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<document id="79A24234D9B4DC4794A90ADAFD581478" ID-CLB-Dataset="22845" ID-DOI="10.11646/zootaxa.4766.2.7" ID-GBIF-Dataset="d7bc2a71-cb55-464e-af0b-533efae08e61" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763525" ID-ZooBank="urn:lsid:zoobank.org:pub:03494EDD-FE6E-4DD1-B137-163A4B266DEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587656660933" checkinUser="carolina" docAuthor="Tri, Ngo Van, Grismer, L. Lee, Thai, Pham Hong &amp; Jr, P. L. Wood" docDate="2020" docId="D325CF056D4B9144FF2F65E1DBB7D752" docLanguage="en" docName="zootaxa.4766.2.7.pdf.imf" docOrigin="Zootaxa 4766 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus culaochamensis Tri &amp; Grismer &amp; Thai &amp; Jr 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="399" masterDocId="2F1CB77D6D48914EFFB8630CDB40D365" masterDocTitle="A new endemic insular Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodctylus) from Quang Nam Province, Central Vietnam" masterLastPageNumber="400" masterPageNumber="389" pageNumber="392" updateTime="1698835666781" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="CDD600EB8629B8684CE4C44348889D4F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EF7E8EDB382FE94CECD013EE3D5936BF">
<mods:title id="0724EBDC01AC64CC3B0F9BACEB8AC944">A new endemic insular Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodctylus) from Quang Nam Province, Central Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="28235FA1F214937390DF452D2F506F48" type="personal">
<mods:role id="C7DA5ACD9BA63EAA045497AEA5D2522D">
<mods:roleTerm id="D18BE9BF0F2954C0D7D1ABAAB5CFEDF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D61B4A7974665B7D5079F261E29AA4D">Tri, Ngo Van</mods:namePart>
</mods:name>
<mods:name id="0A9FA8B3B7EE75F806E651A22B72D6BB" type="personal">
<mods:role id="9DE94E66597B865C7B435ACB0CB5BCBD">
<mods:roleTerm id="4E20816D9268FB0B522AA377BF0FAA41">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB5AB68435DCDD406617C488AEDE8C18">Grismer, L. Lee</mods:namePart>
</mods:name>
<mods:name id="24DC3C382862CDEB458B53B67C96EEC9" type="personal">
<mods:role id="B41CEBA47F8261AC0BC592684A72023B">
<mods:roleTerm id="1C428E699C2088C5E385871D97AE8B00">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0EA9D785C2D197D7A41F65FDF4F08238">Thai, Pham Hong</mods:namePart>
</mods:name>
<mods:name id="BE452CD1308580A1066151A8CD01835B" type="personal">
<mods:role id="4D04A7344D3ECC1463EAE94D14160D60">
<mods:roleTerm id="878F9FF269B0F15FD10DCDA0EF390B12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="84F0D9856801B7E3A65F42444C960C7D">Jr, P. L. Wood</mods:namePart>
</mods:name>
<mods:typeOfResource id="190ED2FDCB086C1C3C7E55AFF4B919CF">text</mods:typeOfResource>
<mods:relatedItem id="4A091CD0EAC0D1EFB1BC85ABD4070E23" type="host">
<mods:titleInfo id="21D4968E77C6E8FF0EB0AAF51AFFF6F1">
<mods:title id="46AF54F20B2CC9E303EA129E5B050B33">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BE4990313AE51D63C8351028F4C76B7C">
<mods:date id="20BE62918E94F1285437FF8E048A0279">2020</mods:date>
<mods:detail id="49AF4FA4BB91F7695556A2AF77D88BAF" type="pubDate">
<mods:number id="472384A5C264918FB72401C11F76FC6C">2020-04-20</mods:number>
</mods:detail>
<mods:detail id="DF0182E6E3065FEB8FCE7ACA5F884932" type="volume">
<mods:number id="853E6D44E0A872D291457853974FC81B">4766</mods:number>
</mods:detail>
<mods:detail id="061F8205B8E2A3D9DE165BB678F43EF9" type="issue">
<mods:number id="791D62B2C5711000463A83C951330238">2</mods:number>
</mods:detail>
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<mods:start id="498705B71A3A3DCDD0B55316E9FC3746">389</mods:start>
<mods:end id="B7274B86F55C39EC427934F5E3A1B01A">400</mods:end>
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<mods:classification id="38DD96385C3D42C8B09D0F2F8AA031EF">journal article</mods:classification>
<mods:identifier id="788D0344606C49ECE0F6905601D89562" type="CLB-Dataset">22845</mods:identifier>
<mods:identifier id="F1D1CF1262BDA26CDC3A0E24C7B7C67B" type="DOI">10.11646/zootaxa.4766.2.7</mods:identifier>
<mods:identifier id="1B62EF8D489CA77E01AC1B0EB81641F9" type="GBIF-Dataset">d7bc2a71-cb55-464e-af0b-533efae08e61</mods:identifier>
<mods:identifier id="AAEBB2394A411C1FA17E36B4C145AC6E" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="2B3E019DC636A8D7B9706830D54AD86C" type="Zenodo-Dep">3763525</mods:identifier>
<mods:identifier id="7DEADE21E9C89F10A45FBD70F7AC3745" type="ZooBank">urn:lsid:zoobank.org:pub:03494EDD-FE6E-4DD1-B137-163A4B266DEB</mods:identifier>
</mods:mods>
<treatment id="D325CF056D4B9144FF2F65E1DBB7D752" ID-DOI="http://doi.org/10.5281/zenodo.3804125" ID-GBIF-Taxon="163736831" ID-Zenodo-Dep="3804125" LSID="urn:lsid:plazi:treatment:D325CF056D4B9144FF2F65E1DBB7D752" httpUri="http://treatment.plazi.org/id/D325CF056D4B9144FF2F65E1DBB7D752" lastPageId="10" lastPageNumber="399" pageId="3" pageNumber="392">
<subSubSection id="13962D986D4B914DFF2F65E1D92CD462" box="[151,620,1773,1800]" pageId="3" pageNumber="392" type="nomenclature">
<paragraph id="5B337E136D4B914DFF2F65E1D92CD462" blockId="3.[151,620,1773,1835]" box="[151,620,1773,1800]" pageId="3" pageNumber="392">
<heading id="007BC97F6D4B914DFF2F65E1D92CD462" bold="true" box="[151,620,1773,1800]" fontSize="10" level="2" pageId="3" pageNumber="392" reason="2">
<emphasis id="69F8A2016D4B914DFF2F65E1D92CD462" bold="true" box="[151,620,1773,1800]" pageId="3" pageNumber="392">
<taxonomicName id="9C8C05906D4B914DFF2F65E1D947D46D" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[151,519,1773,1800]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="392" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D4B914DFF2F65E1D947D46D" bold="true" box="[151,519,1773,1800]" italics="true" pageId="3" pageNumber="392">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D4B914DFDB665E1D92CD462" box="[526,620,1773,1799]" pageId="3" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13962D986D4B914DFF2F641DDBB8D44E" box="[151,248,1809,1835]" pageId="3" pageNumber="392" type="description">
<paragraph id="5B337E136D4B914DFF2F641DDBB8D44E" blockId="3.[151,620,1773,1835]" box="[151,248,1809,1835]" pageId="3" pageNumber="392">
