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<document id="E189E2971E294007A4F9275BA95F87F8" ID-CLB-Dataset="27410" ID-DOI="10.11646/zootaxa.4560.3.3" ID-GBIF-Dataset="91cdc97e-ac24-4299-a478-ccd02ca8e1b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="2627818" ID-ZooBank="120E7D55-C2C4-4A8C-A124-8A69A0B5DB21" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554323075341" checkinUser="plazi" docAuthor="Osawa, Masayuki &amp; Fujita, Yoshihisa" docDate="2019" docId="B60387DAD367FF85FF493C82FC63FE5F" docLanguage="en" docName="zootaxa.4560.3.3.pdf" docOrigin="Zootaxa 4560 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paguristes speciosa McLaughlin &amp; Haig 1989" docType="treatment" docVersion="12" lastPageNumber="477" masterDocId="4A3AFFA2D36AFF8BFFDE3A46FF84FFF3" masterDocTitle="Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan" masterLastPageNumber="482" masterPageNumber="463" pageNumber="476" updateTime="1698718253014" updateUser="ExternalLinkService">
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<mods:title id="5954FD8D3DB32F8925AED7A19DC5067F">Submarine cave hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from three islands of the Ryukyu Islands, southwestern Japan</mods:title>
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<mods:namePart id="AA7547057722DA8D4CEBED70D174E476">Osawa, Masayuki</mods:namePart>
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<mods:namePart id="1934A296579D4063B4A7CF7C8B8CF164">Fujita, Yoshihisa</mods:namePart>
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<treatment id="B60387DAD367FF85FF493C82FC63FE5F" ID-DOI="http://doi.org/10.5281/zenodo.5308574" ID-GBIF-Taxon="154533463" ID-Zenodo-Dep="5308574" LSID="urn:lsid:plazi:treatment:B60387DAD367FF85FF493C82FC63FE5F" httpUri="http://treatment.plazi.org/id/B60387DAD367FF85FF493C82FC63FE5F" lastPageId="14" lastPageNumber="477" pageId="13" pageNumber="476">
<subSubSection id="76B06547D367FF86FF493C82FC88F92C" box="[151,780,1732,1759]" pageId="13" pageNumber="476" type="nomenclature">
<paragraph id="3E1536CCD367FF86FF493C82FC88F92C" blockId="13.[151,780,1732,1792]" box="[151,780,1732,1759]" pageId="13" pageNumber="476">
<heading id="655D81A0D367FF86FF493C82FC88F92C" bold="true" box="[151,780,1732,1759]" fontSize="11" level="1" pageId="13" pageNumber="476" reason="1">
<emphasis id="0CDEEADED367FF86FF493C82FC88F92C" bold="true" box="[151,780,1732,1759]" pageId="13" pageNumber="476">
<emphasis id="0CDEEADED367FF86FF493C82FED5F92D" bold="true" box="[151,337,1732,1758]" italics="true" pageId="13" pageNumber="476">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD367FF86FE863C82FC88F92C" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[344,780,1732,1759]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="0CDEEADED367FF86FE863C82FE39F92D" bold="true" box="[344,445,1732,1758]" italics="true" pageId="13" pageNumber="476">speciosa</emphasis>
<bibRefCitation id="5A3B4B3DD367FF86FE1A3C83FC88F92C" author="McLaughlin, P. A. &amp; Haig, J." box="[452,780,1733,1759]" pageId="13" pageNumber="476" pagination="123 - 171" refId="ref11585" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="76B06547D367FF86FF493CA1FF7FF8F3" box="[151,251,1767,1792]" pageId="13" pageNumber="476" type="description">
<paragraph id="3E1536CCD367FF86FF493CA1FF7FF8F3" blockId="13.[151,780,1732,1792]" box="[151,251,1767,1792]" pageId="13" pageNumber="476">
(
<figureCitation id="A6912A49D367FF86FF413CA1FF77F8F3" box="[159,243,1767,1792]" captionStart="FIGURE 3" captionStartId="11.[151,250,1378,1400]" captionTargetBox="[151,1436,195,1356]" captionTargetId="figure@11.[151,1436,195,1356]" captionTargetPageId="11" captionText="FIGURE 3. Entire animal, dorsal view. A, Pylopaguropsis bellula Osawa &amp; Okuno, 2007, male (sl 4.1 mm), RUMF-ZC-6042 (IE17); B, Pylopaguropsis fimbriata McLaughlin &amp; Haig, 1989, ovigerous female (sl 4.4 mm), RUMF-ZC-6046 (IE179); C, Pylopaguropsis granulata Asakura, 2000, female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31); D, Pylopaguropsis keijii McLaughlin &amp; Haig, 1989, male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87); E, Pylopaguropsis speciosa McLaughlin &amp; Haig, 1989, ovigerous female (sl 3.7 mm), RUMF-ZC-6058 (SH104)." httpUri="https://zenodo.org/record/2627824/files/figure.png" pageId="13" pageNumber="476">Fig. 3E</figureCitation>
