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<document id="2716923FF2BD4C3BD64D168C67AE07D8" ID-DOI="10.5281/zenodo.277211" ID-GBIF-Dataset="02822034-b581-4195-a509-5ee435369d0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="277211" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460276828081" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2011" docId="D34C87B84D2B2635FF44FABE63FBFAAA" docLanguage="en" docName="zt02819p050.pdf" docOrigin="Zootaxa 2819" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Piromis websteri Day 1973" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="2F75FFC04D32262EFFD3FFA9670EFFD2" masterDocTitle="Revision of Piromis Kinberg, 1867 and Pycnoderma Grube, 1877 (Polychaeta: Flabelligeridae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="26" updateTime="1698248591664" updateUser="plazi">
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<mods:title id="A02483CAA33BCA1A1D0C4B6A5E9D4ADD">Revision of Piromis Kinberg, 1867 and Pycnoderma Grube, 1877 (Polychaeta: Flabelligeridae)</mods:title>
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<mods:namePart id="020CF919C3B2DB89702E07D356AB15E8">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:typeOfResource id="552F558CE0988A5221F5D7D675E00D33">text</mods:typeOfResource>
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<mods:date id="F9E2637CA329878F9F03AB7CC63FC30F">2011</mods:date>
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<mods:number id="76A8B6E7718695DC3E1A715C8CD04E25">2819</mods:number>
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<mods:identifier id="09592F4A66FF2F1939A6DA04381F91AF" type="DOI">10.5281/zenodo.277211</mods:identifier>
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<treatment id="D34C87B84D2B2635FF44FABE63FBFAAA" ID-DOI="http://doi.org/10.5281/zenodo.6183601" ID-GBIF-Taxon="119396153" ID-Zenodo-Dep="6183601" LSID="urn:lsid:plazi:treatment:D34C87B84D2B2635FF44FABE63FBFAAA" httpUri="http://treatment.plazi.org/id/D34C87B84D2B2635FF44FABE63FBFAAA" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="13FF65254D2B2637FF44FABE660CFA80" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="5B5A36AE4D2B2637FF44FABE657BFAE3" blockId="25.[151,629,1303,1362]" box="[151,629,1303,1329]" pageId="25" pageNumber="26">
<heading id="001281C24D2B2637FF44FABE657BFAE3" bold="true" box="[151,629,1303,1329]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="6991EABC4D2B2637FF44FABE657BFAE3" bold="true" box="[151,629,1303,1329]" pageId="25" pageNumber="26">
<taxonomicName id="9CE54D2D4D2B2637FF44FABE66ECFAE3" ID-CoL="4J88X" authority="Day, 1973" authorityName="Day" authorityYear="1973" box="[151,482,1303,1329]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="25" pageNumber="26" phylum="Annelida" rank="species" species="websteri">
<emphasis id="6991EABC4D2B2637FF44FABE6651FAE3" bold="true" box="[151,351,1303,1329]" italics="true" pageId="25" pageNumber="26">Piromis websteri</emphasis>
<bibRefCitation id="3F744B5F4D2B2637FEB4FABE66ECFAE3" author="Day" box="[359,482,1303,1329]" pageId="25" pageNumber="26" refString="Day, J. H. (1973) New Polychaeta from Beaufort, with a key to all species recorded from North Carolina. National Oceanographic and Atmospheric Administration, Technical Report of the National Marine Fisheries Service, Circular, 375, 1 - 153." type="journal article" year="1973">Day, 1973</bibRefCitation>
</taxonomicName>
status novo
</emphasis>
</heading>
</paragraph>
<paragraph id="5B5A36AE4D2B2637FF44FA93660CFA80" blockId="25.[151,629,1303,1362]" box="[151,258,1338,1362]" pageId="25" pageNumber="26">
<figureCitation id="C3DE2A2B4D2B2637FF44FA93660CFA80" box="[151,258,1338,1362]" captionStart="FIGURE 11" captionStartId="26.[151,250,1548,1571]" captionTargetBox="[174,1410,665,1524]" captionTargetId="figure@26.[169,1417,661,1528]" captionTargetPageId="26" captionText="FIGURE 11. Piromis websteri Day, 1973, st. n. A. Syntype (USNM 433), complete, lateral view. B – F: non-type specimen (USNM- 53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B – C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm." httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="25" pageNumber="26">Figure 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="13FF65254D2B2637FF44FA2B6579FA65" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="5B5A36AE4D2B2637FF44FA2B6579FA65" blockId="25.[151,787,1410,1463]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="7BF511804D2B2637FF44FA2B6579FA65" pageId="25" pageNumber="26">
