378 lines
41 KiB
XML
378 lines
41 KiB
XML
<document id="91FAEAEB66500F3B160DAE9599ED015F" ID-DOI="10.11646/zootaxa.3024.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5280584" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629739625542" checkinUser="felipe" docAuthor="Taylor, Peter J., Lavrenchenko, Leonid A., Carleton, Michael D., Verheyen, Erik, Bennett, Nigel C., Oosthuizen, Carel J. & Maree, Sarita" docDate="2011" docId="3C3587DEFF86457D7DCCF9910B9B93B1" docLanguage="en" docName="zt03024p066.pdf" docOrigin="Zootaxa 3024 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3024.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Otomys cheesmani Taylor & Lavrenchenko & Carleton & Verheyen & Bennett & Oosthuizen & Maree 2011, new species" docType="treatment" docVersion="5" lastPageNumber="44" masterDocId="C00CFFA6FFAC45567D5BFFE50B729671" masterDocTitle="Specific limits and emerging diversity patterns in East African populations of laminate-toothed rats, genus Otomys (Muridae: Murinae: Otomyini): Revision of the Otomys typus complex 3024" masterLastPageNumber="66" masterPageNumber="1" pageNumber="43" updateTime="1699105037626" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9373147A45E10C754C49A1088C28CD3B">Specific limits and emerging diversity patterns in East African populations of laminate-toothed rats, genus Otomys (Muridae: Murinae: Otomyini): Revision of the Otomys typus complex 3024</mods:title>
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<mods:namePart id="2865920E16BC3378D93A2D0E0F29A8A4">Taylor, Peter J.</mods:namePart>
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<mods:namePart id="A7A2F8EC5AF4E43790A2A07974414BA5">Lavrenchenko, Leonid A.</mods:namePart>
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<mods:namePart id="A0D18F0CE073E9AA12F509EB4438ACB6">Carleton, Michael D.</mods:namePart>
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<mods:namePart id="698F5A1EBD5A7A212BFA8E85C63F59D6">Verheyen, Erik</mods:namePart>
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<mods:namePart id="A936209189D7B51D113F3901BD69C17F">Bennett, Nigel C.</mods:namePart>
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<mods:namePart id="91AC13918F03721A1D7D01DF6B29DC96">Oosthuizen, Carel J.</mods:namePart>
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<mods:namePart id="4529A2096AB7E0678903907F56CBB95B">Maree, Sarita</mods:namePart>
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<mods:title id="E904F4AAA86877F89F24354659E9675F">Zootaxa</mods:title>
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<mods:part id="D4281D98F94C69B178AC340C7831033E">
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<mods:date id="D2D0693F6A81006AA15027C997BC7754">2011</mods:date>
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<mods:number id="E57A92E6CF05BB31F0999CBF71EBC475">2011-09-13</mods:number>
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<mods:number id="6B78A3C729B6E3F210D4847F3C8A84DF">3024</mods:number>
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<mods:url id="B9EEDDB83ADE8C97684DDCB00343293C">https://biotaxa.org/Zootaxa/article/view/zootaxa.3024.1.1</mods:url>
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<mods:classification id="6B3DC9A681B8BD265DF367621A4992C8">journal article</mods:classification>
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<mods:identifier id="6FA52DF9A122977D433E97C6187B02A7" type="DOI">10.11646/zootaxa.3024.1.1</mods:identifier>
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<mods:identifier id="1407106AA87801F2D03974DFF1532201" type="ISSN">11755334</mods:identifier>
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<treatment id="3C3587DEFF86457D7DCCF9910B9B93B1" ID-DOI="http://doi.org/10.5281/zenodo.5285483" ID-Zenodo-Dep="5285483" LSID="urn:lsid:plazi:treatment:3C3587DEFF86457D7DCCF9910B9B93B1" httpUri="http://treatment.plazi.org/id/3C3587DEFF86457D7DCCF9910B9B93B1" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
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<subSubSection id="FC866543FF86457C7DCCF991096390FC" box="[151,529,1651,1678]" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="B42336C8FF86457C7DCCF991096390FC" blockId="42.[151,529,1651,1746]" box="[151,529,1651,1678]" pageId="42" pageNumber="43">
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<heading id="EF6B81A4FF86457C7DCCF991096390FC" bold="true" box="[151,529,1651,1678]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
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<emphasis id="86E8EADAFF86457C7DCCF991096390FC" bold="true" box="[151,529,1651,1678]" pageId="42" pageNumber="43">
