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<document id="DB19B27B06B26E1894E1CB8DC54D3895" ID-CLB-Dataset="39184" ID-DOI="10.11646/zootaxa.4105.5.2" ID-GBIF-Dataset="60ca4dbd-002b-4ca6-999d-9174f9b02324" ID-ISSN="1175-5326" ID-Zenodo-Dep="255425" ID-ZooBank="52401DF9-2901-4AF6-BA17-696D39280B3C" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1461736713249" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2016" docId="E338D67535102168FF05FD5C77076A06" docLanguage="en" docName="zootaxa.4105.5.2.pdf.imf" docOrigin="Zootaxa 4105 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guyalna nigra Boulard 1999" docType="treatment" docVersion="22" lastPageNumber="444" masterDocId="1F01AE0D351E2166FF92FF9876486E39" masterDocTitle="The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species" masterLastPageNumber="454" masterPageNumber="430" pageNumber="444" updateTime="1698625263794" updateUser="plazi">
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<mods:title id="AE88E9C42D868BE9652FB8A5CFB5A45D">The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species</mods:title>
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<subSubSection id="238B34E835102168FF05FD5C77A66CE4" box="[151,494,708,733]" pageId="14" pageNumber="444" type="nomenclature">
<paragraph id="6B2E676335102168FF05FD5C77A66CE4" blockId="14.[151,494,708,769]" box="[151,494,708,733]" pageId="14" pageNumber="444">
<heading id="3066D00F35102168FF05FD5C77A66CE4" bold="true" box="[151,494,708,733]" fontSize="10" level="2" pageId="14" pageNumber="444" reason="2">
<taxonomicName id="AC911CE035102168FF05FD5C77A66CE4" ID-CoL="3HN7C" authority="Boulard, 1999" authorityName="Boulard" authorityYear="1999" box="[151,494,708,733]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="59E5BB7135102168FF05FD5C77A66CE4" bold="true" box="[151,494,708,733]" pageId="14" pageNumber="444">
<emphasis id="59E5BB7135102168FF05FD5C77776CE5" bold="true" box="[151,319,708,733]" italics="true" pageId="14" pageNumber="444">Guyalna nigra</emphasis>
<bibRefCitation id="0F001A9235102168FED4FD5C77A66CE4" author="Boulard, M." box="[326,494,708,733]" firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="14" pageNumber="444" pagination="77 - 116" part="11 / 12" refId="ref12501" refString="Boulard, M. (1999) Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae). Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 11 / 12, 77 - 116." title="Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae)" type="journal article" year="1999">Boulard, 1999</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF05FD7076B16D38" box="[151,249,744,769]" pageId="14" pageNumber="444" type="description">
<paragraph id="6B2E676335102168FF05FD7076B16D38" blockId="14.[151,494,708,769]" box="[151,249,744,769]" pageId="14" pageNumber="444">
(
<figureCitation id="F3AA7BE635102168FF0DFD7076B86D38" box="[159,240,744,769]" captionStart="FIGURE 18" captionStartId="14.[151,250,1637,1659]" captionTargetBox="[151,1436,1143,1615]" captionTargetId="figure@14.[151,1436,1143,1615]" captionTargetPageId="14" captionText="FIGURE 18. Guyalna nigra Boulard, 1999. A, male, Brazil (AFSC). Scale bar: 2 cm; B, timbal cover (compared to image of holotype timbal cover (MNHN )). Scale bar: 2 mm." httpUri="https://zenodo.org/record/255443/files/figure.png" pageId="14" pageNumber="444">Fig. 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF05FCA877BE6D7F" box="[151,502,816,838]" pageId="14" pageNumber="444" type="reference_group">
<paragraph id="6B2E676335102168FF05FCA877BE6D7F" blockId="14.[151,502,816,838]" box="[151,502,816,838]" pageId="14" pageNumber="444">
<treatmentCitationGroup id="4B81404D35102168FF05FCA877BE6D7F" box="[151,502,816,838]" pageId="14" pageNumber="444">