<taxonomicName id="9CE54D2D4D2B2637FF44FA2B6516FA4A" ID-CoL="7D998" authority="Webster, 1879:245" authorityName="Webster" authorityPageNumber="245" authorityYear="1879" box="[151,536,1410,1432]" class="Polychaeta" family="Flabelligeridae" genus="Trophonia" kingdom="Animalia" order="Terebellida" pageId="25" pageNumber="26" phylum="Annelida" rank="species" species="arenosa">
<emphasis id="6991EABC4D2B2637FF44FA2B6659FA4A" box="[151,343,1410,1432]" italics="true" pageId="25" pageNumber="26">Trophonia arenosa</emphasis>
<treatmentCitation id="DA4410BF4D2B2637FE8FFA2B6516FA4A" author="Webster" box="[348,536,1410,1432]" page="245" pageId="25" pageNumber="26" year="1879">
<bibRefCitation id="3F744B5F4D2B2637FE8FFA2B66E1FA4A" author="Webster" box="[348,495,1410,1432]" pageId="25" pageNumber="26" refString="Webster, H. E. (1879) On the Annelida Chaetopoda of the Virginian Coast. Transactions of the Albany Institute, 9, 202 - 269." type="journal article" year="1879">Webster, 1879</bibRefCitation>
:245
</treatmentCitation>
</taxonomicName>
–246, Pl. 7, Figs. 92–97.
<taxonomicName id="9CE54D2D4D2B2637FF44FA0B6511FA65" ID-CoL="4J88X" authority="Day, 1973:109" authorityName="Day" authorityPageNumber="109" authorityYear="1973" box="[151,543,1441,1463]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="25" pageNumber="26" phylum="Annelida" rank="subSpecies" species="eruca" subSpecies="websteri">
<emphasis id="6991EABC4D2B2637FF44FA0B668CFA65" box="[151,386,1441,1463]" italics="true" pageId="25" pageNumber="26">Piromis eruca websteri</emphasis>
<treatmentCitation id="DA4410BF4D2B2637FE5BFA0B6511FA65" author="Day" box="[392,543,1442,1463]" page="109" pageId="25" pageNumber="26" year="1973">
<bibRefCitation id="3F744B5F4D2B2637FE5BFA0B66FDFA65" author="Day" box="[392,499,1442,1463]" pageId="25" pageNumber="26" refString="Day, J. H. (1973) New Polychaeta from Beaufort, with a key to all species recorded from North Carolina. National Oceanographic and Atmospheric Administration, Technical Report of the National Marine Fisheries Service, Circular, 375, 1 - 153." type="journal article" year="1973">Day, 1973</bibRefCitation>
:109
</treatmentCitation>
</taxonomicName>
, no figs.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="13FF65254D2B2634FF44FA4D67C6FD8D" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="5B5A36AE4D2B2637FF44FA4D6244F9B5" blockId="25.[151,1436,1507,2036]" pageId="25" pageNumber="26">
<emphasis id="6991EABC4D2B2637FF44FA4D65B7FA2E" bold="true" box="[151,697,1507,1532]" pageId="25" pageNumber="26">
<typeStatus id="845E880C4D2B2637FF44FA4D67DCFA2E" box="[151,210,1508,1532]" pageId="25" pageNumber="26">Type</typeStatus>
material. Northwestern Atlantic Ocean.
</emphasis>
<typeStatus id="845E880C4D2B2637FD12FA4D6429FA2E" box="[705,807,1508,1532]" pageId="25" pageNumber="26" type="syntype">Syntypes</typeStatus>
(
<collectionCode id="3DF4AE6B4D2B2637FCEBFA4D6485FA2E" LSID="urn:lsid:biocol.org:col:34871" box="[824,907,1508,1532]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
433), H.E. Webster coll., id., Virginia, no further data (two almost complete specimens lacking posterior end, anterior end dissected; there might have been a third specimen since one of the original illustrations indicates a cross section of a median chaetiger; most chaetae broken, sediment cover removed from many areas. Longer
<typeStatus id="845E880C4D2B2637FCFCF9F86488F9BB" box="[815,902,1617,1641]" pageId="25" pageNumber="26" type="syntype">syntype</typeStatus>
<quantity id="9C1D9B4B4D2B2637FC5EF9E664D2F9BA" box="[909,988,1615,1640]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="25" pageNumber="26" unit="mm" value="43.0">43 mm</quantity>
long,
<quantity id="9C1D9B4B4D2B2637FBF1F9E6636DF9BA" box="[1058,1123,1615,1640]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="26" unit="mm" value="5.0">5 mm</quantity>
wide, 63 chaetigers.