<figureCitation id="C3B762966D4B914DFF2F641DDB9FD44E" box="[151,223,1809,1835]" captionStart="FIGURE 1" captionStartId="4.[151,255,1381,1405]" captionTargetBox="[193,1394,183,1342]" captionTargetId="figure@4.[193,1394,180,1354]" captionTargetPageId="4" captionText="FIGURE 1. Inferred phylogenetic relationships Cyrtodactylus culaochamensis sp. nov. The tree is a maximum likelihood topology with BPP and ML nodal support, respectively." figureDoi="http://doi.org/10.5281/zenodo.3763527" httpUri="https://zenodo.org/record/3763527/files/figure.png" pageId="3" pageNumber="392">Figs.1</figureCitation>
,
<figureCitation id="C3B762966D4B914DFF52641DDBB8D44E" box="[234,248,1809,1835]" captionStart="FIGURE 2" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[232,1356,181,1919]" captionTargetId="figure@5.[232,1356,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cyrtodactylus culaochamensis sp. nov. A: Holotype ITBCZ 2494; B: paratype ITBCZ 2495 showing the irregularly shaped dorsal bands and blotches; C: paratype ITBCZ 2498 showing the thin nuchal loop and full tail with 13 light-colored bands alternating with dark-brown bands." figureDoi="http://doi.org/10.5281/zenodo.3763529" httpUri="https://zenodo.org/record/3763529/files/figure.png" pageId="3" pageNumber="392">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="13962D986D4B914AFF2F6455DFF8D56B" lastPageId="4" lastPageNumber="393" pageId="3" pageNumber="392" type="materials_examined">
<paragraph id="5B337E136D4B914DFF2F6455DE72D4DE" blockId="3.[151,1436,1880,1979]" pageId="3" pageNumber="392">
<materialsCitation id="EBE4744E6D4B914DFF2F6455DE72D4DE" ID-GBIF-Occurrence="2608526688" collectingDate="2014-02-15" collectionCode="ITBCZ" collectorName="Ngo Van Tri &amp; N. V." country="Vietnam" county="Central" elevation="50" latitude="15.95" location="Hon Lao Island" longLatPrecision="1283" longitude="108.5" municipality="Hoi An City" pageId="3" pageNumber="392" specimenCode="ITBCZ 2494" specimenCount="1" stateProvince="Quang Nam Province" typeStatus="holotype">
<emphasis id="69F8A2016D4B914DFF2F6455DA4BD416" bold="true" box="[151,267,1881,1907]" pageId="3" pageNumber="392">
<typeStatus id="8437C0B16D4B914DFF2F6455DA46D416" box="[151,262,1881,1907]" pageId="3" pageNumber="392" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="0B2AD6686D4B914DFEAA6455DAE5D416" box="[274,421,1881,1907]" pageId="3" pageNumber="392">ITBCZ 2494</specimenCode>
,
<specimenCount id="4D8AB59A6D4B914DFE086455D965D416" box="[432,549,1881,1907]" pageId="3" pageNumber="392" type="adult">adult male</specimenCount>
collected by
<collectorName id="F6791BC56D4B914DFD046455D806D416" box="[700,838,1881,1907]" pageId="3" pageNumber="392">Ngo Van Tri</collectorName>
(
<collectorName id="F6791BC56D4B914DFCEE6455D8CAD416" box="[854,906,1881,1907]" name="Ngo Van Tri" pageId="3" pageNumber="392">N.V.</collectorName>
T) at 20:30 on
<date id="2F3258D36D4B914DFB896454DFB8D416" box="[1073,1272,1880,1907]" pageId="3" pageNumber="392" value="2014-02-15">
<collectingDate id="3F76A13B6D4B914DFB896454DFB8D416" box="[1073,1272,1880,1907]" pageId="3" pageNumber="392" value="2014-02-15">15 February 2014</collectingDate>
</date>
on the surface of a large rock along a small stream in
<location id="5E5328C86D4B914DFDFE6471D9B5D4F2" LSID="urn:lsid:plazi:treatment:D325CF056D4B9144FF2F65E1DBB7D752:5E5328C86D4B914DFDFE6471D9B5D4F2" box="[582,757,1917,1943]" country="Vietnam" county="Central" latitude="15.95" longLatPrecision="1283" longitude="108.5" municipality="Hoi An City" name="Hon Lao Island" pageId="3" pageNumber="392" stateProvince="Quang Nam Province">Hon Lao Island</location>
,
<location id="5E5328C86D4B914DFCB86471DF37D4F2" LSID="urn:lsid:plazi:treatment:D325CF056D4B9144FF2F65E1DBB7D752:5E5328C86D4B914DFCB86471DF37D4F2" box="[768,1143,1917,1943]" country="Vietnam" county="Central" latitude="15.95" longLatPrecision="1283" longitude="108.5" municipality="Hoi An City" name="Cu Lao Cham Biosphere Reserve" pageId="3" pageNumber="392" stateProvince="Quang Nam Province">Cu Lao Cham Biosphere Reserve</location>
,
<location id="5E5328C86D4B914DFB396471DF8FD4F2" LSID="urn:lsid:plazi:treatment:D325CF056D4B9144FF2F65E1DBB7D752:5E5328C86D4B914DFB396471DF8FD4F2" box="[1153,1231,1917,1943]" country="Vietnam" county="Central" latitude="15.95" longLatPrecision="1283" longitude="108.5" municipality="Hoi An City" name="Tan Hi" pageId="3" pageNumber="392" stateProvince="Quang Nam Province">Tân Hi</location>
ệp
<location id="5E5328C86D4B914DFB486471DE28D4F2" LSID="urn:lsid:plazi:treatment:D325CF056D4B9144FF2F65E1DBB7D752:5E5328C86D4B914DFB486471DE28D4F2" box="[1264,1384,1917,1943]" country="Vietnam" county="Central" latitude="15.95" longLatPrecision="1283" longitude="108.5" municipality="Hoi An City" name="Commune" pageId="3" pageNumber="392" stateProvince="Quang Nam Province">Commune</location>
,
<collectingMunicipality id="BB57E4696D4B914DFACA6471DBB1D4DE" pageId="3" pageNumber="392">Hoi An City</collectingMunicipality>
,
<collectingRegion id="9948B0F16D4B914DFF4564ADDAB2D4DE" box="[253,498,1953,1979]" country="Vietnam" name="Quang Nam" pageId="3" pageNumber="392">Quang Nam Province</collectingRegion>
,
<collectingCounty id="B252069F6D4B914DFE4564ADD90FD4DE" box="[509,591,1953,1979]" pageId="3" pageNumber="392">Central</collectingCounty>
<collectingCountry id="239B3E836D4B914DFDEE64ADD9F5D4DE" box="[598,693,1953,1979]" name="Vietnam" pageId="3" pageNumber="392">Vietnam</collectingCountry>
(around of
<geoCoordinate id="3EB818D46D4B914DFC8064ACD8DDD4DE" box="[824,925,1952,1979]" degrees="15" direction="north" minutes="57" orientation="latitude" pageId="3" pageNumber="392" precision="925" value="15.95">
15
<superScript id="ACF9D35B6D4B914DFCEC64ACD819D4CB" attach="left" box="[852,857,1952,1966]" fontSize="6" pageId="3" pageNumber="392">º</superScript>
57’N
</geoCoordinate>
,
<geoCoordinate id="3EB818D46D4B914DFC1064ADDF55D4DE" box="[936,1045,1952,1979]" degrees="108" direction="east" minutes="30" orientation="longitude" pageId="3" pageNumber="392" precision="925" value="108.5">
108
<superScript id="ACF9D35B6D4B914DFC6A64ACD897D4CB" attach="left" box="[978,983,1952,1966]" fontSize="6" pageId="3" pageNumber="392">º</superScript>
30’E
</geoCoordinate>
) at
<elevation id="D0A199206D4B914DFBF864ACDE6FD4DE" box="[1088,1327,1952,1979]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="392" unit="m" value="50.0">
<quantity id="9C74D3F66D4B914DFBF864ACDF39D4DE" box="[1088,1145,1952,1979]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="392" unit="m" value="50.0">50 m</quantity>
above sea level
</elevation>
.
