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<document id="C30B542E4D6073E286C6C1E8E7E18BDE" ID-CLB-Dataset="20233" ID-DOI="10.5852/ejt.2022.797.1667" ID-GBIF-Dataset="e4b9a92e-23ef-4d55-85c9-ccabcbd8316e" ID-ISSN="2118-9773" ID-Zenodo-Dep="6299487" ID-ZooBank="8DF57743-9C53-4265-BCB5-743276A3A16C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645807222198" checkinUser="felipe" docAuthor="Telnov, Dmitry" docDate="2022" docId="6D52212B21392F65FDB54A4268B4FDC1" docLanguage="en" docName="ejt-797_1-100.pdf" docOrigin="European Journal of Taxonomy 797" docSource="http://dx.doi.org/10.5852/ejt.2022.797.1667" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Macrotomoderus dali Telnov 2022, sp. nov." docType="treatment" docUuid="F4C22B7A-9A8F-4FAE-B190-B28C99E6AB17" docUuidSource="ZooBank" docVersion="10" lastPageNumber="23" masterDocId="916B5953212B2F73FFD24C506A5AFFF0" masterDocTitle="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species" masterLastPageNumber="100" masterPageNumber="1" pageNumber="19" updateTime="1699316540125" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8BA52D114A75F9897778CF47E56779E1">Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species</mods:title>
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<mods:namePart id="D4ACBCB820AFFF481D9B5905FC58FC61">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="EDE8686BA05BA7AC966BE3FB592D9186">Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV – 1004, Rīga, Latvia. Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV – 5401, Daugavpils, Latvia.</mods:affiliation>
<mods:nameIdentifier id="FB6B2721D3FDB7EB88B8129B14C782A8" type="email">anthicus@gmail.com</mods:nameIdentifier>
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<mods:title id="253C152E6A2C9A1E1A5E13C60660BA14">European Journal of Taxonomy</mods:title>
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<mods:date id="BA68C810FE83A33D9619F36C183B6EAE">2022</mods:date>
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<mods:number id="0A76AB30ADAF7A1950B4E8959D314B2D">2022-02-24</mods:number>
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<treatment id="6D52212B21392F65FDB54A4268B4FDC1" ID-DOI="http://doi.org/10.5281/zenodo.6317922" ID-GBIF-Taxon="194038732" ID-Zenodo-Dep="6317922" LSID="urn:lsid:zoobank.org:act:F4C22B7A-9A8F-4FAE-B190-B28C99E6AB17" httpUri="http://treatment.plazi.org/id/6D52212B21392F65FDB54A4268B4FDC1" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19">
<subSubSection id="ADE1C3B621392F61FDB54A426997F9DD" box="[615,973,1554,1581]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="E544903D21392F61FDB54A426997F9DD" blockId="18.[360,1227,1554,1651]" box="[615,973,1554,1581]" pageId="18" pageNumber="19">
<heading id="BE0C275121392F61FDB54A426997F9DD" box="[615,973,1554,1581]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="22FBEBBE21392F61FDB54A426931F9DD" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[615,875,1554,1581]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FDB54A426931F9DD" bold="true" box="[615,875,1554,1581]" italics="true" pageId="18" pageNumber="19">Macrotomoderus dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FCA04A436997F9DD" box="[882,973,1555,1581]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="ADE1C3B621392F61FEBA4A656900F983" pageId="18" pageNumber="19" type="description">
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="E544903D21392F61FD0B4A086900F983" blockId="18.[360,1227,1554,1651]" box="[729,858,1624,1651]" pageId="18" pageNumber="19">
<figureCitation id="7DC08CB821392F61FD0B4A086900F983" box="[729,858,1624,1651]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="20.[189,232,1963,1989]" captionStartId-1="21.[189,232,1928,1954]" captionTargetBox-0="[246,1353,270,1902]" captionTargetBox-1="[264,1326,280,1871]" captionTargetId-0="figure-21@20.[223,1365,265,1922]" captionTargetId-1="figure-12@21.[253,1334,265,1877]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="Fig. 10. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." captionText-1="Fig. 11. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN), aedeagus. A–B. General view, different positions (A combined from two images). C. Gonopore armature in median portion of aedeagus. D. Ditto, stronger magnification. E. Basal portion of apicale. F. Apical portion. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6299509" figureDoi-1="http://doi.org/10.5281/zenodo.6299511" httpUri-0="https://zenodo.org/record/6299509/files/figure.png" httpUri-1="https://zenodo.org/record/6299511/files/figure.png" pageId="18" pageNumber="19">Figs 10–11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621392F60FF6F4ACA6850FEB7" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="E544903D21392F61FF6F4ACA6B93F944" blockId="18.[189,457,1690,1716]" box="[189,457,1690,1716]" pageId="18" pageNumber="19">