</paragraph>
<paragraph id="5B5A36AE4D2B2634FF16F9DA67C6FD8D" blockId="25.[151,1436,1507,2036]" lastBlockId="26.[151,1436,151,607]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="6991EABC4D2B2637FF16F9DA6521F95E" bold="true" box="[197,559,1651,1676]" pageId="25" pageNumber="26">Additional material. Virginia:</emphasis>
Anterior fragment (
<collectionCode id="3DF4AE6B4D2B2637FCC5F9DD6467F95E" LSID="urn:lsid:biocol.org:col:34871" box="[790,873,1652,1676]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
57058), slightly damaged, Chincoteague, Chesapeake Bay,
<date id="2F5B106E4D2B2637FEC9F93E66B9F97D" box="[282,439,1687,1712]" pageId="25" pageNumber="26" value="1965-08-12">12 Aug. 1965</date>
, in eelgrass (dorsal lobes in chaetigers 2–6). Two specimens (VIMS-535), Eelgrass bed, Chincoteague,
<date id="2F5B106E4D2B2637FEECF91566D2F906" box="[319,476,1724,1748]" pageId="25" pageNumber="26" value="1965-08-12">12 Aug. 1965</date>
, M.L. Wass, coll. (both with neurospines in chaetiger 6).
<emphasis id="6991EABC4D2B2637FBB3F915622FF906" bold="true" box="[1120,1313,1723,1748]" pageId="25" pageNumber="26">North Carolina:</emphasis>
Two specimens (
<collectionCode id="3DF4AE6B4D2B2637FF3BF9766635F925" LSID="urn:lsid:biocol.org:col:34871" box="[232,315,1759,1783]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
53880), one complete with swollen posterior end, Banks Channel, Wrightsville Beach, North Carolina, intertidal, muddy sand, no date, T. Fox, coll. (first neurohooks in chaetiger 6; second chaetiger with two ventral tubercles). One specimen (
<collectionCode id="3DF4AE6B4D2B2637FDC9F8816563F892" LSID="urn:lsid:biocol.org:col:34871" box="[538,621,1832,1856]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
53881), Banks Channel, Wrightsville Beach, North Carolina, intertidal, shelly mud,
<date id="2F5B106E4D2B2637FEF2F8E566A0F8B6" box="[289,430,1868,1892]" pageId="25" pageNumber="26" value="1973-07-27">27 Jul. 1973</date>
, C. Jenner, coll. (first neurohooks in chaetiger 6; second chaetiger with two ventral tubercles). One specimen (
<collectionCode id="3DF4AE6B4D2B2637FE5FF8D966D1F85A" LSID="urn:lsid:biocol.org:col:34871" box="[396,479,1904,1928]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
53883), without posterior end, Banks Channel, Wrightsville Beach, North Carolina, sandy mud,
<date id="2F5B106E4D2B2637FEF3F83D66A5F87E" box="[288,427,1940,1964]" pageId="25" pageNumber="26" value="1974-03-08">8 Mar. 1974</date>
, C. Jenner, coll. (first neurohooks in chaetiger 6; second chaetiger with two ventral tubercles). One specimen (
<collectionCode id="3DF4AE6B4D2B2637FE5FF81166D1F802" LSID="urn:lsid:biocol.org:col:34871" box="[396,479,1976,2000]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="26">USNM</collectionCode>
53884), without posterior end, Banks Channel, Wrightsville Beach, North Carolina, soft mud,
<date id="2F5B106E4D2B2637FED8F8756695F826" box="[267,411,2012,2036]" pageId="25" pageNumber="26" value="1974-11-02">2 Nov. 1974</date>
, C. Jenner, coll. (first neurohooks in chaetiger 6; second chaetiger with two ventral tubercles). One specimen (
<collectionCode id="3DF4AE6B4D282634FE5DFF3E66EFFF7D" LSID="urn:lsid:biocol.org:col:34871" box="[398,481,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
53885), from muddy sand, Bogue Sound, Morehead City,
<date id="2F5B106E4D282634FB54FF3E622AFF7D" box="[1159,1316,151,175]" pageId="26" pageNumber="27" value="1975-06-23">23 June 1975</date>
, S.L. Gardiner, coll. One specimen (
<collectionCode id="3DF4AE6B4D282634FE00FF156528FF06" LSID="urn:lsid:biocol.org:col:34871" box="[467,550,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
61742) from Middle Marsh, soft mud,
<date id="2F5B106E4D282634FC28FF15639EFF06" box="[1019,1168,188,212]" pageId="26" pageNumber="27" value="1976-07-23">23 Jul. 1976</date>
.
