale bars 1 mm (A), 200 μm (BF)." figureDoi="http://doi.org/10.5281/zenodo.3602678" httpUri="https://zenodo.org/record/3602678/files/figure.png" pageId="3" pageNumber="484">Figs 2A, B</figureCitation>
,
<figureCitation id="1355FFF3FF99FFA1FBCBA6854DB8F837" box="[1146,1182,1940,1966]" captionStart="FIGURE 4" captionStartId="6.[151,250,1841,1866]" captionTargetBox="[201,1415,290,1703]" captionTargetId="figure@6.[153,1433,249,1731]" captionTargetPageId="6" captionText="FIGURE 4. Collessomma gen. nov. AC. C. sibirica gen. et sp. nov. A, B. Male? PIN No. 5026/206, wing and tarsus. C. Holotype PIN No. 5340/1681, male terminalia. D, E. C. gnoma gen. et sp. nov. Holotype PIN No. 5026/201, wing and male terminalia. F, G. C. mongolica gen. et sp. nov. F. Paratype PIN No. 3559/4154, wing. G. Holotype PIN No. 3559/10039, female terminalia.Abbreviations: aed, aedeagus; cerc, cerci; gst, gonostylus; gx, gonocoxite; TG69, tergites 69; ST6-8, sternites 68. Scale bars 1 mm (A, F), 500 μm (D), 200 μm (C, E, G), 100 μm (B)." figureDoi="http://doi.org/10.5281/zenodo.3602682" httpUri="https://zenodo.org/record/3602682/files/figure.png" pageId="3" pageNumber="484">4C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>