)
</paragraph>
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<subSubSection id="76B06547D367FF86FF493D69FCABF870" pageId="13" pageNumber="476" type="reference_group">
<paragraph id="3E1536CCD367FF86FF493D69FCABF870" blockId="13.[151,1436,1839,1923]" pageId="13" pageNumber="476">
<treatmentCitationGroup id="1EBA11E2D367FF86FF493D69FCABF870" pageId="13" pageNumber="476">
<emphasis id="0CDEEADED367FF86FF493D69FEB1F8B6" box="[151,309,1839,1861]" italics="true" pageId="13" pageNumber="476">Pylopaguropsis</emphasis>
<taxonomicName id="F9AA4D4FD367FF86FEE53D77FD58F8B6" authority="McLaughlin &amp; Haig, 1989: 153" authorityName="McLaughlin &amp; Haig" authorityPageNumber="153" authorityYear="1989" box="[315,732,1839,1862]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="0CDEEADED367FF86FEE53D77FE16F8B5" box="[315,402,1841,1862]" italics="true" pageId="13" pageNumber="476">speciosa</emphasis>
<treatmentCitation id="BF0B10DDD367FF86FE473D69FD58F8B6" author="McLaughlin, P. A. &amp; Haig, J." box="[409,732,1839,1861]" page="153" pageId="13" pageNumber="476" year="1989">
<bibRefCitation id="5A3B4B3DD367FF86FE473D69FD2EF8B6" author="McLaughlin, P. A. &amp; Haig, J." box="[409,682,1839,1861]" pageId="13" pageNumber="476" pagination="123 - 171" refId="ref11585" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra Henderson, P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22 (2), 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
: 153
</treatmentCitation>
</taxonomicName>
, figs. 1c, i, 2c, 3e, 5e, 7f, 9e, 11e, 13e (
<typeStatus id="E111886ED367FF86FBB73D69FB12F8B6" box="[1129,1174,1839,1861]" pageId="13" pageNumber="476">type</typeStatus>
locality: Okinawa Island, Ryukyu Islands).—
<treatmentCitation id="BF0B10DDD367FF86FE573D08FDC7F897" author="Asakura, A." box="[393,579,1870,1892]" page="78" pageId="13" pageNumber="476" year="2000">
<bibRefCitation id="5A3B4B3DD367FF86FE573D08FD9AF897" author="Asakura, A." box="[393,542,1870,1892]" pageId="13" pageNumber="476" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
: 78
</treatmentCitation>
, figs. 1A, B, 4, 5, 19A, 20.—
<treatmentCitation id="BF0B10DDD367FF86FCB53D08FB17F897" author="Osawa, M. &amp; Takeda, M." box="[875,1171,1870,1892]" page="1111" pageId="13" pageNumber="476" year="2004">
<bibRefCitation id="5A3B4B3DD367FF86FCB53D08FBD3F897" author="Osawa, M. &amp; Takeda, M." box="[875,1111,1870,1892]" pageId="13" pageNumber="476" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
: 1111
</treatmentCitation>
.
<taxonomicName id="F9AA4D4FD367FF86FF493D28FF69F870" ID-CoL="a45bf7bc-019d-4e9e-a238-02e442a26045" box="[151,237,1902,1923]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="0CDEEADED367FF86FF493D28FF69F870" box="[151,237,1902,1923]" italics="true" pageId="13" pageNumber="476">Pagurus</emphasis>
</taxonomicName>
sp.—
<treatmentCitation id="BF0B10DDD367FF86FEF93D28FDA1F870" author="Jones, D. S. &amp; Morgan, G. J." box="[295,549,1901,1923]" page="4" pageId="13" pageNumber="476" year="1994">
<bibRefCitation id="5A3B4B3DD367FF86FEF93D28FD8AF870" author="Jones, D. S. &amp; Morgan, G. J." box="[295,526,1901,1923]" pageId="13" pageNumber="476" refId="ref10364" refString="Jones, D. S. &amp; Morgan, G. J. (1994) A Field Guide to Crustaceans of Australian Waters. Reed, Chatswood, 216 pp." type="book" year="1994">Jones &amp; Morgan 1994</bibRefCitation>
: 4
</treatmentCitation>
–5, 124, unnumbered figs.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="76B06547D367FF85FF493DE9FC98FF27" lastPageId="14" lastPageNumber="477" pageId="13" pageNumber="476" type="materials_examined">
<paragraph id="3E1536CCD367FF86FF493DE9FA18F81F" blockId="13.[151,1436,1967,2029]" pageId="13" pageNumber="476">
<materialsCitation id="8EC23C91D367FF86FF493DE9FC08F81E" ID-GBIF-Occurrence="2465041897" collectingDate="2016-08-12" collectionCode="HD" country="Japan" location="main tube, 30 m from entrance, area of rocks" pageId="13" pageNumber="476" specimenCode="RUMF-ZC-6056" specimenCount="3" specimenCount-male="3" stateProvince="Okinawa Island, Hedo Dome cave">
<emphasis id="0CDEEADED367FF86FF493DE9FCA6F83B" bold="true" box="[151,802,1967,1992]" pageId="13" pageNumber="476">
Material examined.
