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<document id="8A73185B8FCA1483A1DABEA136B60292" ID-CLB-Dataset="27410" ID-DOI="10.11646/zootaxa.4560.3.3" ID-GBIF-Dataset="91cdc97e-ac24-4299-a478-ccd02ca8e1b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="2627818" ID-ZooBank="120E7D55-C2C4-4A8C-A124-8A69A0B5DB21" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554323075341" checkinUser="plazi" docAuthor="Osawa, Masayuki &amp; Fujita, Yoshihisa" docDate="2019" docId="B60387DAD36EFF8FFF493891FEC5FB4D" docLanguage="en" docName="zootaxa.4560.3.3.pdf" docOrigin="Zootaxa 4560 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paguristes shidarai Asakura 2004" docType="treatment" docVersion="12" lastPageNumber="467" masterDocId="4A3AFFA2D36AFF8BFFDE3A46FF84FFF3" masterDocTitle="Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan" masterLastPageNumber="482" masterPageNumber="463" pageNumber="467" updateTime="1698718253014" updateUser="ExternalLinkService">
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<mods:title id="F5E735135105B2DA12E17473EA1685A4">Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan</mods:title>
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<emphasis id="0CDEEADED36EFF8FFF493891FD15FD01" bold="true" box="[151,657,727,754]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF493891FEEFFD02" bold="true" box="[151,363,727,753]" italics="true" pageId="4" pageNumber="467">Pseudopaguristes</emphasis>
<taxonomicName id="F9AA4D4FD36EFF8FFEAC3891FD15FD01" authority="Asakura, 2004" authorityName="Asakura" authorityYear="2004" box="[370,657,727,754]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="0CDEEADED36EFF8FFEAC3891FE52FD02" bold="true" box="[370,470,727,753]" italics="true" pageId="4" pageNumber="467">shidarai</emphasis>
<bibRefCitation id="5A3B4B3DD36EFF8FFE02389EFD15FD01" author="Asakura, A." box="[476,657,728,754]" pageId="4" pageNumber="467" pagination="153 - 168" refId="ref9561" refString="Asakura, A. (2004) Pseudopaguristes shidarai, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) from Japan, the fourth species of the genus. Proceedings of the Biological Society of Washington, 117 (2), 153 - 168." type="journal article" year="2004">Asakura, 2004</bibRefCitation>
</taxonomicName>
</emphasis>
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<paragraph id="3E1536CCD36EFF8FFF4938BCFF79FCE0" blockId="4.[151,657,727,787]" box="[151,253,762,787]" pageId="4" pageNumber="467">
(
<figureCitation id="A6912A49D36EFF8FFF4138BCFF70FCE0" box="[159,244,762,787]" captionStart="FIGURE 1" captionStartId="5.[151,250,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@5.[151,1436,193,1705]" captionTargetPageId="5" captionText="FIGURE 1. Entire animal, dorsal view. A, Aniculus retipes Lewinsohn, 1982, male (sl 11.9 mm), RUMF-ZC-6006 (IE141); B, Paguristes jalur Morgan, 1992, male (sl 5.4 mm), RUMF-ZC-6012 (IE69); C, Pseudopaguristes shidarai Asakura, 2004, male (sl 2.6 mm), RUMF-ZC-6020 (HD2017-140); D, Catapaguroides foresti McLaughlin, 2002, male (sl 2.0 mm), RUMFZC-6023 (IE173); E, Catapaguroides iejimensis Osawa &amp; Takeda, 2004, male (sl 1.8 mm), RUMF-ZC-6026 (IE177); F, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, male (sl 2.1 mm), RUMF-ZC-6027 (IE60); G, Catapaguroides longior Komai, Yamada &amp; Shirakawa, 2010, female (sl 1.2 mm), RUMF-ZC-6029 (HD2017-107)." httpUri="https://zenodo.org/record/2627820/files/figure.png" pageId="4" pageNumber="467">Fig. 1C</figureCitation>
)
</paragraph>
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<subSubSection id="76B06547D36EFF8FFF493904FB45FCAB" box="[151,1217,834,856]" pageId="4" pageNumber="467" type="reference_group">