<emphasis id="6991EABC4D282634FB4DFF12626CFF06" bold="true" box="[1182,1378,187,212]" pageId="26" pageNumber="27">
Gulf of
<collectingCountry id="23F2763E4D282634FAD1FF126253FF06" box="[1282,1373,187,212]" name="Mexico" pageId="26" pageNumber="27">Mexico</collectingCountry>
:
</emphasis>
One specimen (UMML-unnumbered), Rookery Bay (26°01ʹ30ʺ N, 81°44ʹ40ʺ W), 32.1.7, no further data. Three anterior fragments (
<collectionCode id="3DF4AE6B4D282634FEC6FEAD6669FECE" LSID="urn:lsid:biocol.org:col:34871" box="[277,359,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
45650), together with an anterior fragment of
<taxonomicName id="9CE54D2D4D282634FCBEFEAC6360FECF" authority="Tampa Bay" authorityName="Tampa Bay" box="[877,1134,260,285]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="robertsi">
<emphasis id="6991EABC4D282634FCBEFEAC64EEFECE" box="[877,992,260,284]" italics="true" pageId="26" pageNumber="27">P. robertsi</emphasis>
, Tampa Bay
</taxonomicName>
, Florida, 1963, J.L. Taylor, coll. (all with neurohooks from chaetiger 6). One specimen (
<collectionCode id="3DF4AE6B4D282634FCE5FE8E6487FEED" LSID="urn:lsid:biocol.org:col:34871" box="[822,905,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
71158),
<quantity id="9C1D9B4B4D282634FC38FE8E6313FEED" box="[1003,1053,295,319]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="26" pageNumber="27" unit="mi" value="3.0">3 mi</quantity>
SE off Dauphin Island (30°11ʹ17ʺ N, 88°07ʹ17ʺ W),
<date id="2F5B106E4D282634FEB2FEE566FAFEB6" box="[353,500,332,356]" pageId="26" pageNumber="27" value="1981-09-18">18 Sep. 1981</date>
, 42 feet, S.L. Gardiner id. Three specimens (
<collectionCode id="3DF4AE6B4D282634FC35FEE56337FEB6" LSID="urn:lsid:biocol.org:col:34871" box="[998,1081,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
61742), one complete, “Middle Marshʺ, North Carolina,
<date id="2F5B106E4D282634FE66FEC66548FE55" box="[437,582,367,391]" pageId="26" pageNumber="27" value="1976-07-23">23 Jul. 1976</date>
, soft mud, S.L. Gardiner, coll. (dorsal, paired tubercles in chaetigers 2–6). One specimen (
<collectionCode id="3DF4AE6B4D282634FE96FE3D6696FE7E" LSID="urn:lsid:biocol.org:col:34871" box="[325,408,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
75920), anterior fragment, damaged, R/V
<emphasis id="6991EABC4D282634FCA7FE3C6360FE7E" box="[884,1134,404,428]" italics="true" pageId="26" pageNumber="27">Captain Brady Joseph</emphasis>
, Sta. M10-2 (29°39ʹ52ʺ N, 93°28ʹ34ʺ W), off
<collectingCountry id="23F2763E4D282634FEB5FE1E66DEFE1D" box="[358,464,439,463]" name="Cameroon" pageId="26" pageNumber="27">Cameron</collectingCountry>
, Louisiana,
<quantity id="9C1D9B4B4D282634FD84FE1E6596FE1D" box="[599,664,439,463]" metricMagnitude="0" metricUnit="m" metricValue="9.8" pageId="26" pageNumber="27" unit="m" value="9.8">9.8 m</quantity>
,
<date id="2F5B106E4D282634FD77FE1E6418FE1D" box="[676,790,439,463]" pageId="26" pageNumber="27" value="1982-02">Feb. 1982</date>
, G.R. Gaston, coll. (hooks start apparently in chaetiger 6). One specimen (
<collectionCode id="3DF4AE6B4D282634FE97FE756698FE26" LSID="urn:lsid:biocol.org:col:34871" box="[324,406,476,500]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
75921), anterior fragment, R/V
<emphasis id="6991EABC4D282634FD2DFE7464F6FE26" box="[766,1016,476,500]" italics="true" pageId="26" pageNumber="27">Captain Brady Joseph</emphasis>
, Sta. M10-2 (29°39ʹ52ʺ N, 93°28ʹ34ʺ W), off
<collectingCountry id="23F2763E4D282634FF2BFE56666CFDC5" box="[248,354,511,535]" name="Cameroon" pageId="26" pageNumber="27">Cameron</collectingCountry>
, Louisiana,
<quantity id="9C1D9B4B4D282634FE21FE566536FDC5" box="[498,568,511,535]" metricMagnitude="0" metricUnit="m" metricValue="9.8" pageId="26" pageNumber="27" unit="m" value="9.8">9.8 m</quantity>
,
<date id="2F5B106E4D282634FD9BFDA965B1FDC5" box="[584,703,511,536]" pageId="26" pageNumber="27" value="1982-04">Apr. 1982</date>
, G.R. Gaston, coll. (hooks start in chaetiger 6). One specimen (