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<taxonomicName id="739C4D4BFF86457C7DCCF9910A0B90FF" ID-CoL="4B7KD" authority="Taylor & Lavrenchenko & Carleton & Verheyen & Bennett & Oosthuizen & Maree, 2011" authorityName="Taylor & Lavrenchenko & Carleton & Verheyen & Bennett & Oosthuizen & Maree" authorityYear="2011" box="[151,377,1652,1678]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="cheesmani" status="sp. nov.">
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<emphasis id="86E8EADAFF86457C7DCCF9910A0B90FF" bold="true" box="[151,377,1652,1678]" italics="true" pageId="42" pageNumber="43">Otomys cheesmani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="9DDB57A1FF86457C7CDAF996096390FC" box="[385,529,1651,1677]" pageId="42" pageNumber="43" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="FC866543FF86457C7DCCF9730AF190A3" pageId="42" pageNumber="43" type="description">
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<paragraph id="B42336C8FF86457C7DCCF9730AF190DF" blockId="42.[151,529,1651,1746]" box="[151,387,1686,1710]" pageId="42" pageNumber="43">
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<figureCitation id="2CA72A4DFF86457C7DCCF9730BAF90DF" box="[151,221,1686,1710]" captionStart="FIGURE 5" captionStartId="24.[151,249,1851,1874]" captionTargetBox="[333,1169,219,1800]" captionTargetId="figure-139@24.[312,1271,193,1835]" captionTargetPageId="24" captionText="FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary)." figureDoi="http://doi.org/10.5281/zenodo.5280596" httpUri="https://zenodo.org/record/5280596/files/figure.png" pageId="42" pageNumber="43">Figs 5</figureCitation>
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,
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<figureCitation id="2CA72A4DFF86457C7DB1F9730B8890DF" box="[234,250,1686,1710]" captionStart="FIGURE 6" captionStartId="26.[151,249,1343,1366]" captionTargetBox="[190,1380,299,1307]" captionTargetId="figure-393@26.[151,1436,279,1318]" captionTargetPageId="26" captionText="FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”)." figureDoi="http://doi.org/10.5281/zenodo.5280598" httpUri="https://zenodo.org/record/5280598/files/figure.png" pageId="42" pageNumber="43">6</figureCitation>
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,
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<figureCitation id="2CA72A4DFF86457C7C5DF9730A5790DF" box="[262,293,1686,1710]" captionStart="FIGURE 17" captionStartId="48.[151,249,1971,1994]" captionTargetBox="[196,1367,183,1936]" captionTargetId="figure-35@48.[196,1384,177,1946]" captionTargetPageId="48" captionText="FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594)." figureDoi="http://doi.org/10.5281/zenodo.5280622" httpUri="https://zenodo.org/record/5280622/files/figure.png" pageId="42" pageNumber="43">17</figureCitation>
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;
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<tableCitation id="F91E0373FF86457C7C6AF9730AF190DF" box="[305,387,1686,1710]" captionStart="TABLE 9" captionStartId="52.[151,234,1198,1221]" captionText="TABLE 9. Summary statistics for taxa recognised by this study." pageId="42" pageNumber="43">Table 9</tableCitation>
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</paragraph>
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<paragraph id="B42336C8FF86457C7DCCF95F0AF190A3" blockId="42.[151,529,1651,1746]" box="[151,387,1722,1746]" pageId="42" pageNumber="43">Cheesman’s Vlei Rat</paragraph>
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</subSubSection>
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<subSubSection id="FC866543FF86457C7DCCF8E60FB49113" pageId="42" pageNumber="43" type="materials_examined">
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<paragraph id="B42336C8FF86457C7DCCF8E60AE4914F" blockId="42.[151,1436,1794,2034]" pageId="42" pageNumber="43">
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<materialsCitation id="04F43C95FF86457C7DCCF8E60AE4914F" collectingDate="1937-10-20" collectionCode="BMNH, R" collectorName="E. Cheesman" pageId="42" pageNumber="43" specimenCount="1" typeStatus="holotype">
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<emphasis id="86E8EADAFF86457C7DCCF8E60A79916A" bold="true" box="[151,267,1795,1819]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">
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<typeStatus id="6B27886AFF86457C7DCCF8E60A74916A" box="[151,262,1795,1819]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="D28DAE0DFF86457C7C49F8E60A14916B" box="[274,358,1795,1818]" pageId="42" pageNumber="43">BMNH</collectionCode>
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1937.2.24.84; adult male, dry skin and skull, collector’s number 7400; collected by
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<collectionCode id="D28DAE0DFF86457C7842F8E60E5C916B" box="[1305,1326,1795,1818]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="42" pageNumber="43">R</collectionCode>
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.