</materialsCitation>
</paragraph>
<caption id="0FF32E9B6D4C914AFF2F6669D9A9D6C7" ID-DOI="http://doi.org/10.5281/zenodo.3763527" ID-Zenodo-Dep="3763527" httpUri="https://zenodo.org/record/3763527/files/figure.png" pageId="4" pageNumber="393" startId="4.[151,255,1381,1405]" targetBox="[193,1394,183,1342]" targetPageId="4">
<paragraph id="5B337E136D4C914AFF2F6669D9A9D6C7" blockId="4.[151,1436,1381,1442]" pageId="4" pageNumber="393">
<emphasis id="69F8A2016D4C914AFF2F6669DA58D618" bold="true" box="[151,280,1381,1405]" pageId="4" pageNumber="393">FIGURE 1.</emphasis>
Inferred phylogenetic relationships
<taxonomicName id="9C8C05906D4C914AFD2F666AD89CD61B" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[663,988,1382,1406]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D4C914AFD2F666AD89CD61B" box="[663,988,1382,1406]" italics="true" pageId="4" pageNumber="393">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D4C914AFC5A666ADF74D618" box="[994,1076,1382,1405]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
The tree is a maximum likelihood topology with BPP and ML nodal support, respectively.
</paragraph>
</caption>
<paragraph id="5B337E136D4C914AFF7F66DCDFF8D56B" blockId="4.[151,1437,1487,2019]" pageId="4" pageNumber="393">
<emphasis id="69F8A2016D4C914AFF7F66DCDA06D68F" bold="true" box="[199,326,1488,1514]" pageId="4" pageNumber="393">
<typeStatus id="8437C0B16D4C914AFF7F66DCDA01D68F" box="[199,321,1488,1514]" pageId="4" pageNumber="393" type="paratype">Paratypes</typeStatus>
.
</emphasis>
The
<typeStatus id="8437C0B16D4C914AFE3D66DCDAB0D68F" box="[389,496,1488,1514]" pageId="4" pageNumber="393" type="paratype">paratypes</typeStatus>
ITBCZ 2495,
<date id="2F3258D36D4C914AFD2566DCD840D68F" box="[669,768,1488,1514]" pageId="4" pageNumber="393" value="2497" valueMax="2498">2497–98</date>
bear the collection data as the
<typeStatus id="8437C0B16D4C914AFBD566DCDF93D68F" box="[1133,1235,1488,1514]" pageId="4" pageNumber="393" type="holotype">holotype</typeStatus>
.
<typeStatus id="8437C0B16D4C914AFB5866DCDE02D68F" box="[1248,1346,1488,1514]" pageId="4" pageNumber="393" type="paratype">Paratype</typeStatus>
ITBCZ 2496 was collected by Phạm Hồng Thái (P.H.T.) at the same locality but on
<date id="2F3258D36D4C914AFC5266F8DFF5D56B" box="[1002,1205,1524,1550]" pageId="4" pageNumber="393" value="2014-02-17">17 February 2014</date>
.
</paragraph>
</subSubSection>
<subSubSection id="13962D986D4C914AFF7F651BDAFFD437" pageId="4" pageNumber="393" type="diagnosis">
<paragraph id="5B337E136D4C914AFF7F651BDAFFD437" blockId="4.[151,1437,1487,2019]" pageId="4" pageNumber="393">
<emphasis id="69F8A2016D4C914AFF7F651BDA02D557" bold="true" box="[199,322,1559,1586]" pageId="4" pageNumber="393">Diagnosis.</emphasis>
<taxonomicName id="9C8C05906D4C914AFEF16514D9E0D557" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[329,672,1560,1586]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="393" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D4C914AFEF16514D9E0D557" box="[329,672,1560,1586]" italics="true" pageId="4" pageNumber="393">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<emphasis id="69F8A2016D4C914AFD1F6514D841D557" bold="true" box="[679,769,1560,1586]" pageId="4" pageNumber="393">
<taxonomicNameLabel id="72CB1F7A6D4C914AFD1F6514D841D557" box="[679,769,1560,1586]" pageId="4" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other congeners by the following combination of characters: maximum SVL
<quantity id="9C74D3F66D4C914AFDA06530D93BD533" box="[536,635,1596,1622]" metricMagnitude="-2" metricUnit="m" metricValue="7.9799999999999995" pageId="4" pageNumber="393" unit="mm" value="79.8">79.8 mm</quantity>
(n=5); original tail length (TL/SVL:1.14–1.31); dark spots and blotches on head edged in lighter coloration; nuchal loop usually fragmented; five or six irregularly shaped, dark, broad, broken dorsal bands; original tail bearing 10–13 dark-brown alternating rings; 2–5 intersupranasals; 11–13 supralabials; nine or 10 infralabials; seven or eight precloacal pores in males; no precloacal pores in females; no enlarged scales beneath thighs; 19–21 interorbitals; 28–32 scales between eyeball and nostril; 45–50 rows of ventral scales; 20–22 irregular, longitudinal rows of smooth and weakly keeled midbody tubercles; 27–32 paravertebral tubercles between limb insertions; 13–15 subdigital lamellae on first toe; 20–23 subdigital lamellae on fourth toe; and slightly enlarged subcaudal scales.
