155 lines
19 KiB
XML
155 lines
19 KiB
XML
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<mods:title id="1B1F616FB9A701C8120616D02C3C645F">New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs</mods:title>
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<mods:name id="49F32533101607375C18EB395E0741CB" type="personal">
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<mods:namePart id="CFA7D708BB2F0B95819C0DA335E6D7E5">Wappler, Torsten</mods:namePart>
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<mods:title id="CD1A60B7662492965E18CDDBFC69624B">Zootaxa</mods:title>
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<mods:date id="BCC34812DCE0FCD9D1233C225FCBAEC2">2003</mods:date>
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<mods:number id="735120822C461AC8C8BA2824A4405914">374</mods:number>
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<mods:identifier id="F48515D2ECC24CAEE6F9ED17EFFBEF26" type="DOI">10.5281/zenodo.156768</mods:identifier>
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<treatment id="D33887DEFF973B0FFEF1014CE719D28C" ID-DOI="http://doi.org/10.5281/zenodo.6273737" ID-GBIF-Taxon="119332624" ID-Zenodo-Dep="6273737" LSID="urn:lsid:zoobank.org:act:5A3BC60C-CEEA-4CC4-8E21-E6F8E1621268" httpUri="http://treatment.plazi.org/id/D33887DEFF973B0FFEF1014CE719D28C" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<subSubSection id="138B6543FF973B0EFEF1014CE70CD271" box="[264,552,1633,1659]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="5B2E36C8FF973B0EFEF1014CE70CD271" blockId="10.[264,552,1633,1659]" box="[264,552,1633,1659]" pageId="10" pageNumber="11">
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<heading id="006681A4FF973B0EFEF1014CE70CD271" bold="true" box="[264,552,1633,1659]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="69E5EADAFF973B0EFEF1014CE70CD271" bold="true" box="[264,552,1633,1659]" pageId="10" pageNumber="11">
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<taxonomicName id="9C914D4BFF973B0EFEF1014CE4EBD271" box="[264,463,1633,1659]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF973B0EFEF1014CE4EBD271" bold="true" box="[264,463,1633,1659]" italics="true" pageId="10" pageNumber="11">Oblongomorpha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF973B0EFE2F014CE70CD271" box="[470,552,1633,1659]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF973B0EFEF10186E6F8D2CC" box="[264,988,1707,1734]" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="5B2E36C8FF973B0EFEF10186E6F8D2CC" blockId="10.[264,1323,1707,1894]" box="[264,988,1707,1734]" pageId="10" pageNumber="11">
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<emphasis id="69E5EADAFF973B0EFEF10186E488D2CF" bold="true" box="[264,428,1707,1733]" pageId="10" pageNumber="11">
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<typeStatus id="842A886AFF973B0EFEF10186E461D2CF" box="[264,325,1707,1733]" pageId="10" pageNumber="11">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="9C914D4BFF973B0EFE4D0181E7ECD2CF" box="[436,712,1708,1733]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lutetia" status="sp. nov.">
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<emphasis id="69E5EADAFF973B0EFE4D0181E7ECD2CF" box="[436,712,1708,1733]" italics="true" pageId="10" pageNumber="11">Oblongomorpha lutetia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF973B0EFD360181E634D2CC" box="[719,784,1708,1734]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF973B0EFEC101FEE7A6D31C" pageId="10" pageNumber="11" type="etymology">
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<paragraph id="5B2E36C8FF973B0EFEC101FEE7A6D31C" blockId="10.[264,1323,1707,1894]" pageId="10" pageNumber="11">
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<emphasis id="69E5EADAFF973B0EFEC101FEE706D2E7" bold="true" box="[312,546,1747,1773]" pageId="10" pageNumber="11">Derivatio nominis:</emphasis>
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The new genusgroup name is a reference to the oblong shape of the body. The name is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF973B0FFEC1000EE1FFD6CC" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="5B2E36C8FF973B0FFEC1000EE1FFD6CC" blockId="10.[264,1323,1707,1894]" lastBlockId="11.[264,1324,284,1670]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<emphasis id="69E5EADAFF973B0EFEC1000EE49ED337" bold="true" box="[312,442,1827,1853]" pageId="10" pageNumber="11">Diagnosis:</emphasis>