<heading id="BE0C275121392F61FF6F4ACA6B93F944" bold="true" box="[189,457,1690,1716]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="D78F4C2F21392F61FF6F4ACA6B93F944" bold="true" box="[189,457,1690,1716]" pageId="18" pageNumber="19">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21392F60FF6F4A986850FEB7" blockId="18.[189,1400,1736,2009]" lastBlockId="19.[189,1398,265,327]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
This species falls in a group of species from continental
<collectingCountry id="9DECD0AD21392F61FCB24A9869FCF912" box="[864,934,1736,1762]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
with pronotum wider than head across eyes. Among them, the males of
<taxonomicName id="22FBEBBE21392F61FD954ABC6920F8F6" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[583,890,1772,1798]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="D78F4C2F21392F61FD954ABC6883F8F6" box="[583,729,1772,1798]" italics="true" pageId="18" pageNumber="19">M. mirabilis</emphasis>
<bibRefCitation id="816AEDCC21392F61FD324ABC6920F8F6" author="Telnov D." box="[736,890,1772,1798]" pageId="18" pageNumber="19" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="22FBEBBE21392F61FC674ABC6F6DF8F6" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[949,1335,1772,1798]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="monstrificabilis">
<emphasis id="D78F4C2F21392F61FC674ABC6ECFF8F6" box="[949,1173,1772,1798]" italics="true" pageId="18" pageNumber="19">M. monstrificabilis</emphasis>
<bibRefCitation id="816AEDCC21392F61FB4F4ABC6F6DF8F6" author="Telnov D." box="[1181,1335,1772,1798]" pageId="18" pageNumber="19" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
have distinct triangular median projection at the head base (not present in
<taxonomicName id="22FBEBBE21392F61FBD44B406E04F8D9" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[1030,1118,1807,1833]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FBD44B406E04F8D9" box="[1030,1118,1807,1833]" italics="true" pageId="18" pageNumber="19">M. dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FBBA4B5F6E90F8D9" box="[1128,1226,1807,1833]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
). The median, dorsal longitudinal carina of pronotal constriction not present in
<taxonomicName id="22FBEBBE21392F61FC1D4B636E7DF8BC" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[975,1063,1842,1868]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FC1D4B636E7DF8BC" box="[975,1063,1842,1868]" italics="true" pageId="18" pageNumber="19">M. dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FBE04B636EC8F8BD" box="[1074,1170,1843,1869]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, weakly present in
<taxonomicName id="22FBEBBE21392F61FF6F4B066B76F89F" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[189,300,1877,1903]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="boops" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FF6F4B066B76F89F" box="[189,300,1877,1903]" italics="true" pageId="18" pageNumber="19">M. boops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FEE74B066BC8F880" box="[309,402,1878,1904]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
(see description above), the eyes are comparatively smaller in
<taxonomicName id="22FBEBBE21392F61FB5A4B066E85F89F" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[1160,1247,1877,1903]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FB5A4B066E85F89F" box="[1160,1247,1877,1903]" italics="true" pageId="18" pageNumber="19">M. dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FB3A4B066F1DF880" box="[1256,1351,1878,1904]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, the head base either only medially subtruncate (broadly subtruncate in
