237 lines
27 KiB
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237 lines
27 KiB
XML
<document id="29E53A6C70697059186938503CCF6BEC" ID-DOI="http://dx.doi.org/10.3897/zookeys.532.6116" ID-GBIF-Dataset="1093c716-87cf-4bb2-bbb5-d9af884a5ad5" ID-PMC="PMC4668894" ID-Pensoft-Pub="1313-2970-532-87" ID-PubMed="26692807" ID-ZooBank="2FAFD349D1C74FEC828A7AD41D6430BD" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-532-87" ModsDocOrigin="ZooKeys 532" ModsDocTitle="A second species of Cheleion from Johor, Malaysia (Coleoptera, Scarabaeidae, Aphodiinae, Stereomerini)" checkinTime="1451243818864" checkinUser="pensoft" docAuthor="Kral, David & Hajek, Jiri" docDate="2015" docId="BE7C3032A6A719ABA82CD2AB2E425313" docLanguage="en" docName="ZooKeys 532: 87-97" docOrigin="ZooKeys 532" docSource="http://dx.doi.org/10.3897/zookeys.532.6116" docTitle="Cheleion jendeki Kral & Hajek, 2015, sp. n." docType="treatment" docUuid="4DF33FD9-FF5D-47A1-8967-06EAEE42349D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="94" masterDocId="FF84FF82037CFF91FF895B540321D929" masterDocTitle="A second species of Cheleion from Johor, Malaysia (Coleoptera, Scarabaeidae, Aphodiinae, Stereomerini)" masterLastPageNumber="97" masterPageNumber="87" pageNumber="88" updateTime="1701316758477" updateUser="plazi">
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<mods:mods id="D95A1E4D68793B6E0502218A0378A5DC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AA677F0DEC1AC5906EEDA262E57ED266">
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<mods:title id="5691F4651183311D548017FED3103927">A second species of Cheleion from Johor, Malaysia (Coleoptera, Scarabaeidae, Aphodiinae, Stereomerini)</mods:title>
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<mods:name id="CA80F268F45F7E82A001DA13C96D8898" type="personal">
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<mods:namePart id="D9F68D6CA57C5E5336A808C340B7057D">Kral, David</mods:namePart>
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<mods:name id="6749E429AE0656155475DBA9980AAE81" type="personal">
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<mods:roleTerm id="BBD525BD20796B614A69B8F0FEC5B767">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3C797CE7E0C4213A6974744D9EEF9245">Hajek, Jiri</mods:namePart>
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<mods:typeOfResource id="852B3096D061D64AAE1CF0F07F7C9E2E">text</mods:typeOfResource>
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<mods:title id="755B8D0A51426D37B0A2A215E831BF83">ZooKeys</mods:title>
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<mods:part id="5A4DEAB20B33D2F768ECBFC6F29EE17A">
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<mods:date id="A0C22C9A1414768621657AEE75792873">2015</mods:date>
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<mods:detail id="70B682BD7B8983815C59BDB0AC19E80F" type="volume">
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<mods:number id="AA8542C6BA89AEB30A574EA7A0FDE631">532</mods:number>
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<mods:start id="29427060437CF32A1F09BF8C70EF0125">87</mods:start>
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<mods:end id="F97433166D94C5655C97FAF484A49B03">97</mods:end>
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<mods:url id="AE39DA57802DA02F20048719BF41A6CD">http://dx.doi.org/10.3897/zookeys.532.6116</mods:url>
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<mods:classification id="C853AA748297F69C37775F6256ED3856">journal article</mods:classification>
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<mods:identifier id="DEB8FBC68B9A00D5C09C092C59ED84FB" type="DOI">http://dx.doi.org/10.3897/zookeys.532.6116</mods:identifier>
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<mods:identifier id="BC4C06B4A99FEF9C326535DEA9977CE6" type="Pensoft-Pub">1313-2970-532-87</mods:identifier>
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</mods:mods>
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<treatment id="BE7C3032A6A719ABA82CD2AB2E425313" ID-GBIF-Taxon="152062982" LSID="urn:lsid:zoobank.org:act:4DF33FD9-FF5D-47A1-8967-06EAEE42349D" httpUri="http://treatment.plazi.org/id/BE7C3032A6A719ABA82CD2AB2E425313" lastPageId="7" lastPageNumber="94" pageId="1" pageNumber="88">
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<subSubSection id="EE958AF195E6F4C6F9F06EC8D7023AA1" pageId="1" pageNumber="88" type="multiple">
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<paragraph id="28E3ED3135D0188E0E29ABA46FAED0C7" pageId="1" pageNumber="88">Taxon classification Animalia Coleoptera Scarabaeidae</paragraph>
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</subSubSection>
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<subSubSection id="F6B481A6562DE77BE77BDC2270AFB8DF" pageId="1" pageNumber="88" type="nomenclature">