<paragraph id="3E1536CCD36EFF8FFF493904FB45FCAB" blockId="4.[151,1217,834,856]" box="[151,1217,834,856]" pageId="4" pageNumber="467">
<treatmentCitationGroup id="1EBA11E2D36EFF8FFF493904FB45FCAB" box="[151,1217,834,856]" pageId="4" pageNumber="467">
<materialsCitation id="8EC23C91D36EFF8FFF493904FB39FCAB" ID-GBIF-Occurrence="2465041903" box="[151,1213,834,856]" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Miyako Island, Ryukyu Islands" typeStatus="type">
<emphasis id="0CDEEADED36EFF8FFF493904FECFFCAB" box="[151,331,834,856]" italics="true" pageId="4" pageNumber="467">Pseudopaguristes</emphasis>
<taxonomicName id="F9AA4D4FD36EFF8FFE8F3904FDF1FCAB" authority="Asakura, 2004: 153" authorityName="Asakura" authorityPageNumber="153" authorityYear="2004" box="[337,629,834,856]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="0CDEEADED36EFF8FFE8F3904FE22FCAB" box="[337,422,834,856]" italics="true" pageId="4" pageNumber="467">shidarai</emphasis>
<treatmentCitation id="BF0B10DDD36EFF8FFE723904FDF1FCAB" author="Asakura, A." box="[428,629,834,856]" page="153" pageId="4" pageNumber="467" year="2004">
<bibRefCitation id="5A3B4B3DD36EFF8FFE723904FDC0FCAB" author="Asakura, A." box="[428,580,834,856]" pageId="4" pageNumber="467" pagination="153 - 168" refId="ref9561" refString="Asakura, A. (2004) Pseudopaguristes shidarai, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) from Japan, the fourth species of the genus. Proceedings of the Biological Society of Washington, 117 (2), 153 - 168." type="journal article" year="2004">Asakura, 2004</bibRefCitation>
: 153
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, fig. 112 (type locality: Miyako Island, Ryukyu Islands)
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="76B06547D36EFF8FFF4939C3FD82FC15" pageId="4" pageNumber="467" type="materials_examined">
<paragraph id="3E1536CCD36EFF8FFF4939C3FD82FC15" blockId="4.[151,1436,901,1214]" pageId="4" pageNumber="467">
<materialsCitation id="8EC23C91D36EFF8FFF4939C3FE29FC31" ID-GBIF-Occurrence="2465041902" collectingDate="2017-05-22" collectionCode="HD" country="Japan" location="main tube" pageId="4" pageNumber="467" specimenCode="RUMF- ZC-6020" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Island, Hedo Dome cave">
<emphasis id="0CDEEADED36EFF8FFF4939C3FCA8FC6D" bold="true" box="[151,812,901,926]" pageId="4" pageNumber="467">
Material examined.
<collectingRegion id="FC6EF82ED36EFF8FFE5739C3FCA3FC6D" box="[393,807,901,926]" pageId="4" pageNumber="467">Okinawa Island, Hedo Dome cave</collectingRegion>
:
</emphasis>
<location id="3B756017D36EFF8FFCEB39C3FC2CFC6D" LSID="urn:lsid:plazi:treatment:B60387DAD36EFF8FFF493891FEC5FB4D:3B756017D36EFF8FFCEB39C3FC2CFC6D" box="[821,936,901,926]" country="Japan" name="main tube" pageId="4" pageNumber="467" stateProvince="Okinawa Island, Hedo Dome cave">main tube</location>
,
<date id="4A14100CD36EFF8FFC6A39C0FBCBFC6D" box="[948,1103,902,926]" pageId="4" pageNumber="467" value="2017-05-22">
<collectingDate id="5A50E9E4D36EFF8FFC6A39C0FBCBFC6D" box="[948,1103,902,926]" pageId="4" pageNumber="467" value="2017-05-22">22 May 2017</collectingDate>
</date>
,
<specimenCount id="28ACFD45D36EFF8FFB8539C0FB23FC6D" box="[1115,1191,901,926]" pageId="4" pageNumber="467" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFB0A39C3FAABFC6D" box="[1236,1327,901,926]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="467" unit="mm" value="2.6">2.6 mm</quantity>
),
<specimenCode id="6E0C9EB7D36EFF8FFA9D39C0FF78FC31" pageId="4" pageNumber="467">RUMF- ZC-6020</specimenCode>
(HD2017-140)
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.