<collectingRegion id="FC6EF82ED367FF86FE583DE9FC99F83B" box="[390,797,1967,1992]" pageId="13" pageNumber="476">Okinawa Island, Hedo Dome cave</collectingRegion>
:
</emphasis>
<location id="3B756017D367FF86FCF63DE9FA9AF83B" LSID="urn:lsid:plazi:treatment:B60387DAD367FF85FF493C82FC63FE5F:3B756017D367FF86FCF63DE9FA9AF83B" box="[808,1310,1967,1992]" country="Japan" name="main tube, 30 m from entrance, area of rocks" pageId="13" pageNumber="476" stateProvince="Okinawa Island, Hedo Dome cave">
main tube,
<quantity id="F9529B29D367FF86FC7A3DF6FC59F834" box="[932,989,1968,1992]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="476" unit="m" value="30.0">30 m</quantity>
from entrance, area of rocks
</location>
,
<date id="4A14100CD367FF86FAF73DF6FF56F81E" pageId="13" pageNumber="476" value="2016-08-12">
<collectingDate id="5A50E9E4D367FF86FAF73DF6FF56F81E" pageId="13" pageNumber="476" value="2016-08-12">12 August 2016</collectingDate>
</date>
,
<specimenCount id="28ACFD45D367FF86FF003D92FEB1F81E" box="[222,309,2004,2029]" pageId="13" pageNumber="476" type="male">3 males</specimenCount>
(sl
<quantity id="F9529B29D367FF86FEBF3D92FE69F81F" box="[353,493,2004,2028]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="13" pageNumber="476" unit="mm" value="2.55" valueMax="2.9" valueMin="2.2">2.2–2.9 mm</quantity>
),
<specimenCode id="6E0C9EB7D367FF86FDDE3D92FD3AF81E" box="[512,702,2004,2029]" pageId="13" pageNumber="476">RUMF-ZC-6056</specimenCode>
(HD
<date id="4A14100CD367FF86FD273D92FCDDF81E" box="[761,857,2004,2029]" pageId="13" pageNumber="476" value="2016" valueMax="2086">2016-86</date>
–88)
</materialsCitation>
;
<materialsCitation id="8EC23C91D367FF86FC463D92FA1CF81F" ID-GBIF-Occurrence="2465041918" box="[920,1432,2004,2029]" collectingDate="2017-05-21" country="Japan" location="main tube" pageId="13" pageNumber="476" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Island, Hedo Dome Cave">
<location id="3B756017D367FF86FC463D92FB8FF81E" LSID="urn:lsid:plazi:treatment:B60387DAD367FF85FF493C82FC63FE5F:3B756017D367FF86FC463D92FB8FF81E" box="[920,1035,2004,2029]" country="Japan" name="main tube" pageId="13" pageNumber="476" stateProvince="Okinawa Island, Hedo Dome Cave">main tube</location>
,
<date id="4A14100CD367FF86FBC93D92FB35F81F" box="[1047,1201,2004,2029]" pageId="13" pageNumber="476" value="2017-05-21">
<collectingDate id="5A50E9E4D367FF86FBC93D92FB35F81F" box="[1047,1201,2004,2029]" pageId="13" pageNumber="476" value="2017-05-21">21 May 2017</collectingDate>
</date>
,
<specimenCount id="28ACFD45D367FF86FB633D92FA8DF81E" box="[1213,1289,2004,2029]" pageId="13" pageNumber="476" type="male">1 male</specimenCount>
(sl
<quantity id="F9529B29D367FF86FAEB3D92FA14F81F" box="[1333,1424,2004,2028]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="476" unit="mm" value="3.1">3.1 mm</quantity>
)
</materialsCitation>
,
</paragraph>
<paragraph id="3E1536CCD364FF85FF493AD1FC98FF27" blockId="14.[151,1437,151,429]" pageId="14" pageNumber="477">
<materialsCitation id="8EC23C91D364FF85FF493AD1FC9CFF27" ID-GBIF-Occurrence="2465041896" collectingDate="2017-09-01" collectionCode="HD" country="Japan" location="entrance" pageId="14" pageNumber="477" specimenCount="1" stateProvince="Shimoji Island, Akuma-no-Yakata cave">
RUMF-ZC-6057 (HD
<date id="4A14100CD364FF85FE473AD1FE7DFF43" box="[409,505,151,176]" pageId="14" pageNumber="477" value="2017" valueMax="2086">2017-86</date>
).