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<paragraph id="2197E318D1CFDBDBC26EB58708EFC8C9" pageId="1" pageNumber="88">
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<taxonomicName id="44F1F099FBA0F112804580827DC7918B" ID-CoL="5Y36J" LSID="http://zoobank.org/4DF33FD9-FF5D-47A1-8967-06EAEE42349D" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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<taxonomicNameLabel id="30FE60D6720DDBA6E7FBDFD63CD9100C" pageId="1" pageNumber="88">sp. n.</taxonomicNameLabel>
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Figures 1, 3, 5, 7, 9, 11, 13-15, 17
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</paragraph>
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</subSubSection>
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<subSubSection id="D0E349F56E9C1ED6BF4786A914BEC042" pageId="1" pageNumber="88" type="type locality">
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<paragraph id="698A81436995C844143D6AA46F56C498" pageId="1" pageNumber="88">Type locality.</paragraph>
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<paragraph id="A0A244FC1C4A3DE2E175D6E5A63EF4D0" pageId="1" pageNumber="88">
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Malaysia, Johor, Endau Rompin National Park,
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<geoCoordinate id="541CB1F28B1713EA0D4F2AD971315D66" direction="north" orientation="latitude" precision="15" value="2.6200001">02°37'12"N</geoCoordinate>
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<geoCoordinate id="8540A3CE947A7969ED77C58F378B4988" direction="east" orientation="longitude" precision="15" value="103.35">103°21'00"E</geoCoordinate>
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, 120-300 m a. s. l.
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</paragraph>
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</subSubSection>
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<subSubSection id="2C29172EC6414E8816E1FB4397C5504B" pageId="1" pageNumber="88" type="type material">
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<paragraph id="90A6D42349367A23DFB54E7C2E02C7CD" pageId="1" pageNumber="88">Type material.</paragraph>
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<paragraph id="5646F7AD0E8D4A064693F27D38428CF4" pageId="1" pageNumber="88">
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Holotype: ♀, "Malaysia, Johor / Endau Rompin NP /
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<geoCoordinate id="657F8B36E9B8973CFA878A6132F4263D" direction="north" orientation="latitude" precision="555" value="2.62">N2.62</geoCoordinate>
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,
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<geoCoordinate id="1AC2EA4F4422354ACCC9DC3976C7CB66" direction="east" orientation="longitude" precision="555" value="103.35">E103.35</geoCoordinate>
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/ 28-31.v.2013, 120-300 m / E. Jendek & O.
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<normalizedToken id="20D3F1AA97F2D8CD52BE83A23465ADD5" originalValue="Šauša">Sausa</normalizedToken>
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leg. [printed] //
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<taxonomicName id="796A07C4C20F944D6D8D21951B005F44" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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sp. nov. / HOLOTYPUS ♀ / David
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<normalizedToken id="5B77FA25251ABF6A1933DA9C21629982" originalValue="Král">Kral</normalizedToken>
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&
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<normalizedToken id="B579C74C6694289F8D75E106347E3D7E" originalValue="Jiří">Jiri</normalizedToken>
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<normalizedToken id="7098B820C26B0D0E40D3520CE29CEDAE" originalValue="Hájek">Hajek</normalizedToken>
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det. 2015 [red, printed]".
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</paragraph>
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</subSubSection>
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<subSubSection id="70DB39CA4DFB69893C9C821CFA57CC1D" lastPageId="6" lastPageNumber="93" pageId="2" pageNumber="89" type="description of female holotype">
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<paragraph id="BD7415D2B6E4D73771934968E016D61F" pageId="2" pageNumber="89">
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<pageBreakToken id="794BBE4B75B92BE2F9D16C1AD76C79E3" pageId="2" pageNumber="89" start="start">Description</pageBreakToken>
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of female holotype.
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</paragraph>
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<paragraph id="23028D59615F29EDC9704AF9653452AE" pageId="2" pageNumber="89">Slightly convex, integument chestnut brown; head appendages and tarsi amber coloured; whole dorsal surface more or less covered with appressed lanceolate scales (Fig. 1).</paragraph>
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<caption id="A5465344CE3512F00DC7C210BB460CEC" pageId="2" pageNumber="89">
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<paragraph id="787CE14C7C37A0841A4C5246AB71E062" pageId="2" pageNumber="89">
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Figures 1-2. Habitus of
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<taxonomicName id="C81E21449012031D9BB11E458FF03EFC" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Cheleion</taxonomicName>
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. 1
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<taxonomicName id="EC3B68D3FEAF273D7413F9F9D99AB8AE" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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sp. n. (♀ holotype) 2
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<taxonomicName id="CC12C82AFE113B40CDE2CD04EB67617E" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(♀ holotype; Malaysia, Pahang, Bukit Fraser).
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</paragraph>
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</caption>
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<paragraph id="BF5E11A74D7F401A05FA3D165E174E4A" pageId="2" pageNumber="89">Head (Figs 1, 3, 7) remarkably transverse, subrectangular in dorsal view, clypeus shiny, impunctate, apically pointed and reflexed under head, frons slightly convex with five straight, anteriorly divergent furrows; posterior transverse furrow across head between posterolateral corners of eyes; occiput with numerous small, longitudinal pits. Surface covered with dense appressed, lanceolate, approximately regularly spaced scales, individual scales separated from each other by less than their diameter (Fig. 7). Antennae long, length equal to width of head, with long macrosetae. Maxillary palpi length equal to length of head, with securiform ultimate palpomere. Labial palpi with long macrosetae apically. Eyes small but visible in dorsal view (Fig. 7).</paragraph>
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<caption id="7BE9A28086F9FD1E37FA43EB70DF9774" pageId="2" pageNumber="89">
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<paragraph id="D248FC03F712A7A1815E8210F12B538E" pageId="2" pageNumber="89">
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Figures 3-6. Habitus of
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<taxonomicName id="42E627BE3733001D7C459F0C86F6D5E4" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Cheleion</taxonomicName>
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. 3, 5
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<taxonomicName id="7352E5110FD6068C815E88BC581D12AE" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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sp. n. (♀ holotype) 4, 6
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<taxonomicName id="02914991A95B1B137C81A7D328466CB4" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(♀ holotype; Malaysia, Pahang, Bukit Fraser) 3, 4 dorsal view 5, 6 ventral view.
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</paragraph>
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</caption>
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<caption id="4B7F9532FFA30C54F52EA65113A71097" pageId="2" pageNumber="89">
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<paragraph id="C102B5ACA82FAA80D313C346645E166A" pageId="2" pageNumber="89">
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Figures 7-12. Details of
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<taxonomicName id="F712F49451980D2E6852C0548AD07EE0" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">Cheleion</taxonomicName>
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. 7, 9, 11
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<taxonomicName id="C0EA2135AC3FB9855D325776B1896386" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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sp. n. (♀ holotype) 8, 10, 12
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<taxonomicName id="87B37F6014EE95CF19E3D2EA58417B52" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(♀ holotype; Malaysia, Pahang, Bukit Fraser) 7-8 head 9-10 pronotum 11-12 base of right elytron.
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</paragraph>
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</caption>
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<paragraph id="1A1642CCB5F3B15A861B50135D2C6608" lastPageId="4" lastPageNumber="91" pageId="2" pageNumber="89">
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Pronotum (Figs 3, 9, 13) large and transverse, anterior edge shallowly bisinuate, sides regularly, broadly rounded, posterior edge with broad medial protrusion. Pronotal disc with seven furrows medially, converging towards middle in hourglass
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<pageBreakToken id="26CB303F68A97745FB34B4C36FBF3636" pageId="3" pageNumber="90" start="start">pattern</pageBreakToken>
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, mid furrow shallower than lateral furrows (Figs 3, 9). Anteromedial disc with distinctly raised knob, posteromedial disc and posterolateral sides with slightly lower, bulbous areas; anterolaterally of the furrows with large, flat elliptical depressions, de
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<pageBreakToken id="5662F280530F62BB8668641F476216DC" pageId="4" pageNumber="91" start="start">lineated</pageBreakToken>
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by furrows. Knob posteriorly and bulbous areas anteriorly with tufts of long dense microtrichiae (= trichomes) (Fig. 13); surface covered with dense apressed, lanceolate, approximately regularly spaced scales, individual scales separated by less their diameter from each other anterolaterally and laterally; scales on knob and bulbous areas smaller and sparser; flat lateral areas with several sparse rather irregularly spaced scales only (Fig. 13).
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</paragraph>
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<caption id="057085409B8024C98995914CD06325EC" pageId="4" pageNumber="91">
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<paragraph id="577FD2165E7E0EC5F32D93906E01188D" pageId="4" pageNumber="91">
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Figures 13-18. Details of
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<taxonomicName id="A9556345DB2D36D37339BB601B4A1C92" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="genus">Cheleion</taxonomicName>
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. 15, 17
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<taxonomicName id="534D07A590F0B56DD6357F30514F2207" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(♀ holotype; Malaysia, Pahang, Bukit Fraser) 13-14, 16, 18
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<taxonomicName id="E4C70BD78E0145D4B8224962E6C523D7" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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sp. n. (♀ holotype) 13 detail of tufts of microtrichiae (= trichomes) in centre of pronotum 14 detail of macrosetation on elytral shoulder 15-16 head and prosternum in ventral view 17-18 meso- and metaventrite in ventral view.
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</paragraph>
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</caption>
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<paragraph id="2A3062C07AA8D0FD8E9BC3734DAC64DC" pageId="4" pageNumber="91">Scutellar shield triangular, notably small (Fig. 11).</paragraph>
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<paragraph id="A2FD0DB72E6E876BFBA52E3DC7CD2864" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92">
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<pageBreakToken id="7D9D0992BF59A68C7FDFEC80E91DF13F" pageId="5" pageNumber="92" start="start">Elytra</pageBreakToken>
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approximately as broad as pronotum and only slightly longer than pronotum and head combined; tapering posteriad, rounded apically. Each elytron with five longitudinal ridges before the lateral edge (Figs 1, 3, 11); ridges of approximately same height, elevated and almost continuous, consisting of longitudinal rows of almost confluent tubercles (Fig. 11); intervals (between ridges) flat, rugose, with irregularly circular pads, each pad bearing lanceolate scale on posterior edge, individual pads sepa
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<pageBreakToken id="AF4065A2B76D2F3DDE735B811A9059AC" pageId="6" pageNumber="93" start="start">rated</pageBreakToken>
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by less their diameter from each other discally, becoming confluent into small rows or groups laterally, especially in humeral area (Figs 11, 14). Epipleura broadly inflexed; posterior two thirds of lateral edge slightly recurved (to allow free movement of metathoracic legs).
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</paragraph>
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<paragraph id="0A9283E8D7895F85A4149A2D5854F2E9" pageId="6" pageNumber="93">Legs short with broad femora and tibiae; tarsi short, tetramerous; claws weak, short, almost straight (Figs 5, 15, 17). Femora shiny, covered with coarse, dense, almost regularly spaced macrosetigerous punctures (Figs 15, 17). Protibiae moderately wide, with finely serrated outer edge and one strong apical lateral tooth, tarsus inserted well before apex (Fig. 15). Meso- and metatibiae broad with finely serrated outer edge and concave apex; each with two inconspicuous terminal spurs and two rows of thick short macrosetae on outer edge (Fig. 17).</paragraph>
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<paragraph id="9B22D3A11B481720C5A51A733290E327" pageId="6" pageNumber="93">Macropterous.</paragraph>
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<paragraph id="F48A88A7B7D5B222751375F1B3932811" pageId="6" pageNumber="93">Pygidium exposed, strongly punctate proximally, less strongly apically, with small emargination on proximal pygidial border.</paragraph>
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<paragraph id="CA3A2CA4BAB5069F216402EE7073CAE7" pageId="6" pageNumber="93">Venter. Prosternal process remarkably elevated, strongly expanded anteriad and posteriad (Figs 5, 15); anterior part grooved longitudinally and sinuate apically, posterior part hastate, surface rugose with marginal bead (Fig. 15). Mesoventrite narrow with alutaceous surface (Fig. 17). Metaventral plate flat, triangular, tapering, widest anteriorly, grooved along midline, surface alutaceous, covered with coarse, dense, almost regularly spaced macrosetigerous punctures (Fig. 17).</paragraph>
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<paragraph id="6508271D2D85727CD6F9E6AF492C9330" pageId="6" pageNumber="93">Five visible abdominal ventrites apparently fused, covered with coarse, dense, almost regularly spaced macrosetigerous punctures.</paragraph>
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</subSubSection>
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<subSubSection id="CB6B994FE3AA26407502A21F9D2204FC" pageId="6" pageNumber="93" type="measurements">
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<paragraph id="B1662ADD56F81A076D2E55C7F92AB7BE" pageId="6" pageNumber="93">Measurements.</paragraph>
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<paragraph id="8532A4563849597F02833A2183F552B5" pageId="6" pageNumber="93">Total body length: 1.9 mm, width at broadest point 0.9 mm.</paragraph>
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</subSubSection>
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<subSubSection id="2D977492990D94033C750937ADFCBBED" lastPageId="7" lastPageNumber="94" pageId="6" pageNumber="93" type="differential diagnosis">
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<paragraph id="4EF07DD7D2C91DC3032A569AE9DB46D3" pageId="6" pageNumber="93">Differential diagnosis.</paragraph>
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<paragraph id="2BF7C7E1918FCF0119F212B0721DEF5E" pageId="6" pageNumber="93">
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The new species is classified in the genus
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<taxonomicName id="7731F3B3A6636E62ECFDB64F6186EEC9" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">Cheleion</taxonomicName>
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mainly by the combination of the following characters: strongly tuberculate and rugose body surface, noticeably long antennae, pronotum with distinct anteromedial knob and bulbous areas medially and laterally and hastate posterior prosternal process.
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<taxonomicName id="B6F59EE89D372D8F455909C17C94A0F2" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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sp. n. is similar and probably closely related to
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<taxonomicName id="3B799B2ACFFB040C18A3BD0056EA4898" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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, the only other known species of the genus, but clearly differs mainly as follows:
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</paragraph>
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<paragraph id="DB7A45B0123F8852B226B84B768A909A" pageId="6" pageNumber="93">
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lateral longitudinal grooves on head straight (Figs 1, 3, 7) (weakly s-shaped in
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<taxonomicName id="6707129C197CFD5E322A5430D299179C" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(Figs 2, 4, 8));
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</paragraph>
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<paragraph id="E6B65862C95F382F2E5BB826B0904D88" pageId="6" pageNumber="93">
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sides of pronotum regularly rounded, maximum width of pronotum in midlength; posterior angles subrectangular (Figs 3, 9, 15) (sides of pronotum more attenuated in basal third, maximum width of pronotum in anterior third; posterior angles obtuse with apparent spiniform scales in
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<taxonomicName id="1FA0476E6F3C1960E671F3C5FBE453BF" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(Figs 4, 10, 16);
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</paragraph>
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<paragraph id="8D581962BE449E31F97AF796644B7B9D" pageId="6" pageNumber="93">
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|
elytral surface at first sight moderately rugose (Figs 1, 3, 11) (more strongly rugose in
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<taxonomicName id="4EFA333D2264B4432AA1B4F0DC141D59" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(Figs 2, 4, 12);
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</paragraph>
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<paragraph id="68CB9547C38FFA20515D4EF3875B9FD7" pageId="6" pageNumber="93">
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|
elytral ridges distinctly elevated (Figs 3, 11) (almost flat in
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|
<taxonomicName id="9EE581B3E66A6B9567A2A2D624E42C87" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(Figs 4, 12);
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</paragraph>
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|
<paragraph id="8B81C52466213E93CE8DA0F91DC50668" pageId="6" pageNumber="93">
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|
elytral ridges continuous, consisting of longitudinal rows of almost confluent tubercles; lateral ridges indistinct (Fig. 3) (rather discontinuous, consisting of tubercles with scanty longitudinal groups of tubercles with scales; all elytral ridges distinct in
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<taxonomicName id="8286DBC02E4CDD836B5B2DCCF5C6386A" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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(Fig. 4));
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|
</paragraph>
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|
<paragraph id="FA32C9C2BA52C02FC5823E046F3219F8" pageId="7" pageNumber="94">
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|
<pageBreakToken id="7F9872E0C9D4A81599F3DE98E11B9E6B" pageId="7" pageNumber="94" start="start">pads</pageBreakToken>
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|
on elytral intervals separated by less their diameter and mostly arranged as triseriate discally (Fig. 11) (confluent to subconfluent and mostly arranged as biseriate in
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|
<taxonomicName id="3449E9BA78A2AF7FCF0DEE5F82B7DFF1" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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|
(Fig. 12));
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|
</paragraph>
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|
<paragraph id="B7B1CC57C2DAA8715F0D49B331784975" pageId="7" pageNumber="94">
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|
marginal bead of posterior part of prosternal process rounded apically (Fig. 15) (angulate apically in
|
|
<taxonomicName id="9C6D93EB3925258DD3BA06EE6A43CE93" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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|
(Fig. 16)).
|
|
</paragraph>
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|
<paragraph id="B17CCC727A7F7A1CDF0F42856EE9D93A" pageId="7" pageNumber="94">
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|
In spite of clear differences mentioned above, we are aware that only single specimens are known for each
|
|
<taxonomicName id="0F8796F61D586EEA7D43EDDE50D20E2C" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">Cheleion</taxonomicName>
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|
species. In addition, both type localities are placed only about 200 km apart, without any distinct barrier between them. Thus, we cannot exclude the possibility that morphological differences of
|
|
<taxonomicName id="61D04C2972A2215BCEF58B8EE6E57E51" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
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|
sp. n. represent only an intraspecific variability of
|
|
<taxonomicName id="CF70693B6AC934A0718BC7FF240757A3" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion malayanum" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="malayanum">Cheleion malayanum</taxonomicName>
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|
, but we consider it quite improbable.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="50ABFBA0D76FD9E8A8D0F1C3152FE0C5" pageId="7" pageNumber="94" type="etymology">
|
|
<paragraph id="49C3E5B3C079712D8798A74A5AB1EA1F" pageId="7" pageNumber="94">Etymology.</paragraph>
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|
<paragraph id="A9A65A072128010D083204B4FB8BF93F" pageId="7" pageNumber="94">
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|
Patronymic; named in honour of our colleague and friend Eduard Jendek (Ottawa, Canada), excellent student in
|
|
<taxonomicName id="DA511D6A681981CB89E9A5C48E04F1AB" family="Buprestidae" lsidName="" pageId="7" pageNumber="94" rank="family">Buprestidae</taxonomicName>
|
|
and collector of the holotype.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6242A232A5F8DB1F455B275319C6AA01" pageId="7" pageNumber="94" type="distribution">
|
|
<paragraph id="F07DE5D1F2F0976BC179D031910175B6" pageId="7" pageNumber="94">Distribution.</paragraph>
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|
<paragraph id="87229993B725F0DB0C2B0426D4DACE77" pageId="7" pageNumber="94">So far known only from the type locality in the Johor Province of continental Malaysia.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BA5E4ABE92B89B688C9ACDE2B44DDB0C" pageId="7" pageNumber="94" type="collecting circumstances">
|
|
<paragraph id="44F91DE7D3329155440E54F912CC2E2B" pageId="7" pageNumber="94">Collecting circumstances.</paragraph>
|
|
<paragraph id="6D287529AF4975DF3AFA27DC35EBFF96" pageId="7" pageNumber="94">Flight intercept trap exposed inside lowland primary tropical forest (Fig. 19; E. Jendek, pers. comm. 2015).</paragraph>
|
|
<caption id="871ACD2E479F48018E40432F027223C4" pageId="7" pageNumber="94">
|
|
<paragraph id="6A2E2E099171925191A8D36FEDEB8F2F" pageId="7" pageNumber="94">
|
|
Figure 19. Lowland primary tropical forest in Endau Rompin NP, habitat of
|
|
<taxonomicName id="EDEACCEB904084EBBC88EDECA99F7224" class="Insecta" family="Scarabaeidae" genus="Cheleion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cheleion jendeki" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="jendeki">Cheleion jendeki</taxonomicName>
|
|
sp. n. (photo E. Jendek).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
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</document> |