</paragraph>
</subSubSection>
<subSubSection id="13962D986D4C9144FF7F6457DAC0D27A" lastPageId="10" lastPageNumber="399" pageId="4" pageNumber="393" type="description">
<paragraph id="5B337E136D4C9149FF7F6457DE7DD39E" blockId="4.[151,1437,1487,2019]" lastBlockId="7.[151,1437,153,539]" lastPageId="7" lastPageNumber="396" pageId="4" pageNumber="393">
<emphasis id="69F8A2016D4C914AFF7F6457D914D413" bold="true" box="[199,596,1883,1910]" pageId="4" pageNumber="393">
Description of
<typeStatus id="8437C0B16D4C914AFECE6450DA9DD413" box="[374,477,1884,1910]" pageId="4" pageNumber="393" type="holotype">holotype</typeStatus>
(
<figureCitation id="C3B762966D4C914AFE536457D905D413" box="[491,581,1883,1910]" captionStart="FIGURE 1" captionStartId="4.[151,255,1381,1405]" captionTargetBox="[193,1394,183,1342]" captionTargetId="figure@4.[193,1394,180,1354]" captionTargetPageId="4" captionText="FIGURE 1. Inferred phylogenetic relationships Cyrtodactylus culaochamensis sp. nov. The tree is a maximum likelihood topology with BPP and ML nodal support, respectively." figureDoi="http://doi.org/10.5281/zenodo.3763527" httpUri="https://zenodo.org/record/3763527/files/figure.png" pageId="4" pageNumber="393">Fig. 1A</figureCitation>
).
</emphasis>
Adult male, SVL
<quantity id="9C74D3F66D4C914AFCA76450D83FD413" box="[799,895,1884,1910]" metricMagnitude="-2" metricUnit="m" metricValue="7.74" pageId="4" pageNumber="393" unit="mm" value="77.4">77.4mm</quantity>
. Head moderately long (HeadL/SVL:0.27), relatively narrow (HeadW/HeadL:0.67), depressed (HeadH/HeadL: 0.34), distinct from neck; lores and interorbital region inflated, canthus rostralis absent, frontonasal region concave; snout elongate (SnEye/HeadL: 0.37), pointed, longer than eye diameter (OrbD/SnEye: 0.68); scales on snout small, rounded, granular, homogeneous, larger than those on occipital region. Eye large (OrbD/HeadL:0.26), iris chestnut with red crenelated pupilar margins; supraciliaries short, bearing tiny conical spines posteriorly. Ear opening oval, oblique, small (EarL/HeadL: 0.10); eye to ear distance greater than diameter of eye (EyeEar/OrbD: 1.02). Rostral scale smooth, wider (
<quantity id="9C74D3F66D4E9148FB246454DFB3D416" box="[1180,1267,1880,1907]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="395" unit="mm" value="2.3">2.3 mm</quantity>
) than high (
<quantity id="9C74D3F66D4E9148FAC0645BDB87D4F2" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="395" unit="mm" value="1.5">1.5 mm</quantity>
), incompletely divided posteriorly by a shallow Y-shaped dorsal groove; two enlarged supranasals separated by one large intersupranasal; rostral in contact with first supralabial and nostril; nostril oval, surrounded by supranasal, rostral, first supralabial, and three enlarged postnasals; three or four rows of small scales separate orbit from supral- abials. Mental triangular, wider (
<quantity id="9C74D3F66D4F9149FDB96395D913D3D6" box="[513,595,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="396" unit="mm" value="2.7">2.7mm</quantity>
) than deep (
<quantity id="9C74D3F66D4F9149FD626395D86FD3D1" box="[730,815,153,180]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="396" unit="mm" value="1.6">1.6mm</quantity>
); one pair of enlarged postmentals, in broad contact medially, bordered laterally by first infralabial and first chinshield, posterolaterally by four small gular scales. Thirteen supralabials and 10 infralabials on each side; 20 interorbital scale rows; 32 scales between eye and nostril.
</paragraph>
<caption id="0FF32E9B6D4D914BFF2F6495D97ED49F" ID-DOI="http://doi.org/10.5281/zenodo.3763529" ID-Zenodo-Dep="3763529" httpUri="https://zenodo.org/record/3763529/files/figure.png" pageId="5" pageNumber="394" startId="5.[151,250,1945,1970]" targetBox="[232,1356,181,1919]" targetPageId="5">
<paragraph id="5B337E136D4D914BFF2F6495D97ED49F" blockId="5.[151,1436,1945,2042]" pageId="5" pageNumber="394">
<emphasis id="69F8A2016D4D914BFF2F6495DA54D4D7" bold="true" box="[151,276,1945,1970]" pageId="5" pageNumber="394">FIGURE 2.</emphasis>
<taxonomicName id="9C8C05906D4D914BFEA36496D913D4D7" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[283,595,1946,1970]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="394" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D4D914BFEA36496D913D4D7" box="[283,595,1946,1970]" italics="true" pageId="5" pageNumber="394">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D4D914BFDE26496D9EDD4D4" box="[602,685,1946,1970]" pageId="5" pageNumber="394" rank="species">
<emphasis id="69F8A2016D4D914BFDE26496D9E7D4D7" bold="true" box="[602,679,1946,1970]" pageId="5" pageNumber="394">sp. nov</emphasis>
.
</taxonomicNameLabel>
A: Holotype ITBCZ 2494; B: paratype ITBCZ 2495 showing the irregularly shaped dorsal bands and blotches; C: paratype ITBCZ 2498 showing the thin nuchal loop and full tail with 13 light-colored bands alternating with dark-brown bands.
</paragraph>
</caption>
<caption id="0FF32E9B6D4E9148FF2F658DD822D587" ID-DOI="http://doi.org/10.5281/zenodo.3763531" ID-Zenodo-Dep="3763531" httpUri="https://zenodo.org/record/3763531/files/figure.png" pageId="6" pageNumber="395" startId="6.[151,250,1665,1690]" targetBox="[151,1436,181,1640]" targetPageId="6">
<paragraph id="5B337E136D4E9148FF2F658DD822D587" blockId="6.[151,1436,1665,1762]" pageId="6" pageNumber="395">
<emphasis id="69F8A2016D4E9148FF2F658DDA55D5FF" bold="true" box="[151,277,1665,1690]" pageId="6" pageNumber="395">FIGURE 3.</emphasis>
<taxonomicName id="9C8C05906D4E9148FEAD658ED90ED5FF" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[277,590,1666,1690]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="395" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D4E9148FEAD658ED90ED5FF" box="[277,590,1666,1690]" italics="true" pageId="6" pageNumber="395">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D4E9148FDEE658ED9EAD5FC" box="[598,682,1666,1690]" pageId="6" pageNumber="395" rank="species">
<emphasis id="69F8A2016D4E9148FDEE658ED9E4D5FF" bold="true" box="[598,676,1666,1690]" pageId="6" pageNumber="395">sp. nov</emphasis>
.