<collectionCode id="3DF4AE6B4D282634FF4DFD8D67FFFDEE" LSID="urn:lsid:biocol.org:col:34871" box="[158,241,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="27">USNM</collectionCode>
75923), R/V
<emphasis id="6991EABC4D282634FE59FD8C658AFDEE" box="[394,644,548,572]" italics="true" pageId="26" pageNumber="27">Captain Brady Joseph</emphasis>
, Sta. 10-6(29°39ʹ52ʺ N, 93°28ʹ34’ W),
<quantity id="9C1D9B4B4D282634FB93FD8D638CFDEE" box="[1088,1154,548,572]" metricMagnitude="0" metricUnit="m" metricValue="9.8" pageId="26" pageNumber="27" unit="m" value="9.8">9.8 m</quantity>
,
<date id="2F5B106E4D282634FB5DFD8C620CFDEE" box="[1166,1282,548,573]" pageId="26" pageNumber="27" value="1982-05">May 1982</date>
, G.R. Gaston, coll.
</paragraph>
</subSubSection>
<caption id="0F9A66264D282634FF44F9A564FDF94C" httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="26" pageNumber="27" targetBox="[174,1410,665,1524]" targetPageId="26">
<paragraph id="5B5A36AE4D282634FF44F9A564FDF94C" blockId="26.[151,1436,1548,1694]" pageId="26" pageNumber="27">
<emphasis id="6991EABC4D282634FF44F9A56614F9F0" bold="true" box="[151,282,1548,1571]" pageId="26" pageNumber="27">FIGURE 11</emphasis>
.
<taxonomicName id="9CE54D2D4D282634FEFAF9A76543F9F0" authority="Day, 1973" authorityName="Day" authorityYear="1973" box="[297,589,1549,1571]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="26" pageNumber="27" phylum="Annelida" rank="species" species="websteri" status="subsp. nov.">
<emphasis id="6991EABC4D282634FEFAF9A76676F9F1" box="[297,376,1550,1571]" italics="true" pageId="26" pageNumber="27">Piromis</emphasis>
<emphasis id="6991EABC4D282634FE52F9A466D9F9F1" box="[385,471,1549,1571]" italics="true" pageId="26" pageNumber="27">websteri</emphasis>
Day, 1973
</taxonomicName>
,
<taxonomicNameLabel id="72A257C74D282634FD89F9A76583F9F1" box="[602,653,1550,1571]" pageId="26" pageNumber="27" rank="subSpecies">st. n.</taxonomicNameLabel>
A. Syntype (USNM 433), complete, lateral view. B–F: non-type specimen (USNM-53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B–C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm.
</paragraph>
</caption>
<subSubSection id="13FF65254D282635FF16F96366BAFB72" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="5B5A36AE4D282634FF16F9636668F8A0" blockId="26.[151,1436,1738,2015]" pageId="26" pageNumber="27">
<emphasis id="6991EABC4D282634FF16F963667DF931" bold="true" box="[197,371,1738,1763]" pageId="26" pageNumber="27">Redescription.</emphasis>
Smaller
<typeStatus id="845E880C4D282634FE08F962653CF931" box="[475,562,1739,1763]" pageId="26" pageNumber="27" type="syntype">syntype</typeStatus>
tapering posteriorly, oval in cross section, sediment cover removed in many areas, remaining as a thin layer of sand grains with clear, eroded areas dorsally and laterally (
<figureCitation id="C3DE2A2B4D282634FB69F9476204F8D4" box="[1210,1290,1774,1798]" captionStart="FIGURE 11" captionStartId="26.[151,250,1548,1571]" captionTargetBox="[174,1410,665,1524]" captionTargetId="figure@26.[169,1417,661,1528]" captionTargetPageId="26" captionText="FIGURE 11. Piromis websteri Day, 1973, st. n. A. Syntype (USNM 433), complete, lateral view. B – F: non-type specimen (USNM- 53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B – C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm." httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="26" pageNumber="27">Fig. 11</figureCitation>
A), sediment cover better conserved in other specimens (
<figureCitation id="C3DE2A2B4D282634FDAAF8BB65C7F8F8" box="[633,713,1810,1834]" captionStart="FIGURE 11" captionStartId="26.[151,250,1548,1571]" captionTargetBox="[174,1410,665,1524]" captionTargetId="figure@26.[169,1417,661,1528]" captionTargetPageId="26" captionText="FIGURE 11. Piromis websteri Day, 1973, st. n. A. Syntype (USNM 433), complete, lateral view. B – F: non-type specimen (USNM- 53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B – C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm." httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="26" pageNumber="27">Fig. 11</figureCitation>
B). Anterior end with five chaetigers directed anteriorly.