<taxonomicName id="22FBEBBE21392F61FC014B296E1BF862" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[979,1089,1912,1938]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="boops" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FC014B296E1BF862" box="[979,1089,1912,1938]" italics="true" pageId="18" pageNumber="19">M. boops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FB9A4B296EFDF863" box="[1096,1191,1913,1939]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
). In
<taxonomicName id="22FBEBBE21392F61FB0E4B296F2DF862" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[1244,1399,1912,1938]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hengduan" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FB0E4B296F2DF862" box="[1244,1399,1912,1938]" italics="true" pageId="18" pageNumber="19">M. hengduan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FF6F4BCC6B42F846" box="[189,280,1948,1974]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
(see description below) the shape of the aedeagus is different, the head base is broadly rounded (truncate medially in
<taxonomicName id="22FBEBBE21392F61FE674B906850F829" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[437,522,1983,2009]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F21392F61FE674B906850F829" box="[437,522,1983,2009]" italics="true" pageId="18" pageNumber="19">M. dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421392F61FDC24BEF6837F829" box="[528,621,1983,2009]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
), the compound eyes are comparatively smaller and not protruding from the lateral outline of the head (the eyes are larger and stronger protruding beyond head outline laterally in
<taxonomicName id="22FBEBBE21382F60FE914D7D6BC2FEB6" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[323,408,300,326]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F21382F60FE914D7D6BC2FEB6" box="[323,408,300,326]" italics="true" pageId="19" pageNumber="20">M. dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421382F60FE4D4D7D6BA7FEB7" box="[415,509,301,327]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621382F60FF6F4D3E6804FE2A" pageId="19" pageNumber="20" type="etymology">
<paragraph id="E544903D21382F60FF6F4D3E6B1EFE78" blockId="19.[189,324,366,392]" box="[189,324,366,392]" pageId="19" pageNumber="20">
<heading id="BE0C275121382F60FF6F4D3E6B1EFE78" bold="true" box="[189,324,366,392]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="D78F4C2F21382F60FF6F4D3E6B1EFE78" bold="true" box="[189,324,366,392]" pageId="19" pageNumber="20">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21382F60FF6F4DCC6804FE2A" blockId="19.[189,1399,412,474]" pageId="19" pageNumber="20">
Named after the ancient Dali Kingdom, situated in modern
<collectingRegion id="273F5EDF21382F60FCA14DCD6E66FE46" box="[883,1084,412,439]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan Province</collectingRegion>
from 937 until
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the area where this new species occurs.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621382F60FF6F4E516982FD60" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="E544903D21382F60FF6F4E516B36FDEA" blockId="19.[189,364,512,539]" box="[189,364,512,539]" pageId="19" pageNumber="20">
<heading id="BE0C275121382F60FF6F4E516B36FDEA" bold="true" box="[189,364,512,539]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="D78F4C2F21382F60FF6F4E516B36FDEA" bold="true" box="[189,364,512,539]" pageId="19" pageNumber="20">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21382F60FF384E7F6982FD60" blockId="19.[189,1399,559,656]" pageId="19" pageNumber="20">
<materialsCitation id="55939A6021382F60FF384E7F6982FD60" ID-GBIF-Occurrence="3705075331" collectingDate="2019-05-20" collectionCode="ZIN" collectorName="Belousov &amp; Davidian &amp; Kabak" country="China" elevation="2620" latitude="25.609167" location="Yunnan" longLatPrecision="21" longitude="99.92" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="D78F4C2F21382F60FF384E7F6B06FDB9" bold="true" box="[234,348,559,585]" pageId="19" pageNumber="20">
<typeStatus id="3A402E9F21382F60FF384E7F6B06FDB9" box="[234,348,559,585]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="9DECD0AD21382F60FF6F4E036B42FD9D" box="[189,280,595,621]" name="China" pageId="19" pageNumber="20">CHINA</collectingCountry>
•
<specimenCount id="F3FD5BB421382F60FEE64E036B17FD9D" box="[308,333,595,621]" pageId="19" pageNumber="20" type="male">♂</specimenCount>
; “CHINA,