<materialsCitation id="8EC23C91D36EFF8FFE6739EFFD86FC15" ID-GBIF-Occurrence="2465041906" collectingDate="2017-09-01" collectionCode="SH" country="Japan" location="entrance" pageId="4" pageNumber="467" specimenCode="RUMF-ZC-6021" specimenCount="1" specimenCount-female="1" stateProvince="Shimoji Island, Akuma-no-Yakata cave">
<emphasis id="0CDEEADED36EFF8FFE6739EFFC1EFC31" bold="true" box="[441,922,937,962]" pageId="4" pageNumber="467">
<collectingRegion id="FC6EF82ED36EFF8FFE6739EFFC11FC31" box="[441,917,937,962]" pageId="4" pageNumber="467">Shimoji Island, Akuma-no-Yakata cave</collectingRegion>
:
</emphasis>
<location id="3B756017D36EFF8FFC7C39ECFB80FC32" LSID="urn:lsid:plazi:treatment:B60387DAD36EFF8FFF493891FEC5FB4D:3B756017D36EFF8FFC7C39ECFB80FC32" box="[930,1028,938,961]" country="Japan" name="entrance" pageId="4" pageNumber="467" stateProvince="Shimoji Island, Akuma-no-Yakata cave">entrance</location>
,
<date id="4A14100CD36EFF8FFBD139ECFB64FC31" box="[1039,1248,937,962]" pageId="4" pageNumber="467" value="2017-09-01">
<collectingDate id="5A50E9E4D36EFF8FFBD139ECFB64FC31" box="[1039,1248,937,962]" pageId="4" pageNumber="467" value="2017-09-01">1 September 2017</collectingDate>
</date>
,
<specimenCount id="28ACFD45D36EFF8FFB3539ECFAC9FC31" box="[1259,1357,937,962]" pageId="4" pageNumber="467" type="female">1 female</specimenCount>
(sl
<quantity id="F9529B29D36EFF8FFAA739ECFF43FC15" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="467" unit="mm" value="2.3">2.3 mm</quantity>
),
<specimenCode id="6E0C9EB7D36EFF8FFF04398BFE1CFC15" box="[218,408,973,998]" pageId="4" pageNumber="467">RUMF-ZC-6021</specimenCode>
(SH105)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36EFF8FFF1939B7FC14FBA1" pageId="4" pageNumber="467" type="distribution">
<paragraph id="3E1536CCD36EFF8FFF1939B7FBD3FBDD" blockId="4.[151,1436,901,1214]" pageId="4" pageNumber="467">
<materialsCitation id="8EC23C91D36EFF8FFF1939B7FEF1FBDE" ID-GBIF-Occurrence="2465041908" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">
<emphasis id="0CDEEADED36EFF8FFF1939B7FEE4FBF9" bold="true" box="[199,352,1009,1034]" pageId="4" pageNumber="467">Distribution.</emphasis>
Ryukyu Islands (Okinawa Island, Kume Island, Miyako Island, and Shimoji Island) in southwestern Japan
</materialsCitation>
;
<materialsCitation id="8EC23C91D36EFF8FFE5E3E50FD17FBDD" ID-GBIF-Occurrence="2465041904" box="[384,659,1045,1070]" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">
<quantity id="F9529B29D36EFF8FFE5E3E50FE67FBDE" box="[384,483,1046,1070]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="4" pageNumber="467" unit="m" value="30.0" valueMax="40.0" valueMin="20.0">2040 m</quantity>
(
<bibRefCitation id="5A3B4B3DD36EFF8FFE2F3E53FD17FBDD" author="Asakura, A." box="[497,659,1045,1070]" pageId="4" pageNumber="467" pagination="153 - 168" refId="ref9561" refString="Asakura, A. (2004) Pseudopaguristes shidarai, a new species of hermit crab (Crustacea: Decapoda: Diogenidae) from Japan, the fourth species of the genus. Proceedings of the Biological Society of Washington, 117 (2), 153 - 168." type="journal article" year="2004">Asakura 2004</bibRefCitation>
</materialsCitation>
;