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Body oblong, surface areolate. Head very short, protruding little in front of the antennal insertion. No cephalic spines or tubercles. Eyes globular, strongly projecting laterally of the head. Antennae long and thin, first antennal segment 3 to 4 times as long as second, 3rd prolonged, 4th clubbed and covered with dense setae. Labium short, not reaching middle of metasternum. Pronotum with a ringlike vesicular at the anterior margin. Paranota absent. Triangular areolate posterior pronotal process is well developed, extremely elongated backwards along hemelytra covering whole clavi and approximately half of discoidal area. Pronotum with 3 longitudinal carinae. Macropterous form. Outline of hemelytra distinctly sinuate about midlength, apical rounded. Costal and subcostal area inconsicuous. Discoidal occupying nearly 0.54 of hemelytron length. Sutural area (membrane) in broadest place with 11–12 of angled cells. Legs moderately long. Tarsi 2segmented, with second segment definitely broader and longer than segment I. Upper surface of segment II slightly convex, lower surface flat and covered with bristlelike hairs.
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</paragraph>
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</subSubSection>
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<subSubSection id="138B6543FF963B0FFEC105F9E719D28C" pageId="11" pageNumber="12" type="discussion">
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<paragraph id="5B2E36C8FF963B0FFEC105F9E6A1D0AC" blockId="11.[264,1324,284,1670]" pageId="11" pageNumber="12">
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<emphasis id="69E5EADAFF963B0FFEC105F9E4ECD6E4" bold="true" box="[312,456,724,750]" pageId="11" pageNumber="12">Comments:</emphasis>
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The development of a celllike hind outgrowth of the pronotum that completely covers the clavi, a short and broad head, and the absence of additional protruding transverse veins on the hemelytra (plesiomorphic!) clearly classify
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<taxonomicName id="9C914D4BFF963B0FFBBF0409E02CD737" box="[1094,1288,804,829]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF963B0FFBBF0409E02CD737" box="[1094,1288,804,829]" italics="true" pageId="11" pageNumber="12">Oblongomorpha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF963B0FFAED0409E41ED76C" pageId="11" pageNumber="12" rank="genus">n. gen.</taxonomicNameLabel>
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in the subfamily
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<taxonomicName id="9C914D4BFF963B0FFDE80461E758D76C" box="[529,636,844,870]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
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. The general structure of the antennae and especially the presence of clearly modified second tarsal segment, with bristlelike hairs in a ventral concavity, unequivocally place
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<taxonomicName id="9C914D4BFF963B0FFDA904B1E636D7BF" box="[592,786,924,949]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF963B0FFDA904B1E636D7BF" box="[592,786,924,949]" italics="true" pageId="11" pageNumber="12">Oblongomorpha</emphasis>
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</taxonomicName>
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in the tribe
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<taxonomicName id="9C914D4BFF963B0FFC5904B1E136D7BC" box="[928,1042,924,950]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Litadeini">
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<emphasis id="69E5EADAFF963B0FFC5904B1E136D7BC" bold="true" box="[928,1042,924,950]" pageId="11" pageNumber="12">Litadeini</emphasis>
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</taxonomicName>
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. Even if the second tarsal segment is not quite as wide as in most of members of the tribe (e.g.