<collectingRegion id="273F5EDF21382F60FE064E03686BFD9D" box="[468,561,595,621]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan</collectingRegion>
,
<locationDeviation id="F4F6891F21382F60FDEF4E036ED1FD9D" box="[573,1163,594,621]" pageId="19" pageNumber="20">
W Dali City, SW Yangbi,
<quantity id="22033DD821382F60FCA64E0269F3FD9C" box="[884,937,594,620]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="20" unit="km" value="1.0">1km</quantity>
N Laoheshang Mt
</locationDeviation>
,
<quantity id="22033DD821382F60FB4A4E026EAFFD9D" box="[1176,1269,594,621]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="19" pageNumber="20" unit="m" value="2620.0">
<elevation id="6ED6770E21382F60FB4A4E026EAFFD9D" box="[1176,1269,594,621]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="19" pageNumber="20" unit="m" value="2620.0">2620 m</elevation>
</quantity>
,
<date id="9145B6FD21382F60FAD14E036F2DFD9D" box="[1283,1399,595,621]" pageId="19" pageNumber="20" value="2019-05-20">
<collectingDate id="81014F1521382F60FAD14E036F2DFD9D" box="[1283,1399,595,621]" pageId="19" pageNumber="20" value="2019-05-20">20.v.2019</collectingDate>
</date>
<collectorName id="480EF5EB21382F60FF6F4E266B77FD60" box="[189,301,630,656]" pageId="19" pageNumber="20">Belousov</collectorName>
,
<collectorName id="480EF5EB21382F60FEEB4E266BFFFD60" box="[313,421,630,656]" pageId="19" pageNumber="20">Davidian</collectorName>
&amp;
<collectorName id="480EF5EB21382F60FE184E26684DFD60" box="[458,535,630,656]" pageId="19" pageNumber="20">Kabak</collectorName>
leg. //
<geoCoordinate id="80CFF6FA21382F60FDBA4E2568A9FD60" box="[616,755,628,656]" degrees="25" direction="north" minutes="36" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="33" value="25.609167">25°36′33″N</geoCoordinate>
<geoCoordinate id="80CFF6FA21382F60FD284E2669D2FD60" box="[762,904,628,656]" degrees="99" direction="east" minutes="55" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="12" value="99.92">99°55′12″E</geoCoordinate>
”;
<collectionCode id="83EA08F821382F60FC4D4E26698EFD60" box="[927,980,630,656]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="20" type="Museum">ZIN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621382F65FF6F4EE76BF1FEA1" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20" type="description">
<paragraph id="E544903D21382F60FF6F4EE76B14FD21" blockId="19.[189,334,695,721]" box="[189,334,695,721]" pageId="19" pageNumber="20">
<heading id="BE0C275121382F60FF6F4EE76B14FD21" bold="true" box="[189,334,695,721]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="D78F4C2F21382F60FF6F4EE76B14FD21" bold="true" box="[189,334,695,721]" pageId="19" pageNumber="20">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21382F60FF6F4EB66B80FCB6" blockId="19.[189,1399,741,839]" pageId="19" pageNumber="20">
MEASUREMENTS.
<typeStatus id="3A402E9F21382F60FEAD4EB66BB5FCF0" box="[383,495,742,768]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="22033DD821382F60FD034EB5696AFCF0" box="[721,816,741,768]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="19" pageNumber="20" unit="mm" value="4.5">4.5 mm</quantity>
; head
<quantity id="22033DD821382F60FCAD4EB66981FCF0" box="[895,987,742,768]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="19" pageNumber="20" unit="mm" value="0.9">0.9 mm</quantity>
long, across eyes
<quantity id="22033DD821382F60FB6B4EB66F79FCF0" box="[1209,1315,742,768]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="19" pageNumber="20" unit="mm" value="0.96">0.96 mm</quantity>
broad, pronotum
<quantity id="22033DD821382F60FEE94F596BFFFCD3" box="[315,421,777,804]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="19" pageNumber="20" unit="mm" value="1.35">1.35 mm</quantity>
long, maximum width
<quantity id="22033DD821382F60FD114F596968FCD3" box="[707,818,777,804]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="19" pageNumber="20" unit="mm" value="1.05">1.05 mm</quantity>
, minimum width
<quantity id="22033DD821382F60FBD84F596E30FCD3" box="[1034,1130,777,803]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="mm" value="0.4">0.4 mm</quantity>
, elytra
<quantity id="22033DD821382F60FB174F596F6BFCD3" box="[1221,1329,777,804]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="19" pageNumber="20" unit="mm" value="2.25">2.25 mm</quantity>
long,
<quantity id="22033DD821382F60FF6F4F7C6B4DFCB7" box="[189,279,812,839]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="mm" value="1.5">1.5 mm</quantity>
combined wide.