<emphasis id="0CDEEADED364FF85FDCB3AD1FB94FF43" bold="true" box="[533,1040,151,176]" pageId="14" pageNumber="477">
<collectingRegion id="FC6EF82ED364FF85FDCB3AD1FB8FFF43" box="[533,1035,151,176]" pageId="14" pageNumber="477">Shimoji Island, Akuma-no-Yakata cave</collectingRegion>
:
</emphasis>
<location id="3B756017D364FF85FBFF3ADEFB07FF5C" LSID="urn:lsid:plazi:treatment:B60387DAD367FF85FF493C82FC63FE5F:3B756017D364FF85FBFF3ADEFB07FF5C" box="[1057,1155,152,175]" country="Japan" name="entrance" pageId="14" pageNumber="477" stateProvince="Shimoji Island, Akuma-no-Yakata cave">entrance</location>
,
<date id="4A14100CD364FF85FB463ADEFAFDFF43" box="[1176,1401,151,176]" pageId="14" pageNumber="477" value="2017-09-01">
<collectingDate id="5A50E9E4D364FF85FB463ADEFAFDFF43" box="[1176,1401,151,176]" pageId="14" pageNumber="477" value="2017-09-01">1 September 2017</collectingDate>
</date>
,
<specimenCount id="28ACFD45D364FF85FA503ADEFEDCFF26" pageId="14" pageNumber="477">1 ovigerous female</specimenCount>
(sl
<quantity id="F9529B29D364FF85FE5C3AFAFE59FF27" box="[386,477,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="477" unit="mm" value="3.7">3.7 mm</quantity>
), RUMF-ZC-6058 (SH104)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="76B06547D364FF85FF193A99FA08FEEF" pageId="14" pageNumber="477" type="distribution">
<paragraph id="3E1536CCD364FF85FF193A99FA19FF0B" blockId="14.[151,1437,151,429]" box="[199,1437,223,248]" pageId="14" pageNumber="477">
<emphasis id="0CDEEADED364FF85FF193A99FEE4FF0B" bold="true" box="[199,352,223,248]" pageId="14" pageNumber="477">Distribution.</emphasis>
Japan (Ryukyu and Izu Islands) and Australia;
<quantity id="F9529B29D364FF85FCAF3A99FC50FF04" box="[881,980,223,248]" metricMagnitude="1" metricUnit="m" metricValue="3.55" metricValueMax="6.1" metricValueMin="1.0" pageId="14" pageNumber="477" unit="m" value="35.5" valueMax="61.0" valueMin="10.0">10–61 m</quantity>
(
<bibRefCitation id="5A3B4B3DD364FF85FC3F3A99FB06FF0B" author="Asakura, A." box="[993,1154,223,248]" pageId="14" pageNumber="477" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FB503AA6FA15FF0B" author="Osawa, M. &amp; Takeda, M." box="[1166,1425,223,248]" pageId="14" pageNumber="477" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
).