<typeStatus id="845E880C4D282634FA93F8BB6292F8F8" box="[1344,1436,1810,1834]" pageId="26" pageNumber="27" type="syntype">Syntype</typeStatus>
<quantity id="9C1D9B4B4D282634FF44F89F67E9F89C" box="[151,231,1846,1870]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="27" unit="mm" value="28.0">28 mm</quantity>
long,
<quantity id="9C1D9B4B4D282634FEE3F89F667CF89C" box="[304,370,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="26" pageNumber="27" unit="mm" value="3.0">3 mm</quantity>
wide, cephalic cage
<quantity id="9C1D9B4B4D282634FDB2F89F65AAF89C" box="[609,676,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="27" unit="mm" value="4.0">4 mm</quantity>
long, 58 chaetigers (with a longitudinal dissection, ventrally along chaetigers 45–52).
</paragraph>
<paragraph id="5B5A36AE4D282635FF14F8D76406FF06" blockId="26.[151,1436,1738,2015]" lastBlockId="27.[151,1436,151,1400]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Cephalic hood not exposed, anterior end retracted (original dissection by Day; other specimens with everted anterior end show a short cephalic hood, as long as chaetiger 1 length). Anterior end details based on non-type materials (
<figureCitation id="C3DE2A2B4D282634FEDEF86F6652F80C" box="[269,348,1990,2014]" captionStart="FIGURE 11" captionStartId="26.[151,250,1548,1571]" captionTargetBox="[174,1410,665,1524]" captionTargetId="figure@26.[169,1417,661,1528]" captionTargetPageId="26" captionText="FIGURE 11. Piromis websteri Day, 1973, st. n. A. Syntype (USNM 433), complete, lateral view. B – F: non-type specimen (USNM- 53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B – C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm." httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="26" pageNumber="27">Fig. 11</figureCitation>
C; VIMS-535). Prostomium low; eyes large, slightly pigmented. Caruncle with diffuse pigmentation. Another specimen (USNM 53883), with anterior end everted; prostomium low; eyes large, fused. Caruncle with median keel pale, lateral ridges basally pigmented.
</paragraph>
<paragraph id="5B5A36AE4D292635FF14FF766633FE55" blockId="27.[151,1436,151,1400]" pageId="27" pageNumber="28">Cephalic hood short, margin entire, not crenulated or papillated. Palps long, corrugated, with black spots; palp keels low (larger specimens with pigmented. swollen palp keels). Mouth with two thick lateral lips, dorsal lip reduced, ventral lip thin; buccal organ exposed. Branchiae cirriform, placed on a tongue-shaped lobe, separated in two lateral groups, each with about 60 branchial filaments. Size relationship with palps not discernible. Nephridial lobes not seen.</paragraph>
<paragraph id="5B5A36AE4D292635FF14FE3D662AFE26" blockId="27.[151,1436,151,1400]" pageId="27" pageNumber="28">Cephalic cage chaetae slightly longer than body width; cephalic cage made by the first three chaetigers, but longest chaetae in chaetigers 1–2; chaetae arranged in a tuft. Notopodia 1–2 with 2–3 chaetae, neuropodia 1–2 with 3–4 chaetae.</paragraph>
<paragraph id="5B5A36AE4D292635FF14FE566672FD3D" blockId="27.[151,1436,151,1400]" pageId="27" pageNumber="28">
Anterior dorsal margin of chaetiger 1 with a median projection, distally eroded; chaetigers 2–3 with pair of dorsolateral elongated papillae, and single long notopodial papilla. Dorsal tubercles eroded in
<typeStatus id="845E880C4D292635FB6EFD8C622CFDEF" box="[1213,1314,549,573]" pageId="27" pageNumber="28" type="syntype">syntypes</typeStatus>
, present in non-type specimens from North Carolina. Post-cephalic cage chaetigers not elongated. Chaetal transition from cephalic cage to body chaetae abrupt; chaetiger 4 with short notochaetae, bifid neurohooks replacing capillaries (after Webster and Day; no chaetae left, but confirmed in the additional materials). Chaetiger 1 with two ventrolateral lobes, branched, on its anterior margin, better preserved in some specimens (
<figureCitation id="C3DE2A2B4D292635FBF2FD1D637FFD1E" box="[1057,1137,692,716]" captionStart="FIGURE 11" captionStartId="26.[151,250,1548,1571]" captionTargetBox="[174,1410,665,1524]" captionTargetId="figure@26.[169,1417,661,1528]" captionTargetPageId="26" captionText="FIGURE 11. Piromis websteri Day, 1973, st. n. A. Syntype (USNM 433), complete, lateral view. B – F: non-type specimen (USNM- 53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B – C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm." httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="27" pageNumber="28">Fig. 11</figureCitation>
B, USNM 53883). Gonopodial lobes not seen.