</paragraph>
<paragraph id="E544903D21382F60FF6F4F3E68DAF857" blockId="19.[187,1401,877,1959]" pageId="19" pageNumber="20">
Dorsum and venter uniformly brown. Mouthparts, antennae, palps and legs yellowish-brown. Head subtriangular with moderately large, ovoid compound eyes, which are protruding beyond head outline laterally. Tempora converging towards head base, about one third shorter of eye length. Temporal angles broadly rounded, head base broadly rounded, occiput declivous posterodorsally. Head dorsal punctures minute and inconspicuous. Head dorsal setae inconspicuous, distinctly denser posterior to eyes than on frons. Antennae long, extending towards anterior third of elytra. Antennomere three about 1.25× as long as antennomere two, antennomeres 9–10 transverse, of which penultimate antennomere strongly transverse. Terminal antennomere broadly triangular with rounded apex, twice as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum broad, anterior lobe significantly wider than posterior and slightly wider than head across eyes. Postmedian lateral constriction broad and medially deeply notched. Front margin of anterior lobe broadly rounded, truncate medially, dorsally without anterior rim. Anterior lobe large and broad, inverted triangular, almost twice as long as much narrower posterior lobe. Front margin of anterior lobe medially somewhat flattened and with a group of golden, very dense setae; some (central-positioned) setae are significantly longer than other, stronger curved and touching head base (
<figureCitation id="7DC08CB821382F60FDE9490A6891FA84" box="[571,715,1370,1396]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[246,1353,270,1902]" captionTargetId="figure-21@20.[223,1365,265,1922]" captionTargetPageId="20" captionText="Fig. 10. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299509" httpUri="https://zenodo.org/record/6299509/files/figure.png" pageId="19" pageNumber="20">Fig. 10B–C</figureCitation>
). Anterior lobe slightly convex in lateral view (
<figureCitation id="7DC08CB821382F60FB2E490A6F33FA84" box="[1276,1385,1370,1396]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[246,1353,270,1902]" captionTargetId="figure-21@20.[223,1365,265,1922]" captionTargetPageId="20" captionText="Fig. 10. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299509" httpUri="https://zenodo.org/record/6299509/files/figure.png" pageId="19" pageNumber="20">Fig. 10C</figureCitation>
). Lateral constriction continues onto disc in lateral view, shallow (
<figureCitation id="7DC08CB821382F60FC0C492D6E15FA68" box="[990,1103,1405,1432]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[246,1353,270,1902]" captionTargetId="figure-21@20.[223,1365,265,1922]" captionTargetPageId="20" captionText="Fig. 10. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299509" httpUri="https://zenodo.org/record/6299509/files/figure.png" pageId="19" pageNumber="20">Fig. 10C</figureCitation>
). Lateral pronotal fovea moderately broad at lower external margin of pronotum, widens upwards towards pronotal disc in lateral view, external margins protruding into a pair of strongly obtuse (in lateral view), moderately widely separated denticles (in lateral view). Cavity in lateral wall of pronotum between lateral denticles broad, elliptical. In dorsal view, lateral pronotal fovea broad and deep, anterior and posterior pair of denticles appear acute angulate, partially concealed by dense appressed setae from adjacent portions of fovea (
<figureCitation id="7DC08CB821382F60FEE74A016BFEF99B" box="[309,420,1617,1643]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[246,1353,270,1902]" captionTargetId="figure-21@20.[223,1365,265,1922]" captionTargetPageId="20" captionText="Fig. 10. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299509" httpUri="https://zenodo.org/record/6299509/files/figure.png" pageId="19" pageNumber="20">Fig. 10B</figureCitation>
). Pronotal punctures minute on disc; lateral constriction dorsally with irregularly sized, dense punctures (
<figureCitation id="7DC08CB821382F60FE054A246818F97E" box="[471,578,1652,1678]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[246,1353,270,1902]" captionTargetId="figure-21@20.[223,1365,265,1922]" captionTargetPageId="20" captionText="Fig. 10. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299509" httpUri="https://zenodo.org/record/6299509/files/figure.png" pageId="19" pageNumber="20">Fig. 10B</figureCitation>