<taxonomicName id="AC911CE035102168FF05FCA877BA6D7F" ID-CoL="3HN7C" authority="Boulard, 1999" authorityName="Boulard" authorityPageNumber="100" authorityYear="1999" box="[151,498,816,838]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="59E5BB7135102168FF05FCA877636D7F" box="[151,299,816,838]" italics="true" pageId="14" pageNumber="444">Guyalna nigra</emphasis>
<treatmentCitation id="EA30417235102168FEA3FCA877BA6D7F" author="Boulard" box="[305,498,816,838]" page="100" pageId="14" pageNumber="444" year="1999">
<bibRefCitation id="0F001A9235102168FEA3FCA877BA6D7F" author="Boulard, M." box="[305,498,816,838]" firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" page="100" pageId="14" pageNumber="444" pagination="77 - 116" part="11 / 12" refId="ref12501" refString="Boulard, M. (1999) Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae). Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 11 / 12, 77 - 116." title="Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae)" type="journal article" year="1999">Boulard 1999: 100</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF05FCEB74DC6DB5" box="[151,660,883,908]" pageId="14" pageNumber="444" type="materials_examined">
<paragraph id="6B2E676335102168FF05FCEB74DC6DB5" blockId="14.[151,1436,883,1088]" box="[151,660,883,908]" pageId="14" pageNumber="444">
<materialsCitation id="DBF96D3E35102168FF05FCEB74DC6DB5" ID-GBIF-Occurrence="4115800316" box="[151,660,883,908]" country="Brazil" location="Buzios" municipality="Buzios" pageId="14" pageNumber="444" specimenCount="1" stateProvince="Rio de Janiero" typeStatus="type">
<emphasis id="59E5BB7135102168FF05FCEB777D6DB5" bold="true" box="[151,309,883,908]" pageId="14" pageNumber="444">
<typeStatus id="B42AD9C135102168FF05FCEB769A6DB5" box="[151,210,883,908]" pageId="14" pageNumber="444">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="8B4AFD1935102168FEAEFCEB77C56DB5" box="[316,397,883,908]" pageId="14" pageNumber="444">Búzios</collectingMunicipality>
,
<collectingRegion id="A955A98135102168FE0AFCEB74756DB5" box="[408,573,883,908]" pageId="14" pageNumber="444">Rio de Janiero</collectingRegion>
,
<collectingCountry id="138627F335102168FDD5FCEB74C76DB5" box="[583,655,883,908]" name="Brazil" pageId="14" pageNumber="444">Brazil</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF55FC0F722C6DCE" pageId="14" pageNumber="444" type="discussion">
<paragraph id="6B2E676335102168FF55FC0F722C6DCE" blockId="14.[151,1436,883,1088]" pageId="14" pageNumber="444">
<emphasis id="59E5BB7135102168FF55FC0F77736D89" bold="true" box="[199,315,919,944]" pageId="14" pageNumber="444">Remarks.</emphasis>
The species is black and is similar in appearance but larger than
<emphasis id="59E5BB7135102168FB9CFC0072116D96" box="[1038,1113,920,943]" italics="true" pageId="14" pageNumber="444">G. c u t a</emphasis>
,
<taxonomicName id="AC911CE035102168FBF8FC0072856D89" box="[1130,1229,919,944]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="species" species="nadae">
<emphasis id="59E5BB7135102168FBF8FC0072856D89" box="[1130,1229,919,944]" italics="true" pageId="14" pageNumber="444">G. nadae</emphasis>
</taxonomicName>
and
<taxonomicName id="AC911CE035102168FA91FC0073DD6D89" box="[1283,1429,919,944]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="species" species="platyrhina">
<emphasis id="59E5BB7135102168FA91FC0073DD6D89" box="[1283,1429,919,944]" italics="true" pageId="14" pageNumber="444">G. platyrhina</emphasis>
</taxonomicName>
. It possesses a timbal cover that expands over the lateral timbal cavity with the rounded apex (