</taxonomicNameLabel>
, Holotype ITBCZ 2494, A: Rostral with one large internasal; B: Mental and two enlarged postmentals; C: Dorsum with weakly keeked tubercles; D: Subimbricate ventral scales; E: dorsal surface of a weakly segmented tail lacking tubercles; F: enlarged subcaudal scales.
</paragraph>
</caption>
<paragraph id="5B337E136D4F9149FF7F6209DA56D17E" blockId="7.[151,1437,153,539]" pageId="7" pageNumber="396">Body slender, elongate (TrunkL/SVL: 0.42). Dorsal scales conical; regularly distributed weakly keeled tubercles (4–6 times size of adjacent scales), extend from occiput to base of tail; tubercles arranged in 22 irregular rows at midbody, smallest on flanks and in occipital region; 27 paravertebral tubercles. Ventral scales larger than dorsals, smooth, relatively round, subimbricate, largest posteriorly; 50 ventral scale rows with smooth tubercles in scattering on lateral folds; gular region bearing relatively homogeneous, smooth scales. Precloacal groove absent; precloacal scales enlarged; a series of seven precloacal pores in a chevron pattern; no enlarged femoral scales beneath each thigh. Scales on palm and hind limbs granular with scattered, weakly keeled tubercles that are smaller than those on dorsum.</paragraph>
<caption id="0FF32E9B6D4F9149FF2F67E7D922D629" ID-DOI="http://doi.org/10.5281/zenodo.3763535" ID-Zenodo-Dep="3763535" httpUri="https://zenodo.org/record/3763535/files/figure.png" pageId="7" pageNumber="396" startId="7.[151,250,1259,1284]" targetBox="[177,1411,561,1235]" targetPageId="7">
<paragraph id="5B337E136D4F9149FF2F67E7D922D629" blockId="7.[151,1437,1259,1356]" pageId="7" pageNumber="396">
<emphasis id="69F8A2016D4F9149FF2F67E7DA53D661" bold="true" box="[151,275,1259,1284]" pageId="7" pageNumber="396">FIGURE 4.</emphasis>
A: Ventral view of tail of
<taxonomicName id="9C8C05906D4F9149FDA667E0D815D661" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[542,853,1260,1284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D4F9149FDA667E0D815D661" box="[542,853,1260,1284]" italics="true" pageId="7" pageNumber="396">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D4F9149FCE367E0D8EED661" box="[859,942,1260,1284]" pageId="7" pageNumber="396" rank="species">
<emphasis id="69F8A2016D4F9149FCE367E0D8E7D661" bold="true" box="[859,935,1260,1284]" pageId="7" pageNumber="396">sp. nov</emphasis>
.
</taxonomicNameLabel>
, ITBCZ 2494, showing light-colored, subcaudal rings and very slightly enlarged subcaudals; B: Precloacal region of
<taxonomicName id="9C8C05906D4F9149FCEF661DDF58D64D" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[855,1048,1296,1320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="396" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D4F9149FCEF661DDF58D64D" box="[855,1048,1296,1320]" italics="true" pageId="7" pageNumber="396">C. culaochamensis</emphasis>
</taxonomicName>
<emphasis id="69F8A2016D4F9149FBA7661CDF2CD64D" bold="true" box="[1055,1132,1296,1320]" pageId="7" pageNumber="396">
<taxonomicNameLabel id="72CB1F7A6D4F9149FBA7661CDF2CD64D" box="[1055,1132,1296,1320]" pageId="7" pageNumber="396" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
ITBCZ 2494, showing seven precloacal cloacal pores in a chevron pattern.
</paragraph>
</caption>
<paragraph id="5B337E136D4F9149FF7F6677DAFAD508" blockId="7.[151,1436,1402,2041]" pageId="7" pageNumber="396">Fore and hind limbs moderately slender (ForeL/SVL: 0.14; CrusL/SVL: 0.16); digits moderately slender, strongly inflected at basal interphalangeal joints, all bearing slightly curved claws; basal subdigital lamellae nearly as broad as digit and lacking a scansorial surface: 5–6–7–7–7 manus; 5–8–8–9–9 pes; narrow lamellae distal to digital inflection and not including ventral claw sheath: 8–7–10–10–7 manus; 7–9–12–12–11 pes; one or two rows of small, non-lamellar granules between basal and distal lamellar series; interdigital webbing present but weakly developed. Length of digits in mm: (manus)III (6.9)&gt;IV (6.0)&gt; II (4.7)&gt; V (4.5)&gt;I (3.6); (pes):IV (7.3)&gt; V (7.2)&gt; III (7.2)&gt; II (5.8)&gt; I (3.7).</paragraph>
<paragraph id="5B337E136D4F9149FF7F657ADAF0D445" blockId="7.[151,1436,1402,2041]" pageId="7" pageNumber="396">
Original tail length
<quantity id="9C74D3F66D4F9149FE27657BD942D5F4" box="[415,514,1655,1681]" metricMagnitude="-2" metricUnit="m" metricValue="9.07" pageId="7" pageNumber="396" unit="mm" value="90.7">90.7 mm</quantity>
(TL/SVL: 1.27), slender, tapering to a point; longer than SVL; two smooth, whitish postcloacal tubercles at base; base of tail bearing five parasagittal and three longitudinal rows of keeled tubercles, followed by two rows of smaller tubercles on the first four segments of tail, decreasing to one row towards the posterior segment and absent posteriorly to that; subcaudals smooth, subimbricate, all subcaudals very slightly enlarged throughout length of tail.
</paragraph>
<paragraph id="5B337E136D4F9149FF7F6426DE70D49C" blockId="7.[151,1436,1402,2041]" pageId="7" pageNumber="396">
<emphasis id="69F8A2016D4F9149FF7F6426DAB6D420" bold="true" box="[199,502,1834,1861]" pageId="7" pageNumber="396">
Coloration in life (
<figureCitation id="C3B762966D4F9149FE196426DAA7D420" box="[417,487,1834,1861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[232,1356,181,1919]" captionTargetId="figure@5.[232,1356,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cyrtodactylus culaochamensis sp. nov. A: Holotype ITBCZ 2494; B: paratype ITBCZ 2495 showing the irregularly shaped dorsal bands and blotches; C: paratype ITBCZ 2498 showing the thin nuchal loop and full tail with 13 light-colored bands alternating with dark-brown bands." figureDoi="http://doi.org/10.5281/zenodo.3763529" httpUri="https://zenodo.org/record/3763529/files/figure.png" pageId="7" pageNumber="396">Fig. 2</figureCitation>
).
