Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species Author Telnov, Dmitry F78B0D3E-E409-4054-A834-E6C06918B62F Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom & Institute of Biology, University of Latvia, O. Vācieša iela 4, LV – 1004, Rīga, Latvia. Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV – 5401, Daugavpils, Latvia. anthicus@gmail.com text European Journal of Taxonomy 2022 2022-02-24 797 1 100 http://dx.doi.org/10.5852/ejt.2022.797.1667 journal article 20233 10.5852/ejt.2022.797.1667 e4b9a92e-23ef-4d55-85c9-ccabcbd8316e 2118-9773 6299487 8DF57743-9C53-4265-BCB5-743276A3A16C Macrotomoderus dali sp. nov. urn:lsid:zoobank.org:act: F4C22B7A-9A8F-4FAE-B190-B28C99E6AB17 Figs 10–11 Differential diagnosis This species falls in a group of species from continental China with pronotum wider than head across eyes. Among them, the males of M. mirabilis Telnov, 2018 and M. monstrificabilis Telnov, 2018 have distinct triangular median projection at the head base (not present in M. dali sp. nov. ). The median, dorsal longitudinal carina of pronotal constriction not present in M. dali sp. nov. , weakly present in M. boops sp. nov. (see description above), the eyes are comparatively smaller in M. dali sp. nov. , the head base either only medially subtruncate (broadly subtruncate in M. boops sp. nov. ). In M. hengduan sp. nov. (see description below) the shape of the aedeagus is different, the head base is broadly rounded (truncate medially in M. dali sp. nov. ), the compound eyes are comparatively smaller and not protruding from the lateral outline of the head (the eyes are larger and stronger protruding beyond head outline laterally in M. dali sp. nov. ). Etymology Named after the ancient Dali Kingdom, situated in modern Yunnan Province from 937 until 1382 in the area where this new species occurs. Type material Holotype CHINA • ♂ ; “CHINA, Yunnan , W Dali City, SW Yangbi, 1km N Laoheshang Mt , 2620 m , 20.v.2019 Belousov , Davidian & Kabak leg. // 25°36′33″N 99°55′12″E ”; ZIN . Description MEASUREMENTS. Holotype , total body length 4.5 mm ; head 0.9 mm long, across eyes 0.96 mm broad, pronotum 1.35 mm long, maximum width 1.05 mm , minimum width 0.4 mm , elytra 2.25 mm long, 1.5 mm combined wide. Dorsum and venter uniformly brown. Mouthparts, antennae, palps and legs yellowish-brown. Head subtriangular with moderately large, ovoid compound eyes, which are protruding beyond head outline laterally. Tempora converging towards head base, about one third shorter of eye length. Temporal angles broadly rounded, head base broadly rounded, occiput declivous posterodorsally. Head dorsal punctures minute and inconspicuous. Head dorsal setae inconspicuous, distinctly denser posterior to eyes than on frons. Antennae long, extending towards anterior third of elytra. Antennomere three about 1.25× as long as antennomere two, antennomeres 9–10 transverse, of which penultimate antennomere strongly transverse. Terminal antennomere broadly triangular with rounded apex, twice as long as penultimate antennomere. Terminal maxillary palpomere securiform. Pronotum broad, anterior lobe significantly wider than posterior and slightly wider than head across eyes. Postmedian lateral constriction broad and medially deeply notched. Front margin of anterior lobe broadly rounded, truncate medially, dorsally without anterior rim. Anterior lobe large and broad, inverted triangular, almost twice as long as much narrower posterior lobe. Front margin of anterior lobe medially somewhat flattened and with a group of golden, very dense setae; some (central-positioned) setae are significantly longer than other, stronger curved and touching head base ( Fig. 10B–C ). Anterior lobe slightly convex in lateral view ( Fig. 10C ). Lateral constriction continues onto disc in lateral view, shallow ( Fig. 10C ). Lateral pronotal fovea moderately broad at lower external margin of pronotum, widens upwards towards pronotal disc in lateral view, external margins protruding into a pair of strongly obtuse (in lateral view), moderately widely separated denticles (in lateral view). Cavity in lateral wall of pronotum between lateral denticles broad, elliptical. In dorsal view, lateral pronotal fovea broad and deep, anterior and posterior pair of denticles appear acute angulate, partially concealed by dense appressed setae from adjacent portions of fovea ( Fig. 10B ). Pronotal punctures minute on disc; lateral constriction dorsally with irregularly sized, dense punctures ( Fig. 10B ). Dorsal pronotal setae rather long and dense. Scutellar shield minute, triangular. Elytra dorsally elongate elliptical, flattened in lateral view, widened laterally around midlength, lateral margins evenly broadly rounded; shoulders obsolete (apterous species). Elytral punctures inconspicuous, sparse, not becoming much smaller and sparser towards apices. Elytral setae long and sparse, suberect. Male tergite and morphological sternite VII broadly rounded at posterior margin. Aedeagus as in Fig. 11 , large and robust, with rounded apex and peculiar, double gonopore armature consists of numerous irregularly shaped constituting pieces intermixed with very delicate, foam-like constituting pieces. Large constituting pieces of armature becoming distinctly smaller in apical and basal portions of aedeagus. Fig. 10. Macrotomoderus dali sp. nov. , holotype, ♂ (ZIN). A . Habitus, dorsal view. B . Forebody, dorsal view. C . Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale. Fig. 11. Macrotomoderus dali sp. nov. , holotype, ♂ (ZIN), aedeagus. A–B . General view, different positions (A combined from two images). C . Gonopore armature in median portion of aedeagus. D . Ditto, stronger magnification. E . Basal portion of apicale. F . Apical portion. Not to scale. Sexual dimorphism Female is unknown. Ecology Collected at 2620 m elevation. Distribution Known only from Yunnan Province , SW China .