<materialsCitation id="8EC23C91D36EFF8FFD403E53FC23FBDD" ID-GBIF-Occurrence="2465041901" box="[670,935,1045,1070]" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">
<bibRefCitation id="5A3B4B3DD36EFF8FFD403E53FC23FBDD" author="Rahayu, D. L. &amp; Osawa, M." box="[670,935,1045,1070]" pageId="4" pageNumber="467" pagination="176 - 190" refId="ref13115" refString="Rahayu, D. L. &amp; Osawa, M. (2012) Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa, 3367, 176 - 190." type="journal article" year="2012">Rahayu &amp; Osawa 2012</bibRefCitation>
</materialsCitation>
;
<materialsCitation id="8EC23C91D36EFF8FFC6C3E50FBD7FBDD" ID-GBIF-Occurrence="2465041917" box="[946,1107,1045,1070]" country="Japan" pageId="4" pageNumber="467" specimenCount="1" stateProvince="Ryukyu Islands (okinawa Island, Kume Island, Miyako Island, And Shimoji Island)">present study)</materialsCitation>
.
</paragraph>
<paragraph id="3E1536CCD36EFF8FFF193E7FFC14FBA1" blockId="4.[151,1436,901,1214]" box="[199,912,1081,1106]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193E7FFEAEFBA1" bold="true" box="[199,298,1081,1106]" pageId="4" pageNumber="467">Habitat.</emphasis>
Coral reefs; fore-reef slopes (
<bibRefCitation id="5A3B4B3DD36EFF8FFDA63E7FFC00FBA1" author="Rahayu, D. L. &amp; Osawa, M." box="[632,900,1081,1106]" pageId="4" pageNumber="467" pagination="176 - 190" refId="ref13115" refString="Rahayu, D. L. &amp; Osawa, M. (2012) Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa, 3367, 176 - 190." type="journal article" year="2012">Rahayu &amp; Osawa 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D36EFF8FFF193E1BFEC5FB4D" pageId="4" pageNumber="467" type="discussion">
<paragraph id="3E1536CCD36EFF8FFF193E1BFEC5FB4D" blockId="4.[151,1436,901,1214]" pageId="4" pageNumber="467">
<emphasis id="0CDEEADED36EFF8FFF193E1BFEBFFB85" bold="true" box="[199,315,1117,1142]" pageId="4" pageNumber="467">Remarks.</emphasis>
In the Ryukyu Islands,
<taxonomicName id="F9AA4D4FD36EFF8FFD9C3E18FD33FB86" authorityName="Asakura" authorityYear="2004" box="[578,695,1117,1141]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shidarai">
<emphasis id="0CDEEADED36EFF8FFD9C3E18FD33FB86" box="[578,695,1117,1141]" italics="true" pageId="4" pageNumber="467">P. shidarai</emphasis>
</taxonomicName>
has been recorded from cryptic environments of the shallow coral reefs such as small crevices and horizontal holes, and submarine caves (
<bibRefCitation id="5A3B4B3DD36EFF8FFC133EC7FB5FFB69" author="Rahayu, D. L. &amp; Osawa, M." box="[973,1243,1153,1178]" pageId="4" pageNumber="467" pagination="176 - 190" refId="ref13115" refString="Rahayu, D. L. &amp; Osawa, M. (2012) Hermit crabs (Crustacea: Decapoda: Diogenidae) from the KUMEJIMA 2009 Expedition, Japan. Zootaxa, 3367, 176 - 190." type="journal article" year="2012">Rahayu &amp; Osawa 2012</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD36EFF8FFB363EC4FF57FB4D" author="Osawa, M. &amp; Fujita, Y." pageId="4" pageNumber="467" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
;
<tableCitation id="73280377D36EFF8FFF033EE3FEB7FB4D" box="[221,307,1189,1214]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="4" pageNumber="467">Table 2</tableCitation>
).
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</treatment>
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