</emphasis>
Top of head brown with dark blotches or spots edged in lighter coloration; chestnut iris with crenelated vertical pupil, eye rings yellowish; nuchal loop fragmented; two irregularly shaped, dark bands anterior to forelimb insertions that grade into four broken bands posteriorly that resemble transversely arranged, squarish blotches; small dark blotches scattered on flanks; dorsal surface of limbs brown, bearing irregularly shaped darker bands; all dark markings on body and limbs are edged with light-colored tubercles; original tail bearing 10 dark and light-brown alternating rings. Ventral surface of body pinkish bearing scattered yellowish spots.
</paragraph>
<caption id="0FF32E9B6D409146FF2F64A5D954D483" ID-DOI="http://doi.org/10.5281/zenodo.3763537" ID-Zenodo-Dep="3763537" httpUri="https://zenodo.org/record/3763537/files/figure.png" pageId="8" pageNumber="397" startId="8.[151,250,1961,1986]" targetBox="[164,1423,181,1936]" targetPageId="8">
<paragraph id="5B337E136D409146FF2F64A5D954D483" blockId="8.[151,1437,1961,2022]" pageId="8" pageNumber="397">
<emphasis id="69F8A2016D409146FF2F64A5DA52D4A7" bold="true" box="[151,274,1961,1986]" pageId="8" pageNumber="397">FIGURE 5.</emphasis>
A: Hon Lao Island to which
<taxonomicName id="9C8C05906D409146FD8D64A6D82BD4A7" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[565,875,1962,1986]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="397" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D409146FD8D64A6D82BD4A7" box="[565,875,1962,1986]" italics="true" pageId="8" pageNumber="397">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D409146FCC964A6D881D4A7" box="[881,961,1961,1986]" pageId="8" pageNumber="397" rank="species">
<emphasis id="69F8A2016D409146FCC964A6D8FCD4A7" bold="true" box="[881,956,1962,1986]" pageId="8" pageNumber="397">sp. nov</emphasis>
.
</taxonomicNameLabel>
is endemic; B: Granite boulder, riparian microhabitat of
<taxonomicName id="9C8C05906D409146FF4664C2DAFDD483" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[254,445,1998,2022]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="397" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D409146FF4664C2DAFDD483" box="[254,445,1998,2022]" italics="true" pageId="8" pageNumber="397">C. culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D409146FE7C64C2D954D483" box="[452,532,1998,2022]" pageId="8" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="0FF32E9B6D419147FF2F6496D91FD4B2" ID-DOI="http://doi.org/10.5281/zenodo.3763541" ID-Zenodo-Dep="3763541" httpUri="https://zenodo.org/record/3763541/files/figure.png" pageId="9" pageNumber="398" startId="9.[151,250,1946,1971]" targetBox="[289,1295,187,1913]" targetPageId="9">
<paragraph id="5B337E136D419147FF2F6496D91FD4B2" blockId="9.[151,1436,1946,2007]" pageId="9" pageNumber="398">
<emphasis id="69F8A2016D419147FF2F6496DA52D4D6" bold="true" box="[151,274,1946,1971]" pageId="9" pageNumber="398">FIGURE 6.</emphasis>
Map showing the location of the type locality of
<taxonomicName id="9C8C05906D419147FCB86496DF76D4D7" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[768,1078,1946,1970]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="398" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D419147FCB86496DF76D4D7" box="[768,1078,1946,1970]" italics="true" pageId="9" pageNumber="398">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D419147FB836496DFCCD4D7" box="[1083,1164,1946,1970]" pageId="9" pageNumber="398" rank="species">
<emphasis id="69F8A2016D419147FB836496DFC6D4D7" bold="true" box="[1083,1158,1946,1970]" pageId="9" pageNumber="398">sp. nov</emphasis>
.
</taxonomicNameLabel>
in Cu lao Cham Biosphere Reserve Quang Nam, Vietnam (filled circle).
</paragraph>
</caption>
<paragraph id="5B337E136D429144FF7F6394DAC0D27A" blockId="10.[151,1437,152,1079]" pageId="10" pageNumber="399">
<emphasis id="69F8A2016D429144FF7F6394DAD9D3D6" bold="true" box="[199,409,152,179]" pageId="10" pageNumber="399">
Variation (
<figureCitation id="C3B762966D429144FEFD6394DACAD3D6" box="[325,394,152,179]" captionStart="FIGURE 2" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[232,1356,181,1919]" captionTargetId="figure@5.[232,1356,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cyrtodactylus culaochamensis sp. nov. A: Holotype ITBCZ 2494; B: paratype ITBCZ 2495 showing the irregularly shaped dorsal bands and blotches; C: paratype ITBCZ 2498 showing the thin nuchal loop and full tail with 13 light-colored bands alternating with dark-brown bands." figureDoi="http://doi.org/10.5281/zenodo.3763529" httpUri="https://zenodo.org/record/3763529/files/figure.png" pageId="10" pageNumber="399">Fig. 2</figureCitation>
).
</emphasis>
Color pattern of the
<typeStatus id="8437C0B16D429144FDC56395D9A8D3D6" box="[637,744,153,179]" pageId="10" pageNumber="399" type="paratype">paratypes</typeStatus>
resembles that of the
<typeStatus id="8437C0B16D429144FC606394DF7ED3D7" box="[984,1086,152,178]" pageId="10" pageNumber="399" type="holotype">holotype</typeStatus>
. ITBCZ 2496 bears a complete, thin, dark nuchal loop and the first dorsal body band is U-shaped. ITBCZ 2498 has a portion of regenerated tail is brown, scattered with small dark spots. ITBCZ 2494, showing light, subcaudal rings. Variation in scale counts is presented in
<tableCitation id="160E4BA86D429144FE9E6208DA3AD27A" box="[294,378,260,287]" captionStart="TABLE 1" captionStartId="1.[151,239,442,466]" captionTargetPageId="1" captionText="TABLE 1. Measurements and meristics of Cyrtodactylus culaochamensis sp. nov." httpUri="http://table.plazi.org/id/0FF32E9B6D49914FFF2F62B6DF47D2B6" pageId="10" pageNumber="399" tableUuid="0FF32E9B6D49914FFF2F62B6DF47D2B6">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="13962D986D429144FF7F6225DA3BD2EF" pageId="10" pageNumber="399" type="etymology">
<paragraph id="5B337E136D429144FF7F6225DA3BD2EF" blockId="10.[151,1437,152,1079]" pageId="10" pageNumber="399">
<emphasis id="69F8A2016D429144FF7F6225DA0ED226" bold="true" box="[199,334,297,323]" pageId="10" pageNumber="399">Etymology.</emphasis>
The specific epithet is derived from the name of Cu Lao Cham Mountain in Cu Lao Cham Biosphere Reserve. Suggested common names: English: Cù Lao Chàm Bent–toed Gecko; Vietnamese: Thằn lằn chân ngón Cù Lao Chàm.