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<taxonomicName id="9C914D4BFF963B0FFB9804E9E19FD7D7" box="[1121,1211,964,989]" class="Insecta" family="Tingidae" genus="Litadea" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="69E5EADAFF963B0FFB9804E9E19FD7D7" box="[1121,1211,964,989]" italics="true" pageId="11" pageNumber="12">Litadea</emphasis>
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</taxonomicName>
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<collectingCountry id="23867658FF963B0FFB3A04E9E02FD7D4" box="[1219,1291,964,990]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
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) it is even elongated and decidedly broader than the first tarsal segment. The decision of describing a new genus for the fossil species instead of including it in the genus
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<taxonomicName id="9C914D4BFF963B0FFB460339E00FD027" box="[1215,1323,1044,1069]" class="Insecta" family="Tingidae" genus="Oecharis" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="69E5EADAFF963B0FFB460339E00FD027" box="[1215,1323,1044,1069]" italics="true" pageId="11" pageNumber="12">Oecharis</emphasis>
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</taxonomicName>
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(only genus among the
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<taxonomicName id="9C914D4BFF963B0FFDDA0311E7A9D05C" box="[547,653,1084,1110]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Litadeini">Litadeini</taxonomicName>
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with the same tarsal modification as in the fossil specimen) is mainly based on the absence of clearly prominent cyst formed by the paranota and the general structure of the hemelytra with small cells.
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</paragraph>
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<paragraph id="5B2E36C8FF963B0FFEC10399E470D14C" blockId="11.[264,1324,284,1670]" pageId="11" pageNumber="12">
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The current classification of this tribe contains 13 extant genera (
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<bibRefCitation id="3F004B39FF963B0FFBD70399E1DCD0C4" author="Froeschner" box="[1070,1272,1204,1230]" pageId="11" pageNumber="18" refString="Froeschner, R. C. (2001) Lace bug genera of the world, II: Subfamily Tinginae: Tribes Litadeini and Ypsotingini (Heteroptera: Tingidae). Smithsonian Contributions to Zoology, 611, 1 - 28." type="journal article" year="2001">Froeschner 2001</bibRefCitation>
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). So far no representative of the
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<taxonomicName id="9C914D4BFF963B0FFDAB03F1E798D0FC" box="[594,700,1244,1270]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Litadeini">Litadeini</taxonomicName>
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has been reported from the fossil record. Lace bugs belonging to the tribe
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<taxonomicName id="9C914D4BFF963B0FFDEF0229E7A4D114" box="[534,640,1284,1310]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" tribe="Litadeini">Litadeini</taxonomicName>
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range to be PanTropical without exception (
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<bibRefCitation id="3F004B39FF963B0FFB5F0229E463D14C" author="Froeschner" pageId="11" pageNumber="18" refString="Froeschner, R. C. (2001) Lace bug genera of the world, II: Subfamily Tinginae: Tribes Litadeini and Ypsotingini (Heteroptera: Tingidae). Smithsonian Contributions to Zoology, 611, 1 - 28." type="journal article" year="2001">Froeschner 2001</bibRefCitation>
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).
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</paragraph>
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<paragraph id="5B2E36C8FF963B0FFEC10279E6D2D23C" blockId="11.[264,1324,284,1670]" pageId="11" pageNumber="12">
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The oldest representative of the
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<taxonomicName id="9C914D4BFF963B0FFD3C0279E608D164" box="[709,812,1364,1390]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
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(
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<emphasis id="69E5EADAFF963B0FFCB80279E10CD167" box="[833,1064,1364,1389]" italics="true" pageId="11" pageNumber="12">Archepopovia yurii</emphasis>
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Golub) has yet been recorded from Baltic amber. Judging on the characteristic features of this specimen the subfamily
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<taxonomicName id="9C914D4BFF963B0FFE730289E4D5D1B4" box="[394,497,1444,1470]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
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with their main features was in the stage of formation during the Eocene (
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<bibRefCitation id="3F004B39FF963B0FFE8902E1E720D1EC" author="Golub" box="[368,516,1484,1510]" pageId="11" pageNumber="18" refString="Golub, V. B. (2001) Archepopovia yurii n. gen. n. sp. a new remarkable lace bug from Baltic amber, with some notes on phylogeny and classification of Tingidae (Heteroptera, Tingidae). Mitteilungen aus dem Geologisch-Palaontologischen Institut der Universitat Hamburg, 85, 263 - 276." type="journal article" year="2001">Golub 2001</bibRefCitation>
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).