</paragraph>
<paragraph id="5B5A36AE4D292635FF14FD5564BDFC8E" blockId="27.[151,1436,151,1400]" pageId="27" pageNumber="28">Anterior parapodia projecting more than median or posterior ones; those as lower chaetal lobes. Parapodia placed over the body corners in cross section, at about the same distance to each other. Notopodia with long pre- and postchaetal papillae, the latter longer, especially on chaetigers 1–4.</paragraph>
<paragraph id="5B5A36AE4D292635FF14FCCE64ABFBE6" blockId="27.[151,1436,151,1400]" pageId="27" pageNumber="28">
Median notochaetae in transverse row, with about six chaetae per bundle (up to
<quantity id="9C1D9B4B4D292635FB51FCCE63B1FCAD" box="[1154,1215,871,895]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="27" pageNumber="28" unit="in" value="11.0">11 in</quantity>
larger specimens), reduced to 5 (
<quantity id="9C1D9B4B4D292635FEE2FC256652FC76" box="[305,348,908,932]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="27" pageNumber="28" unit="in" value="7.0">7 in</quantity>
larger specimens) in posterior chaetigers, about 1/3 as long as body width; all notochaetae multiarticulated capillaries (
<figureCitation id="C3DE2A2B4D292635FE49FC0666E4FC15" box="[410,490,943,967]" captionStart="FIGURE 11" captionStartId="26.[151,250,1548,1571]" captionTargetBox="[174,1410,665,1524]" captionTargetId="figure@26.[169,1417,661,1528]" captionTargetPageId="26" captionText="FIGURE 11. Piromis websteri Day, 1973, st. n. A. Syntype (USNM 433), complete, lateral view. B – F: non-type specimen (USNM- 53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B – C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm." httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="27" pageNumber="28">Fig. 11</figureCitation>
D) with short articles basally, longer medially and distally. Neuropodia less prominent than notopodia; long papillae mostly postchaetal. From chaetiger 4, 6–7 multiarticulate bidentate neurohooks, arranged in a J-shape. Anterior and median chaetigers with neurochaetae with long articles (
<figureCitation id="C3DE2A2B4D292635FB68FC5E6203FBDD" box="[1211,1293,1015,1039]" captionStart="FIGURE 11" captionStartId="26.[151,250,1548,1571]" captionTargetBox="[174,1410,665,1524]" captionTargetId="figure@26.[169,1417,661,1528]" captionTargetPageId="26" captionText="FIGURE 11. Piromis websteri Day, 1973, st. n. A. Syntype (USNM 433), complete, lateral view. B – F: non-type specimen (USNM- 53883). B. Anterior end, dorsal view. C. Anterior end, ventral view, head exposed (Br: branchiae, CH: cephalic hood, DL: dorsal lip, LL: lateral lip, Pa: palp, PK: palp keel, VL: ventral lip). D. Notochaetae from chaetiger 20, showing basal, medial and distal regions. E. Neurochaetae from chaetiger 35, insert showing tips. G. Neurochaetae from chaetiger 125, insert showing tips. Scale bars: A: 2.5 mm, B – C: 0.5 mm, D: 70 µm, E: 150 µm, F: 50 µm." httpUri="https://zenodo.org/record/277222/files/figure.png" pageId="27" pageNumber="28">Fig. 11</figureCitation>
E); posterior chaetigers with 4–5 long articles. Tips bidentate, accessory tooth thin.