</paragraph>
</subSubSection>
<subSubSection id="13962D986D429144FF7F6298DA1CD1E3" pageId="10" pageNumber="399" type="distribution">
<paragraph id="5B337E136D429144FF7F6298DA1CD1E3" blockId="10.[151,1437,152,1079]" pageId="10" pageNumber="399">
<emphasis id="69F8A2016D429144FF7F6298D918D2CA" bold="true" box="[199,600,404,431]" pageId="10" pageNumber="399">Distribution and Natural History.</emphasis>
<taxonomicName id="9C8C05906D429144FDE76298D8F6D2CB" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[607,950,404,430]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D429144FDE76298D8F6D2CB" box="[607,950,404,430]" italics="true" pageId="10" pageNumber="399">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<emphasis id="69F8A2016D429144FC056299DF56D2CA" bold="true" box="[957,1046,405,431]" pageId="10" pageNumber="399">
<taxonomicNameLabel id="72CB1F7A6D429144FC056299DF56D2CA" box="[957,1046,405,431]" pageId="10" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently known only from Hon Lao Island of Cu Lao Cham Biosphere Reserve, Hoi An Town in
<collectingRegion id="9948B0F16D429144FCD762B5D8B7D2B6" box="[879,1015,441,467]" country="Vietnam" name="Quang Nam" pageId="10" pageNumber="399">Quang Nam</collectingRegion>
City, Central
<collectingCountry id="239B3E836D429144FB2A62B4DFB5D2B7" box="[1170,1269,440,466]" name="Vietnam" pageId="10" pageNumber="399">Vietnam</collectingCountry>
. All specimens were collected at night on rock outcrops composed of large granitic boulders along a small, permanently flowing stream. Other specimens were observed on tree branches and dry tree roots beneath the canopy, 0.3 m–
<quantity id="9C74D3F66D429144FAAE610CDE13D17E" box="[1302,1363,512,539]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="399" unit="m" value="1.5">1.5 m</quantity>
above forest floor during the rainy season. ITBCZ
<date id="2F3258D36D429144FD386128D9BED15A" box="[640,766,548,575]" pageId="10" pageNumber="399" value="2259" valueMax="2260">2259–2260</date>
were collected by local people in Cu Lao Cham Islands during April of 2009. These specimens were put into 40% alcohol and became severely degraded and were not included in the
<typeStatus id="8437C0B16D429144FF656161DA4DD1E2" box="[221,269,621,647]" pageId="10" pageNumber="399">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="13962D986D429144FF7F619CDBB7D752" pageId="10" pageNumber="399" type="discussion">
<paragraph id="5B337E136D429144FF7F619CDBB7D752" blockId="10.[151,1437,152,1079]" pageId="10" pageNumber="399">
<emphasis id="69F8A2016D429144FF7F619CDA2DD1CE" bold="true" box="[199,365,656,683]" pageId="10" pageNumber="399">Comparisons.</emphasis>
<taxonomicName id="9C8C05906D429144FECF619CD992D1CF" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[375,722,656,682]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D429144FECF619CD992D1CF" box="[375,722,656,682]" italics="true" pageId="10" pageNumber="399">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<emphasis id="69F8A2016D429144FD64619DD879D1CE" bold="true" box="[732,825,657,683]" pageId="10" pageNumber="399">
<taxonomicNameLabel id="72CB1F7A6D429144FD64619DD879D1CE" box="[732,825,657,683]" pageId="10" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the other members in its clade (
<figureCitation id="C3B762966D429144FAF5619CDED1D1CE" box="[1357,1425,656,683]" captionStart="FIGURE 1" captionStartId="4.[151,255,1381,1405]" captionTargetBox="[193,1394,183,1342]" captionTargetId="figure@4.[193,1394,180,1354]" captionTargetPageId="4" captionText="FIGURE 1. Inferred phylogenetic relationships Cyrtodactylus culaochamensis sp. nov. The tree is a maximum likelihood topology with BPP and ML nodal support, respectively." figureDoi="http://doi.org/10.5281/zenodo.3763527" httpUri="https://zenodo.org/record/3763527/files/figure.png" pageId="10" pageNumber="399">Fig. 1</figureCitation>
) as outlined below. From
<taxonomicName id="9C8C05906D429144FE0461B8D856D1AB" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[444,790,692,718]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D429144FE0461B8D856D1AB" box="[444,790,692,718]" italics="true" pageId="10" pageNumber="399">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D429144FC9961B9D8C0D1AA" box="[801,896,693,719]" pageId="10" pageNumber="399" rank="species">
<emphasis id="69F8A2016D429144FC9961B9D839D1AA" bold="true" box="[801,889,693,719]" pageId="10" pageNumber="399">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from
<taxonomicName id="9C8C05906D429144FBA761B9DA43D197" authority="Hguyen, Orlov, &amp; Darevsky" authorityName="Hguyen, Orlov, &amp; Darevsky" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="badenensis">
<emphasis id="69F8A2016D429144FBA761B9DF80D1AB" box="[1055,1216,692,719]" italics="true" pageId="10" pageNumber="399">C. badenensis</emphasis>
Hguyen, Orlov, &amp; Darevsky
</taxonomicName>
,
<taxonomicName id="9C8C05906D429144FEB761D5D956D196" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1921" box="[271,534,728,755]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="condorensis">
<emphasis id="69F8A2016D429144FEB761D5DAF6D197" box="[271,438,728,755]" italics="true" pageId="10" pageNumber="399">C. condorensis</emphasis>
(Smith)
</taxonomicName>
,
<taxonomicName id="9C8C05906D429144FD9961D5D8DCD197" authority="Chan &amp; Norhayati" authorityName="Chan &amp; Norhayati" authorityYear="2010" box="[545,924,728,755]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="leegrismeri">
<emphasis id="69F8A2016D429144FD9961D5D982D197" box="[545,706,728,755]" italics="true" pageId="10" pageNumber="399">C. leegrismeri</emphasis>
Chan &amp; Norhayati
</taxonomicName>
,
<taxonomicName id="9C8C05906D429144FC1F61D5DFC6D196" authority="Ngo" authorityName="Ngo" box="[935,1158,729,755]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="eisenmanae">
<emphasis id="69F8A2016D429144FC1F61D5DF0CD197" box="[935,1100,729,755]" italics="true" pageId="10" pageNumber="399">C. eisenmanae</emphasis>
Ngo
</taxonomicName>