<figureCitation id="F3AA7BE635102168FB3EFC2372B76DEA" box="[1196,1279,955,980]" captionStart="FIGURE 18" captionStartId="14.[151,250,1637,1659]" captionTargetBox="[151,1436,1143,1615]" captionTargetId="figure@14.[151,1436,1143,1615]" captionTargetPageId="14" captionText="FIGURE 18. Guyalna nigra Boulard, 1999. A, male, Brazil (AFSC). Scale bar: 2 cm; B, timbal cover (compared to image of holotype timbal cover (MNHN )). Scale bar: 2 mm." httpUri="https://zenodo.org/record/255443/files/figure.png" pageId="14" pageNumber="444">Fig. 18</figureCitation>
). Body length
<quantity id="AC69CA8635102168FF05FC47775A6DCE" box="[151,274,991,1016]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="14" pageNumber="444" unit="mm" value="19.0" valueMax="22.0" valueMin="16.0">1622 mm</quantity>
, fore wing length
<quantity id="AC69CA8635102168FE4CFC4774236DCE" box="[478,619,991,1016]" metricMagnitude="-2" metricUnit="m" metricValue="2.575" metricValueMax="2.9" metricValueMin="2.25" pageId="14" pageNumber="444" unit="mm" value="25.75" valueMax="29.0" valueMin="22.5">22.529 mm</quantity>
(
<bibRefCitation id="0F001A9235102168FDE8FC4772106DCE" author="Boulard, M." box="[634,1112,991,1016]" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="14" pageNumber="444" pagination="77 - 116" part="11 / 12" refId="ref12501" refString="Boulard, M. (1999) Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae). Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 11 / 12, 77 - 116." title="Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae)" type="journal article" year="1999">Boulard 1999 and specimens in the AFSC</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF55FB9B77076A06" pageId="14" pageNumber="444" type="distribution">
<paragraph id="6B2E676335102168FF55FB9B77076A06" blockId="14.[151,1436,883,1088]" pageId="14" pageNumber="444">
<emphasis id="59E5BB7135102168FF55FB9B77286A25" bold="true" box="[199,352,1027,1052]" pageId="14" pageNumber="444">Distribution.</emphasis>
The species has been reported previously from
<collectingCountry id="138627F335102168FC39FB9B75A76A25" box="[939,1007,1027,1052]" name="Brazil" pageId="14" pageNumber="444">Brazil</collectingCountry>
and
<collectingCountry id="138627F335102168FBA0FB9B72966A25" box="[1074,1246,1027,1052]" name="French Guiana" pageId="14" pageNumber="444">French Guiana</collectingCountry>
(
<bibRefCitation id="0F001A9235102168FB61FB9B73DF6A22" author="Boulard, M." box="[1267,1431,1027,1052]" firstAuthor="Boulard" journalOrPublisher="Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea" pageId="14" pageNumber="444" pagination="77 - 116" part="11 / 12" refId="ref12501" refString="Boulard, M. (1999) Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae). Ecole Pratique des Hautes Etudes, Travaux du Laboratoire Biologie et Evolution des Insectes Hemipteroidea, 11 / 12, 77 - 116." title="Postures de cymbalisation, cymbalisations et cartes d'identite acoustique des cigales 3. - Especes tropicales des savanes et milieux ouverts (Cicadoidea, Cicadidae et Tibicinidae)" type="journal article" year="1999">Boulard 1999</bibRefCitation>
;
<bibRefCitation id="0F001A9235102168FF05FBBF770B6A06" author="Sanborn, A. F." box="[151,323,1063,1088]" firstAuthor="Sanborn" journalOrPublisher="Zoosystema" pageId="14" pageNumber="444" pagination="377 - 418" part="33" refId="ref14039" refString="Sanborn, A. F. (2011 a) Checklist of the cicadas of French Guiana including new records and the description of nine new species (Insecta, Hemiptera, Cicadoidea, Cicadidae). Zoosystema, 33, 377 - 418. http: // dx. doi. org / 10.5252 / z 2011 n 3 a 6" title="Checklist of the cicadas of French Guiana including new records and the description of nine new species (Insecta, Hemiptera, Cicadoidea, Cicadidae)" type="journal article" year="2011" yearSuffix="a">Sanborn 2011a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>