</paragraph>
<paragraph id="5B5A36AE4D292635FF14FBE966BAFB72" blockId="27.[151,1436,151,1400]" pageId="27" pageNumber="28">Posterior end damaged in Virginia specimens (specimens from Alabama and another one from North Carolina with terminal anus (another specimen with dorsoterminal anus), without anal cirri, but with marginal papillae, and a ventral elongated lobe).</paragraph>
</subSubSection>
<subSubSection id="13FF65254D292635FF16FB056238FA86" pageId="27" pageNumber="28" type="discussion">
<paragraph id="5B5A36AE4D292635FF16FB056238FA86" blockId="27.[151,1436,151,1400]" pageId="27" pageNumber="28">
<emphasis id="6991EABC4D292635FF16FB056637FB16" bold="true" box="[197,313,1196,1220]" pageId="27" pageNumber="28">Remarks.</emphasis>
<taxonomicName id="9CE54D2D4D292635FE92FB04658FFB16" authority="Day, 1973" authorityName="Day" authorityYear="1973" box="[321,641,1196,1221]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="27" pageNumber="28" phylum="Annelida" rank="species" species="websteri">
<emphasis id="6991EABC4D292635FE92FB04650FFB16" box="[321,513,1196,1220]" italics="true" pageId="27" pageNumber="28">Piromis websteri</emphasis>
<bibRefCitation id="3F744B5F4D292635FDDAFB04658FFB16" author="Day" box="[521,641,1196,1221]" pageId="27" pageNumber="28" refString="Day, J. H. (1973) New Polychaeta from Beaufort, with a key to all species recorded from North Carolina. National Oceanographic and Atmospheric Administration, Technical Report of the National Marine Fisheries Service, Circular, 375, 1 - 153." type="journal article" year="1973">Day, 1973</bibRefCitation>
</taxonomicName>
, originally described as a subspecies, does closely resemble
<emphasis id="6991EABC4D292635FAE6FB0467E0FB3A" italics="true" pageId="27" pageNumber="28">P. k is e m - boanus,</emphasis>
but differs sufficiently to warrant full species status and is herein so elevated.
<taxonomicName id="9CE54D2D4D292635FB59FB786245FB3A" box="[1162,1355,1232,1256]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="27" pageNumber="28" phylum="Annelida" rank="species" species="websteri">
<emphasis id="6991EABC4D292635FB59FB786245FB3A" box="[1162,1355,1232,1256]" italics="true" pageId="27" pageNumber="28">Piromis websteri</emphasis>
</taxonomicName>
and
<taxonomicName id="9CE54D2D4D292635FA54FB786626FADE" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="27" pageNumber="28" phylum="Annelida" rank="species" species="kisemboanus">
<emphasis id="6991EABC4D292635FA54FB786626FADE" italics="true" pageId="27" pageNumber="28">P. kisemboanus</emphasis>
</taxonomicName>
differ in the relative size of the sediment particles that adhere to the tunic and on the relative number of notochaetae. Thus, in
<emphasis id="6991EABC4D292635FE7DFAB0652BFAE2" box="[430,549,1304,1328]" italics="true" pageId="27" pageNumber="28">P. w e b s t e r i</emphasis>
the sediment grains are fine and there are about eight notochaetae per bundle, while in
<taxonomicName id="9CE54D2D4D292635FF2AFA9466ABFA86" box="[249,421,1340,1364]" class="Polychaeta" family="Flabelligeridae" genus="Piromis" kingdom="Animalia" order="Terebellida" pageId="27" pageNumber="28" phylum="Annelida" rank="species" species="kisemboanus">
<emphasis id="6991EABC4D292635FF2AFA9466ABFA86" box="[249,421,1340,1364]" italics="true" pageId="27" pageNumber="28">P. kisemboanus</emphasis>
</taxonomicName>
the sediment particles are larger and there are about four notochaetae per bundle.
</paragraph>
</subSubSection>
<subSubSection id="13FF65254D292635FF16FAF663FBFAAA" box="[197,1269,1375,1400]" pageId="27" pageNumber="28" type="distribution">
<paragraph id="5B5A36AE4D292635FF16FAF663FBFAAA" blockId="27.[151,1436,151,1400]" box="[197,1269,1375,1400]" pageId="27" pageNumber="28">
<emphasis id="6991EABC4D292635FF16FAF66650FAAA" bold="true" box="[197,350,1375,1400]" pageId="27" pageNumber="28">Distribution.</emphasis>
Intertidal or shallow water in Virginia, North Carolina and in the Gulf of
<collectingCountry id="23F2763E4D292635FB4BFAC963FFFAAA" box="[1176,1265,1376,1400]" name="Mexico" pageId="27" pageNumber="28">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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