</paragraph>
<paragraph id="3E1536CCD364FF85FF193B42FA08FEEF" blockId="14.[151,1437,151,429]" box="[199,1420,260,285]" pageId="14" pageNumber="477">
<emphasis id="0CDEEADED364FF85FF193B42FEAEFEEE" bold="true" box="[199,298,260,285]" pageId="14" pageNumber="477">Habitat.</emphasis>
Coral reefs, often in rocky areas or submarine caves (
<bibRefCitation id="5A3B4B3DD364FF85FC583B42FBF5FEEF" author="Kato, S. &amp; Okuno, J." box="[902,1137,260,285]" pageId="14" pageNumber="477" refId="ref10558" refString="Kato, S. &amp; Okuno, J. (2001) Shrimps and crabs of Hachijo Island. TBS-Britannica Co. Ltd, Tokyo, 160 pp. [in Japanese]" type="book" year="2001">Kato &amp; Okuno 2001</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FBA53B42FA04FEEF" author="Osawa, M. &amp; Takeda, M." box="[1147,1408,260,285]" pageId="14" pageNumber="477" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="76B06547D364FF85FF193B61FC63FE5F" pageId="14" pageNumber="477" type="discussion">
<paragraph id="3E1536CCD364FF85FF193B61FC63FE5F" blockId="14.[151,1437,151,429]" pageId="14" pageNumber="477">
<emphasis id="0CDEEADED364FF85FF193B61FEBFFEB3" bold="true" box="[199,315,295,320]" pageId="14" pageNumber="477">Remarks.</emphasis>
<emphasis id="0CDEEADED364FF85FE9C3B6EFDDCFECC" box="[322,600,296,320]" italics="true" pageId="14" pageNumber="477">
Pylopaguropsis
<taxonomicName id="F9AA4D4FD364FF85FE293B6EFDDCFECC" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[503,600,296,319]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="477" phylum="Arthropoda" rank="species" species="speciosa">speciosa</taxonomicName>
</emphasis>
has been often recorded from coral reefs including submarine caves in the Ryukyu Islands (
<bibRefCitation id="5A3B4B3DD364FF85FE843B0AFE7BFE97" author="Asakura, A." box="[346,511,332,357]" pageId="14" pageNumber="477" pagination="70 - 108" refId="ref9456" refString="Asakura, A. (2000) A review of Japanese species of Pylopaguropsis Alcock, 1905 (Decapoda: Anomura: Paguridae). Crustacean Research, 29, 70 - 108. https: // doi. org / 10.18353 / crustacea. 29.0 _ 70" type="journal article" year="2000">Asakura 2000</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FDD33B0AFC9FFE97" author="Osawa, M. &amp; Takeda, M." box="[525,795,332,357]" pageId="14" pageNumber="477" pagination="1097 - 1132" refId="ref12989" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38 (9), 1097 - 1132. https: // doi. org / 10.1080 / 0022293031000079589" type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
;
<bibRefCitation id="5A3B4B3DD364FF85FCF73B0AFBA3FE96" author="Osawa, M. &amp; Fujita, Y." box="[809,1063,332,357]" pageId="14" pageNumber="477" pagination="37 - 56" refId="ref12816" refString="Osawa, M. &amp; Fujita, Y. (2016) Stomatopods and decapods of Axiidea, Gebiidea and Anomura (Crustacea: Malacostraca) from Irabu-jima and Shimojijima Islands, Miyako Group, southern Ryukyus, Japan. Fauna Ryukyuana, 28, 37 - 56. [in Japanese with English abstract]" type="journal article" year="2016">Osawa &amp; Fujita 2016</bibRefCitation>
;
<tableCitation id="73280377D364FF85FBE83B0AFB0BFE97" box="[1078,1167,332,357]" captionStart="TABLE 2" captionStartId="6.[147,233,596,618]" captionText="TABLE 2. Submarine caves surveyed and hermit crab species collected in three islands of the Ryukyu Islands, southwestern Japan." pageId="14" pageNumber="477">Table 2</tableCitation>
). The coloration of the species is very characteristic and readily distinguished from other congeners; the right chela is magenta and the left cheliped and ambulatory legs are bright orange with white stripes (
<figureCitation id="A6912A49D364FF85FC5D3BD2FC5FFE5F" box="[899,987,404,429]" captionStart="FIGURE 3" captionStartId="11.[151,250,1378,1400]" captionTargetBox="[151,1436,195,1356]" captionTargetId="figure@11.[151,1436,195,1356]" captionTargetPageId="11" captionText="FIGURE 3. Entire animal, dorsal view. A, Pylopaguropsis bellula Osawa &amp; Okuno, 2007, male (sl 4.1 mm), RUMF-ZC-6042 (IE17); B, Pylopaguropsis fimbriata McLaughlin &amp; Haig, 1989, ovigerous female (sl 4.4 mm), RUMF-ZC-6046 (IE179); C, Pylopaguropsis granulata Asakura, 2000, female (sl 4.8 mm), RUMF-ZC-6052 (HD2017-31); D, Pylopaguropsis keijii McLaughlin &amp; Haig, 1989, male (sl 1.9 mm), RUMF-ZC-6055 (HD2017-87); E, Pylopaguropsis speciosa McLaughlin &amp; Haig, 1989, ovigerous female (sl 3.7 mm), RUMF-ZC-6058 (SH104)." httpUri="https://zenodo.org/record/2627824/files/figure.png" pageId="14" pageNumber="477">Fig. 3E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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