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<taxonomicName id="9C914D4BFF963B0FFDE202E1E7F9D1EF" box="[539,733,1484,1509]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF963B0FFDE202E1E7F9D1EF" box="[539,733,1484,1509]" italics="true" pageId="11" pageNumber="12">Oblongomorpha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF963B0FFD1E02E1E61DD1EC" box="[743,825,1484,1510]" pageId="11" pageNumber="12" rank="genus">n. gen.</taxonomicNameLabel>
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and
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<taxonomicName id="9C914D4BFF963B0FFC8F02E1E109D1EF" box="[886,1069,1484,1509]" class="Insecta" family="Tingidae" genus="Chorotingioites" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF963B0FFC8F02E1E109D1EF" box="[886,1069,1484,1509]" italics="true" pageId="11" pageNumber="12">Chorotingioites</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="72D657A1FF963B0FFBCE02E1E1AED1EC" box="[1079,1162,1484,1510]" pageId="11" pageNumber="12" rank="genus">n. gen.</taxonomicNameLabel>
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clearly show that the differentiation of the
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<taxonomicName id="9C914D4BFF963B0FFD8B02D9E7FDD204" box="[626,729,1524,1550]" class="Insecta" family="Tingidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Tinginae">Tinginae</taxonomicName>
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has taken place in this period or even earlier as already supposed by
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<bibRefCitation id="3F004B39FF963B0FFE060131E785D23C" author="Golub" box="[511,673,1564,1590]" pageId="11" pageNumber="18" refString="Golub, V. B. (2001) Archepopovia yurii n. gen. n. sp. a new remarkable lace bug from Baltic amber, with some notes on phylogeny and classification of Tingidae (Heteroptera, Tingidae). Mitteilungen aus dem Geologisch-Palaontologischen Institut der Universitat Hamburg, 85, 263 - 276." type="journal article" year="2001">Golub (2001)</bibRefCitation>
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, and
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<bibRefCitation id="3F004B39FF963B0FFD270131E6D6D23C" author="Golub" box="[734,1010,1564,1590]" pageId="11" pageNumber="18" refString="Golub, V. B. & Popov, Y. A. (1999) Composition and evolution of Cretaceous and Cenozoic faunas of bugs of the superfamily Tingoidea (Heteroptera: Cimicomorpha). AMBA / AM / PFICM 98 / 1.99: Proceedings of the First International Palaeoentomological Conference, Moscow 1998, 33 - 39." type="book chapter" year="1999">Golub & Popov (1999)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="5B2E36C8FF963B0FFEC10169E719D28C" blockId="11.[264,1324,284,1670]" pageId="11" pageNumber="12">
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A reconstruction of the
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<typeStatus id="842A886AFF963B0FFD9F0169E7BCD254" box="[614,664,1604,1630]" pageId="11" pageNumber="12">type</typeStatus>
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species of
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<taxonomicName id="9C914D4BFF963B0FFCD40169E6CBD257" box="[813,1007,1604,1629]" class="Insecta" family="Tingidae" genus="Oblongomorpha" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="69E5EADAFF963B0FFCD40169E6CBD257" box="[813,1007,1604,1629]" italics="true" pageId="11" pageNumber="12">Oblongomorpha</emphasis>
|
||
</taxonomicName>
|
||
based on the description below is shown in
|
||
<figureCitation id="C3AA2A4DFF963B0FFE1D0141E71DD28C" box="[484,569,1644,1670]" captionStart="FIGURE 11" captionStartId="12.[264,367,1810,1834]" captionTargetBox="[360,1234,313,1795]" captionTargetId="figure@12.[336,1250,294,1806]" captionTargetPageId="12" captionText="FIGURE 11. Oblongomorpha lutetia gen. et sp. n., reconstruction of the holotype, (macropterous form), compression fossil; coll. Forschungsinstitut Senckenberg (SMF), Frankfurt am Main (Germany), specimen SMF MeI 10508; scale bar represents 1 mm." httpUri="https://zenodo.org/record/156773/files/figure.png" pageId="11" pageNumber="12">Fig. 11</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |