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<document id="0C34D6656B4729B28BAB717748A3F285" ID-DOI="10.5281/zenodo.5325118" ID-ISSN="0374-1036" ID-Zenodo-Dep="5325118" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630142892026" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2010" docId="BE6487EEB035FFCFFE006F85A1BD163C" docLanguage="en" docName="ActaEntMusNatPra.2010.50.1.131-144.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 50 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus chappuisi Bernhauer 1939" docType="treatment" docVersion="4" lastPageNumber="134" masterDocId="425DFF96B036FFCCFF836F5FA373106D" masterDocTitle="A revision of the Afrotropical species of the Philonthus arrowianus species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="144" masterPageNumber="131" pageNumber="134" updateTime="1699163914098" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0C8B97079A0547D181C3BACB4D822CAE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E13D749CEB1DE3C9E1E7E05DC0EE7945">A revision of the Afrotropical species of the Philonthus arrowianus species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
</mods:titleInfo>
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<mods:namePart id="E1096EC2FC1CF1E9A271AAC160646D75">Hromádka, Lubomír</mods:namePart>
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<mods:typeOfResource id="93F128D0F8D5E8ECE17DF9A3A1388AC4">text</mods:typeOfResource>
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<mods:title id="B583D2D4BF237A44394DE5501BE3B600">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="E3D4415A211304070B61CB25C99DBDA2">
<mods:date id="A74ABA4ABBB2660A2233A0770DADCBFF">2010</mods:date>
<mods:detail id="581DFF5EF94B887EBE46EF906FC646A1" type="pubDate">
<mods:number id="743E42E921B6B6D176E5CE667656FBF8">2010-06-30</mods:number>
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<mods:number id="43EAD4B562AD25F545C448F7697F39C9">50</mods:number>
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<mods:classification id="28E7E7DED84FDF401B7529CC581734A2">journal article</mods:classification>
<mods:identifier id="784F8A3F7EC6F2BEBCF16A8FCD4A15A1" type="DOI">10.5281/zenodo.5325118</mods:identifier>
<mods:identifier id="ECF3ED2E98D7CF22D0BDA9860C17A9B9" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="593571DDCC77B473A5E9906E13310B6A" type="Zenodo-Dep">5325118</mods:identifier>
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<treatment id="BE6487EEB035FFCFFE006F85A1BD163C" ID-DOI="http://doi.org/10.5281/zenodo.5344213" ID-Zenodo-Dep="5344213" LSID="urn:lsid:plazi:treatment:BE6487EEB035FFCFFE006F85A1BD163C" httpUri="http://treatment.plazi.org/id/BE6487EEB035FFCFFE006F85A1BD163C" lastPageNumber="134" pageId="3" pageNumber="134">
<subSubSection id="7ED76573B035FFCFFE006F85A05C109E" box="[387,815,218,243]" pageId="3" pageNumber="134" type="nomenclature">
<paragraph id="367236F8B035FFCFFE006F85A05C109E" blockId="3.[387,815,218,243]" box="[387,815,218,243]" pageId="3" pageNumber="134">
<heading id="6D3A8194B035FFCFFE006F85A05C109E" bold="true" box="[387,815,218,243]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="134" reason="3">
<taxonomicName id="F1CD4D7BB035FFCFFE006F85A05C109E" ID-CoL="6V9J2" authority="Bernhauer, 1939" authorityName="Bernhauer" authorityYear="1939" box="[387,815,218,243]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="chappuisi">
<emphasis id="04B9EAEAB035FFCFFE006F85A05C109E" bold="true" box="[387,815,218,243]" pageId="3" pageNumber="134">
<emphasis id="04B9EAEAB035FFCFFE006F85A11D109E" bold="true" box="[387,622,218,243]" italics="true" pageId="3" pageNumber="134">Philonthus chappuisi</emphasis>
Bernhauer, 1939
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7ED76573B035FFCFFDAE6FA6A1F61161" box="[557,645,249,268]" pageId="3" pageNumber="134" type="description">
<paragraph id="367236F8B035FFCFFDAE6FA6A1F61161" blockId="3.[113,1089,249,417]" box="[557,645,249,268]" pageId="3" pageNumber="134">
(
<figureCitation id="AEF62A7DB035FFCFFDB06FA6A10C1161" box="[563,639,249,268]" captionStart="Figs" captionStartId="4.[151,190,1437,1456]" captionTargetBox="[227,1054,205,1388]" captionTargetId="figure-10@4.[217,1062,205,1409]" captionTargetPageId="4" captionText="Figs. 1–14. Philonthus species. 1–5 – P. arrowianus Bernhauer, 1931 (1 – aedeagus, ventral view; 2 – aedeagus, lateral view; 3 – apex of of paramere with sensory peg setae, ventral view; 4 – apical portion of male sternite VIII, ventral view; 5 – male sternite IX, ventral view). 6–9 – P. chappuisi Bernhauer, 1939 (6 – aedeagus, ventral view; 7 – aedeagus, lateral view; 8 – apex of paramere with sensory peg setae, ventral view; 9 – male sternite IX, ventral view). 10–14 – P. currax sp. nov. (10 – aedeagus, ventral view; 11 – aedeagus without paramere, ventral view; 12 – aedeagus, lateral view; 13 – apex of paramere with sensory peg setae; 14 – apical portion of male sternite VIII, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5325120" httpUri="https://zenodo.org/record/5325120/files/figure.png" pageId="3" pageNumber="134">Figs 6–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7ED76573B035FFCFFFF26E42A2931142" box="[113,480,284,303]" pageId="3" pageNumber="134" type="reference_group">
<paragraph id="367236F8B035FFCFFFF26E42A2931142" blockId="3.[113,1089,249,417]" box="[113,480,284,303]" pageId="3" pageNumber="134">
<taxonomicName id="F1CD4D7BB035FFCFFFF26E42A2AF1142" ID-CoL="6V9J2" authority="Bernhauer, 1939: 84" authorityName="Bernhauer" authorityPageNumber="84" authorityYear="1939" box="[113,476,284,303]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="chappuisi">
<emphasis id="04B9EAEAB035FFCFFFF26E42A2551142" box="[113,294,285,303]" italics="true" pageId="3" pageNumber="134">Philonthus chappuisi</emphasis>
Bernhauer, 1939: 84
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7ED76573B035FFCFFFF26E61A2C211CC" pageId="3" pageNumber="134" type="materials_examined">
<paragraph id="367236F8B035FFCFFFF26E61A108113C" blockId="3.[113,1089,249,417]" box="[113,635,317,337]" pageId="3" pageNumber="134">
<emphasis id="04B9EAEAB035FFCFFFF26E61A399113C" bold="true" box="[113,234,317,337]" pageId="3" pageNumber="134">
<typeStatus id="E976885AB035FFCFFFF26E61A3ED113C" box="[113,158,318,337]" pageId="3" pageNumber="134">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="4EDA7668B035FFCFFF6C6E61A25A113C" box="[239,297,318,337]" name="Kenya" pageId="3" pageNumber="134">Kenya</collectingCountry>
, Mt. Aberdare, Kinangop,
<quantity id="F1359B1DB035FFCFFD8E6E61A13D113C" box="[525,590,318,337]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="134" unit="m" value="2600.0">2600 m</quantity>
a.s.l.
</paragraph>
<paragraph id="367236F8B035FFCFFFF26E06A2C211CC" blockId="3.[113,1089,249,417]" pageId="3" pageNumber="134">
<emphasis id="04B9EAEAB035FFCFFFF26E06A23D1101" bold="true" box="[113,334,344,364]" pageId="3" pageNumber="134">Type material examined.</emphasis>
<materialsCitation id="86A53CA5B035FFCFFED16E06A2C211CC" collectionCode="FMNH" collectorName="Mission de l'Omo, C &amp; Arambourg, P. A &amp; R. Jeannel" country="Kenya" county="Mission de l'Omo" elevation="2600" location="Mais forest" municipality="Ouest" pageId="3" pageNumber="134" specimenCount="1" typeStatus="holotype">
<typeStatus id="E976885AB035FFCFFED16E06A2D71107" box="[338,420,345,364]" pageId="3" pageNumber="134" type="holotype">HOLOTYPE</typeStatus>
: J, ‛
<collectingCountry id="4EDA7668B035FFCFFE4D6E06A16B1101" box="[462,536,345,364]" name="Kenya" pageId="3" pageNumber="134">KENYA</collectingCountry>
,
<location id="33126023B035FFCFFD9C6E06A10E1101" LSID="urn:lsid:plazi:treatment:BE6487EEB035FFCFFE006F85A1BD163C:33126023B035FFCFFD9C6E06A10E1101" box="[543,637,345,364]" country="Kenya" county="Mission de l'Omo" municipality="Ouest" name="Mais forest" pageId="3" pageNumber="134">Mais forest</location>
<location id="33126023B035FFCFFD026E06A1A41101" LSID="urn:lsid:plazi:treatment:BE6487EEB035FFCFFE006F85A1BD163C:33126023B035FFCFFD026E06A1A41101" box="[641,727,345,364]" country="Kenya" county="Mission de l'Omo" municipality="Ouest" name="Kinangop" pageId="3" pageNumber="134">Kinangop</location>
,
<location id="33126023B035FFCFFD5E6E06A03C1101" LSID="urn:lsid:plazi:treatment:BE6487EEB035FFCFFE006F85A1BD163C:33126023B035FFCFFD5E6E06A03C1101" box="[733,847,345,364]" country="Kenya" county="Mission de l'Omo" municipality="Ouest" name="Mt. Aberdare" pageId="3" pageNumber="134">Mt.Aberdare</location>
,ver´s
<collectingMunicipality id="D616AC82B035FFCFFC006E06A0C71101" box="[899,948,345,364]" pageId="3" pageNumber="134">Ouest</collectingMunicipality>
<quantity id="F1359B1DB035FFCFFC3A6E06A0861101" box="[953,1013,345,364]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="134" unit="m" value="2600.0">
<elevation id="BDE0D1CBB035FFCFFC3A6E06A0861101" box="[953,1013,345,364]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="134" unit="m" value="2600.0">2600m</elevation>
</quantity>
,
<collectorName id="9B38532EB035FFCFFC7F6E06A3AE11EB" pageId="3" pageNumber="134">
<collectingCounty id="DF134E74B035FFCFFC7F6E06A3B711EB" pageId="3" pageNumber="134">Mission de l´Omo</collectingCounty>
, C
</collectorName>
.
<collectorName id="9B38532EB035FFCFFF606E2CA20011EB" box="[227,371,371,391]" pageId="3" pageNumber="134">Arambourg, P.A</collectorName>
. Chappuis &amp;
<collectorName id="9B38532EB035FFCFFE666E2BA14C11EB" box="[485,575,371,391]" pageId="3" pageNumber="134">R. Jeannel</collectorName>
, 1932 – 33. //
<taxonomicName id="F1CD4D7BB035FFCFFD356E2BA0BB11EB" authority="Bernhauer" authorityName="Bernhauer" authorityYear="1939" box="[694,968,371,390]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="chappuisi">
<emphasis id="04B9EAEAB035FFCFFD356E2BA01911EB" box="[694,874,372,390]" italics="true" pageId="3" pageNumber="134">Philonthus chappuisi</emphasis>
Bernhauer
</taxonomicName>
, TYPE [ochre oblong label, handwritten]’ (
<collectionCode id="50DCAE3DB035FFCFFEE76ED1A2D411CC" box="[356,423,398,417]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="134" type="Museum">FMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7ED76573B035FFCFFFF26EE6A22F159C" pageId="3" pageNumber="134" type="description">
<paragraph id="367236F8B035FFCFFFF26EE6A08911BC" blockId="3.[113,1090,441,1618]" box="[113,1018,441,465]" pageId="3" pageNumber="134">
<emphasis id="04B9EAEAB035FFCFFFF26EE6A26411BC" bold="true" box="[113,279,441,465]" pageId="3" pageNumber="134">Redescription.</emphasis>
Body length
<quantity id="F1359B1DB035FFCFFE246EE6A17B11BC" box="[423,520,441,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="3" pageNumber="134" unit="mm" value="10.9">10.9 mm</quantity>
, length of fore body (to end of elytra) 5.0 mm.
</paragraph>
<paragraph id="367236F8B035FFCFFF136E86A397121C" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Colouration. Head black, maxillary and labial palpi yellow-brown, mandibles dark brown, antennae black, base of antennomere 1 yellow-brown, clypeus along anterior margin and antennal sockets narrowly yellow-brown, pronotum black-brown, elytra dark brown with strong coppery shine, abdomen black-brown, femora and tarsi yellow-brown, tibiae of all legs black.</paragraph>
<paragraph id="367236F8B035FFCFFF136D26A117137C" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Head rather flat, slightly wider than long (ratio 30: 28), parallel-sided, posterior angles marked, eyes slightly projecting, somewhat shorter than temples (ratio 10: 11), between eyes with four punctures, distance between medial interocular punctures five times the distance between medial and lateral puncture; temporal area with many coarse punctures; surface with microsculpture consisting of transverse waves.</paragraph>
<paragraph id="367236F8B035FFCFFF136C45A1C2131C" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Antennae long, when reclined exceeding posterior margin of pronotum by length of antennomere 11. All antennomeres longer than wide. Relative lengths of antennomeres: 1 = 12; 2 = 6; 3 = 7; 4–6 = 6; 7 = 5; 8 = 4.5; 9–10 = 4; 11 = 7.5.</paragraph>
<paragraph id="367236F8B035FFCFFF136C25A749139C" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Pronotum slightly wider than long (ratio 40: 38), hardly narrowed anteriad, each dorsal row with four punctures, distance between punctures 3–4 somewhat smaller than distance between punctures 2–3, each sublateral row with two punctures, puncture 1 situated on level between punctures 2–3 of dorsal rows, surface with microsculpture similar to that on head.</paragraph>
<paragraph id="367236F8B035FFCFFF136CA6A14C145C" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Scutellum finely and densely punctate, punctures equal in size to eye facets, separated by distance smaller than diameter of puncture.</paragraph>
<paragraph id="367236F8B035FFCFFF136B66A27114DC" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Elytra combined distinctly wider than long (ratio 52: 46), slightly widened posteriad, very densely and finely punctate, diameter of punctures somewhat larger than eye facets, distance between punctures smaller than diameter of punctures, surface without microsculpture; setation dark.</paragraph>
<paragraph id="367236F8B035FFCFFF136BE6A201153C" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Abdomen wide, slightly narrowed anteriad and posteriad from visible tergite III. First four visible tergites with two basal lines, elevated area between lines impunctate, punctation of visible tergites somewhat finer than that on elytra, becoming finer and sparser towards posterior margin of each tergite, surface between punctures without microsculpture; setation similar to that on elytra.</paragraph>
<paragraph id="367236F8B035FFCFFF136A05A29715FC" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">Legs. Metatibia vaguely longer than metatarsus (ratio 35: 33), lengths of metatarsomeres: 1 = 10; 2 = 5; 3 = 4.5; 4 = 4; 5 = 8.</paragraph>
<paragraph id="367236F8B035FFCFFF136AC6A07615BC" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">
Male. Protarsomeres 1–3 markedly dilated and sub-bilobed, each covered with modified pale setae ventrally, sternite IX (
<figureCitation id="AEF62A7DB035FFCFFE486AE6A17F15BC" box="[459,524,1465,1489]" captionStart="Figs" captionStartId="4.[151,190,1437,1456]" captionTargetBox="[227,1054,205,1388]" captionTargetId="figure-10@4.[217,1062,205,1409]" captionTargetPageId="4" captionText="Figs. 1–14. Philonthus species. 1–5 – P. arrowianus Bernhauer, 1931 (1 – aedeagus, ventral view; 2 – aedeagus, lateral view; 3 – apex of of paramere with sensory peg setae, ventral view; 4 – apical portion of male sternite VIII, ventral view; 5 – male sternite IX, ventral view). 6–9 – P. chappuisi Bernhauer, 1939 (6 – aedeagus, ventral view; 7 – aedeagus, lateral view; 8 – apex of paramere with sensory peg setae, ventral view; 9 – male sternite IX, ventral view). 10–14 – P. currax sp. nov. (10 – aedeagus, ventral view; 11 – aedeagus without paramere, ventral view; 12 – aedeagus, lateral view; 13 – apex of paramere with sensory peg setae; 14 – apical portion of male sternite VIII, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5325120" httpUri="https://zenodo.org/record/5325120/files/figure.png" pageId="3" pageNumber="134">Fig. 9</figureCitation>
), aedeagus (
<figureCitation id="AEF62A7DB035FFCFFD136AE6A18B15BC" box="[656,760,1465,1489]" captionStart="Figs" captionStartId="4.[151,190,1437,1456]" captionTargetBox="[227,1054,205,1388]" captionTargetId="figure-10@4.[217,1062,205,1409]" captionTargetPageId="4" captionText="Figs. 1–14. Philonthus species. 1–5 – P. arrowianus Bernhauer, 1931 (1 – aedeagus, ventral view; 2 – aedeagus, lateral view; 3 – apex of of paramere with sensory peg setae, ventral view; 4 – apical portion of male sternite VIII, ventral view; 5 – male sternite IX, ventral view). 6–9 – P. chappuisi Bernhauer, 1939 (6 – aedeagus, ventral view; 7 – aedeagus, lateral view; 8 – apex of paramere with sensory peg setae, ventral view; 9 – male sternite IX, ventral view). 10–14 – P. currax sp. nov. (10 – aedeagus, ventral view; 11 – aedeagus without paramere, ventral view; 12 – aedeagus, lateral view; 13 – apex of paramere with sensory peg setae; 14 – apical portion of male sternite VIII, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5325120" httpUri="https://zenodo.org/record/5325120/files/figure.png" pageId="3" pageNumber="134">Figs. 6–8</figureCitation>
).
</paragraph>
<paragraph id="367236F8B035FFCFFF136A86A22F159C" blockId="3.[113,1090,441,1618]" box="[144,348,1497,1521]" pageId="3" pageNumber="134">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="7ED76573B035FFCFFFF26AA6A07C165C" pageId="3" pageNumber="134" type="diagnosis">
<paragraph id="367236F8B035FFCFFFF26AA6A07C165C" blockId="3.[113,1090,441,1618]" pageId="3" pageNumber="134">
<emphasis id="04B9EAEAB035FFCFFFF26AA6A218167C" bold="true" box="[113,363,1529,1553]" pageId="3" pageNumber="134">Differential diagnosis.</emphasis>
<taxonomicName id="F1CD4D7BB035FFCFFEF26AA4A120167F" authorityName="Bernhauer" authorityYear="1939" box="[369,595,1531,1554]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="chappuisi">
<emphasis id="04B9EAEAB035FFCFFEF26AA4A120167F" box="[369,595,1531,1554]" italics="true" pageId="3" pageNumber="134">Philonthus chappuisi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="F1CD4D7BB035FFCFFD5F6AA4A02E167F" authorityName="Tottenham" authorityYear="1949" box="[732,861,1531,1554]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="134" phylum="Arthropoda" rank="species" species="trilobatus">
<emphasis id="04B9EAEAB035FFCFFD5F6AA4A02E167F" box="[732,861,1531,1554]" italics="true" pageId="3" pageNumber="134">P. trilobatus</emphasis>
</taxonomicName>
but differs from it by wider elytra, lighter legs and a different shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="7ED76573B035FFCFFFF26966A1BD163C" box="[113,718,1593,1618]" pageId="3" pageNumber="134" type="distribution">
<paragraph id="367236F8B035FFCFFFF26966A1BD163C" blockId="3.[113,1090,441,1618]" box="[113,718,1593,1618]" pageId="3" pageNumber="134">
<emphasis id="04B9EAEAB035FFCFFFF26966A277163C" bold="true" box="[113,260,1593,1617]" pageId="3" pageNumber="134">Distribution.</emphasis>
<collectingCountry id="4EDA7668B035FFCFFE896965A221163F" box="[266,338,1594,1618]" name="Kenya" pageId="3" pageNumber="134">Kenya</collectingCountry>
, known only from the
<typeStatus id="E976885AB035FFCFFDC76965A101163F" box="[580,626,1594,1618]" pageId="3" pageNumber="134">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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