). Dorsal pronotal setae rather long and dense. Scutellar shield minute, triangular. Elytra dorsally elongate elliptical, flattened in lateral view, widened laterally around midlength, lateral margins evenly broadly rounded; shoulders obsolete (apterous species). Elytral punctures inconspicuous, sparse, not becoming much smaller and sparser towards apices. Elytral setae long and sparse, suberect. Male tergite and morphological sternite VII broadly rounded at posterior margin. Aedeagus as in
<figureCitation id="7DC08CB821382F60FE304B746863F8CE" box="[482,569,1828,1854]" captionStart="Fig" captionStartId="21.[189,232,1928,1954]" captionTargetBox="[264,1326,280,1871]" captionTargetId="figure-12@21.[253,1334,265,1877]" captionTargetPageId="21" captionText="Fig. 11. Macrotomoderus dali sp. nov., holotype, ♂ (ZIN), aedeagus. A–B. General view, different positions (A combined from two images). C. Gonopore armature in median portion of aedeagus. D. Ditto, stronger magnification. E. Basal portion of apicale. F. Apical portion. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299511" httpUri="https://zenodo.org/record/6299511/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
, large and robust, with rounded apex and peculiar, double gonopore armature consists of numerous irregularly shaped constituting pieces intermixed with very delicate, foam-like constituting pieces. Large constituting pieces of armature becoming distinctly smaller in apical and basal portions of aedeagus.
</paragraph>
<caption id="B184C0B5213F2F67FF6F4BFB69B0F818" ID-DOI="http://doi.org/10.5281/zenodo.6299509" ID-Zenodo-Dep="6299509" httpUri="https://zenodo.org/record/6299509/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1963,1989]" targetBox="[246,1353,270,1902]" targetPageId="20">
<paragraph id="E544903D213F2F67FF6F4BFB69B0F818" blockId="20.[188,1399,1963,2025]" pageId="20" pageNumber="21">
<emphasis id="D78F4C2F213F2F67FF6F4BFB6B40F835" bold="true" box="[189,282,1963,1989]" pageId="20" pageNumber="21">Fig. 10.</emphasis>
<taxonomicName id="22FBEBBE213F2F67FEF24BFB6844F835" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[288,542,1963,1989]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F213F2F67FEF24BFB6844F835" box="[288,542,1963,1989]" italics="true" pageId="20" pageNumber="21">Macrotomoderus dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF154213F2F67FDF74BFC68DBF836" box="[549,641,1964,1990]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZIN).
<emphasis id="D78F4C2F213F2F67FCA24BFB69DFF835" bold="true" box="[880,901,1963,1989]" pageId="20" pageNumber="21">A</emphasis>
. Habitus, dorsal view.
<emphasis id="D78F4C2F213F2F67FB424BFB6EFEF835" bold="true" box="[1168,1188,1963,1989]" pageId="20" pageNumber="21">B</emphasis>
. Forebody, dorsal view.
<emphasis id="D78F4C2F213F2F67FED14B9E6B42F818" bold="true" box="[259,280,1998,2024]" pageId="20" pageNumber="21">C</emphasis>
. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale.
</paragraph>
</caption>
<caption id="B184C0B5213E2F66FF6F4BD86EA3F818" ID-DOI="http://doi.org/10.5281/zenodo.6299511" ID-Zenodo-Dep="6299511" httpUri="https://zenodo.org/record/6299511/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1928,1954]" targetBox="[264,1326,280,1871]" targetPageId="21">
<paragraph id="E544903D213E2F66FF6F4BD86EA3F818" blockId="21.[189,1399,1928,2025]" pageId="21" pageNumber="22">
<emphasis id="D78F4C2F213E2F66FF6F4BD86B47F852" bold="true" box="[189,285,1928,1954]" pageId="21" pageNumber="22">Fig. 11.</emphasis>
<taxonomicName id="22FBEBBE213E2F66FEFA4BD86870F852" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[296,554,1928,1954]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="dali" status="sp. nov.">
<emphasis id="D78F4C2F213E2F66FEFA4BD86870F852" box="[296,554,1928,1954]" italics="true" pageId="21" pageNumber="22">Macrotomoderus dali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF154213E2F66FDE74BD868CCF852" box="[565,662,1928,1954]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (ZIN), aedeagus.