, and
<taxonomicName id="9C8C05906D429144FB7A61D5DE39D196" authority="Ngo" authorityName="Ngo" box="[1218,1401,729,755]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="grismeri">
<emphasis id="69F8A2016D429144FB7A61D5DE02D197" box="[1218,1346,729,755]" italics="true" pageId="10" pageNumber="399">C. grismeri</emphasis>
Ngo
</taxonomicName>
by the absence as opposed to the presence of enlarged femoral scales. It differs from
<taxonomicName id="9C8C05906D429144FBFC61F1DE7DD072" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1921" box="[1092,1341,764,791]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="69F8A2016D429144FBFC61F1DF9ED073" box="[1092,1246,764,791]" italics="true" pageId="10" pageNumber="399">C. irregularis</emphasis>
(Smith)
</taxonomicName>
by having 20–20 longitudinal rows of body tubercles as opposed to 17. It differs from
<taxonomicName id="9C8C05906D429144FB9D602DDE38D05F" authority="Ngo &amp; Chan" authorityName="Ngo &amp; Chan" authorityYear="2010" box="[1061,1400,800,827]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="yangbayensis">
<emphasis id="69F8A2016D429144FB9D602DDF9ED05F" box="[1061,1246,800,827]" italics="true" pageId="10" pageNumber="399">C. yangbayensis</emphasis>
Ngo &amp; Chan
</taxonomicName>
by having a smaller maximum SVL of
<quantity id="9C74D3F66D429144FD956049D9D2D03A" box="[557,658,837,863]" metricMagnitude="-2" metricUnit="m" metricValue="7.9799999999999995" pageId="10" pageNumber="399" unit="mm" value="79.8">79.8 mm</quantity>
versus
<quantity id="9C74D3F66D429144FD516049D81ED03A" box="[745,862,837,863]" metricMagnitude="-1" metricUnit="m" metricValue="1.091" pageId="10" pageNumber="399" unit="mm" value="109.1">109.1 mm</quantity>
. longitudinal rows of body tubercles as opposed to 17.
<taxonomicName id="9C8C05906D429144FF7A6064D95BD0E7" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[194,539,872,898]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D429144FF7A6064D95BD0E7" box="[194,539,872,898]" italics="true" pageId="10" pageNumber="399">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D429144FD9B6065D93FD0E6" box="[547,639,873,899]" pageId="10" pageNumber="399" rank="species">
<emphasis id="69F8A2016D429144FD9B6065D938D0E6" bold="true" box="[547,632,873,899]" pageId="10" pageNumber="399">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from
<taxonomicName id="9C8C05906D429144FCAF6065D8DCD0E6" authority="Ngo" authorityName="Ngo" box="[791,924,872,899]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="dati">
<emphasis id="69F8A2016D429144FCAF6065D824D0E7" box="[791,868,872,899]" italics="true" pageId="10" pageNumber="399">C. dati</emphasis>
Ngo
</taxonomicName>
by having seven or eight precloacal pores as opposed to five or six and no femoral prores as opposed to six or seven.
<taxonomicName id="9C8C05906D429144FC706080DE5FD0C3" authority="Tri &amp; Grismer &amp; Thai &amp; Jr, 2020" authorityName="Tri &amp; Grismer &amp; Thai &amp; Jr" authorityYear="2020" box="[968,1311,908,934]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="culaochamensis" status="sp. nov.">
<emphasis id="69F8A2016D429144FC706080DE5FD0C3" box="[968,1311,908,934]" italics="true" pageId="10" pageNumber="399">Cyrtodactylus culaochamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="72CB1F7A6D429144FA9F6081DEC2D0C2" box="[1319,1410,909,935]" pageId="10" pageNumber="399" rank="species">
<emphasis id="69F8A2016D429144FA9F6081DE3CD0C2" bold="true" box="[1319,1404,909,935]" pageId="10" pageNumber="399">sp. nov</emphasis>
.
</taxonomicNameLabel>
is most closely related to
<taxonomicName id="9C8C05906D429144FE2F60BDDF7DD0AF" authority="Rosler, Nguyen, Vu, Ngo &amp; Ziegler" authorityName="Rosler, Nguyen, Vu, Ngo &amp; Ziegler" box="[407,1085,944,971]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="399" phylum="Chordata" rank="species" species="pseudoquadrivirgatus">
<emphasis id="69F8A2016D429144FE2F60BDD9ECD0AF" box="[407,684,944,971]" italics="true" pageId="10" pageNumber="399">C. pseudoquadrivirgatus</emphasis>
Rösler, Nguyen, Vu, Ngo &amp; Ziegler
</taxonomicName>
from which it differs by having all the subcaudals being only slightly enlarged at best (
<figureCitation id="C3B762966D429144FD4960D8D807D08A" box="[753,839,980,1007]" captionStart="FIGURE 4" captionStartId="7.[151,250,1259,1284]" captionTargetBox="[177,1411,561,1235]" captionTargetId="figure@7.[177,1411,561,1235]" captionTargetPageId="7" captionText="FIGURE 4. A: Ventral view of tail of Cyrtodactylus culaochamensis sp. nov., ITBCZ 2494, showing light-colored, subcaudal rings and very slightly enlarged subcaudals; B: Precloacal region of C. culaochamensis sp. nov ITBCZ 2494, showing seven precloacal cloacal pores in a chevron pattern." figureDoi="http://doi.org/10.5281/zenodo.3763535" httpUri="https://zenodo.org/record/3763535/files/figure.png" pageId="10" pageNumber="399">Fig. 4A</figureCitation>
) versus having two medial rows of distinctly enlarged subcaudals (Rösler
<emphasis id="69F8A2016D429144FECB60F5DAEBD777" box="[371,427,1016,1042]" italics="true" pageId="10" pageNumber="399">et al.</emphasis>
2008;
<figureCitation id="C3B762966D429144FE4160F4D913D776" box="[505,595,1016,1043]" captionStart="FIGURE 2" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[232,1356,181,1919]" captionTargetId="figure@5.[232,1356,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. Cyrtodactylus culaochamensis sp. nov. A: Holotype ITBCZ 2494; B: paratype ITBCZ 2495 showing the irregularly shaped dorsal bands and blotches; C: paratype ITBCZ 2498 showing the thin nuchal loop and full tail with 13 light-colored bands alternating with dark-brown bands." figureDoi="http://doi.org/10.5281/zenodo.3763529" httpUri="https://zenodo.org/record/3763529/files/figure.png" pageId="10" pageNumber="399">Fig. 2D</figureCitation>
). These two species differ by an uncorrected pairwise sequence divergence of 7.2%.
</paragraph>
</subSubSection>
</treatment>
</document>
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