<emphasis id="D78F4C2F213E2F66FBC44BD86E14F852" bold="true" box="[1046,1102,1928,1954]" pageId="21" pageNumber="22">A–B</emphasis>
. General view, different positions (A combined from two images).
<emphasis id="D78F4C2F213E2F66FD034BFB68BCF835" bold="true" box="[721,742,1963,1989]" pageId="21" pageNumber="22">C</emphasis>
. Gonopore armature in median portion of aedeagus.
<emphasis id="D78F4C2F213E2F66FF6F4B9E6A88F818" bold="true" box="[189,210,1998,2024]" pageId="21" pageNumber="22">D</emphasis>
. Ditto, stronger magnification.
<emphasis id="D78F4C2F213E2F66FD914B9E680DF818" bold="true" box="[579,599,1998,2024]" pageId="21" pageNumber="22">E</emphasis>
. Basal portion of apicale.
<emphasis id="D78F4C2F213E2F66FC594B9E69C7F818" bold="true" box="[907,925,1998,2024]" pageId="21" pageNumber="22">F</emphasis>
. Apical portion. Not to scale.
</paragraph>
</caption>
<paragraph id="E544903D213D2F65FF6F4D586BEAFED2" blockId="22.[189,432,264,290]" box="[189,432,264,290]" pageId="22" pageNumber="23">
<heading id="BE0C2751213D2F65FF6F4D586BEAFED2" bold="true" box="[189,432,264,290]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="D78F4C2F213D2F65FF6F4D586BEAFED2" bold="true" box="[189,432,264,290]" pageId="22" pageNumber="23">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D213D2F65FF6F4D676BF1FEA1" blockId="22.[189,427,311,337]" box="[189,427,311,337]" pageId="22" pageNumber="23">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B6213D2F65FF6F4D29687FFE31" box="[189,549,377,450]" pageId="22" pageNumber="23" type="biology_ecology">
<paragraph id="E544903D213D2F65FF6F4D296B7AFE63" blockId="22.[189,288,377,403]" box="[189,288,377,403]" pageId="22" pageNumber="23">
<heading id="BE0C2751213D2F65FF6F4D296B7AFE63" bold="true" box="[189,288,377,403]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="D78F4C2F213D2F65FF6F4D296B7AFE63" bold="true" box="[189,288,377,403]" pageId="22" pageNumber="23">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D213D2F65FF6F4DF7687FFE31" blockId="22.[189,549,423,450]" box="[189,549,423,450]" pageId="22" pageNumber="23">
Collected at
<quantity id="22033DD8213D2F65FE834DF76BF0FE32" box="[337,426,423,450]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="22" pageNumber="23" unit="m" value="2620.0">2620 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B6213D2F65FF6F4DB868B4FDC1" box="[189,750,488,562]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="E544903D213D2F65FF6F4DB86B0CFDF2" blockId="22.[189,342,488,514]" box="[189,342,488,514]" pageId="22" pageNumber="23">
<heading id="BE0C2751213D2F65FF6F4DB86B0CFDF2" bold="true" box="[189,342,488,514]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="D78F4C2F213D2F65FF6F4DB86B0CFDF2" bold="true" box="[189,342,488,514]" pageId="22" pageNumber="23">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D213D2F65FF6F4E4868B4FDC1" blockId="22.[189,750,535,562]" box="[189,750,535,562]" pageId="22" pageNumber="23">
Known only from
<collectingRegion id="273F5EDF213D2F65FE474E48683BFDC1" box="[405,609,535,562]" country="China" name="Yunnan" pageId="22" pageNumber="23">Yunnan Province</collectingRegion>
, SW
<collectingCountry id="9DECD0AD213D2F65FD724E4768B0FDC1" box="[672,746,535,561]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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