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<collectorName id="1969531EFF86457C7869F8E60BBB914F" pageId="42" pageNumber="43">E. Cheesman</collectorName>
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,
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<date id="C0221008FF86457C7D88F8C30AE0914F" box="[211,402,1830,1854]" pageId="42" pageNumber="43" value="1937-10-20">
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<collectingDate id="D066E9E0FF86457C7D88F8C30AE0914F" box="[211,402,1830,1854]" pageId="42" pageNumber="43" value="1937-10-20">20 October 1937</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="B42336C8FF86457C7D9EF8AE0FB49113" blockId="42.[151,1436,1794,2034]" box="[197,1222,1866,1891]" pageId="42" pageNumber="43">
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<emphasis id="86E8EADAFF86457C7D9EF8AE0A119112" bold="true" box="[197,355,1866,1891]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Type locality.</emphasis>
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<materialsCitation id="04F43C95FF86457C7C31F8AF089C9113" box="[362,1006,1866,1890]" country="Ethiopia" elevation="2100" location="Dangila" pageId="42" pageNumber="43" specimenCount="1">
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<collectingCountry id="CC8B7658FF86457C7C31F8AF0ABE9113" box="[362,460,1866,1890]" name="Ethiopia" pageId="42" pageNumber="43">Ethiopia</collectingCountry>
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, vicinity of
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<location id="B1436013FF86457C7F09F8AF09DF9113" LSID="urn:lsid:plazi:treatment:3C3587DEFF86457D7DCCF9910B9B93B1:B1436013FF86457C7F09F8AF09DF9113" box="[594,685,1866,1890]" country="Ethiopia" name="Dangila" pageId="42" pageNumber="43">Dangila</location>
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,
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<locationDeviation id="A5912FEAFF86457C7FECF8AF08FF9113" box="[695,909,1866,1890]" location="Lake Tana" pageId="42" pageNumber="43">
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<quantity id="73649B2DFF86457C7FECF8AF098E9113" box="[695,764,1866,1890]" metricMagnitude="4" metricUnit="m" metricValue="6.6" pageId="42" pageNumber="43" unit="km" value="66.0">66 km</quantity>
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S Lake Tana
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</locationDeviation>
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,
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<quantity id="73649B2DFF86457C7EC3F8AF089C9113" box="[920,1006,1866,1890]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="42" pageNumber="43" unit="m" value="2100.0">
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<elevation id="3FB1D1FBFF86457C7EC3F8AF089C9113" box="[920,1006,1866,1890]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="42" pageNumber="43" unit="m" value="2100.0">2100 m</elevation>
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</quantity>
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</materialsCitation>
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;
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<geoCoordinate id="D1A8500FFF86457C7EA2F8AF0F2A9113" box="[1017,1112,1866,1891]" degrees="11" direction="north" minutes="16" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="11.266666">11°16’N</geoCoordinate>
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,
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<geoCoordinate id="D1A8500FFF86457C793FF8AF0FB09113" box="[1124,1218,1866,1891]" degrees="36" direction="east" minutes="51" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="36.85">36°51’E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="FC866543FF86457D7D9EF88B09879785" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="diagnosis">
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<paragraph id="B42336C8FF86457C7D9EF88B08309183" blockId="42.[151,1436,1794,2034]" pageId="42" pageNumber="43">
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<emphasis id="86E8EADAFF86457C7D9EF88B0A4D91F6" bold="true" box="[197,319,1902,1927]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Diagnosis.</emphasis>
|
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The largest representative of the
|
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<taxonomicName id="739C4D4BFF86457C7FE6F88B082491F7" authorityName="sensu Bohmann" authorityYear="1952" box="[701,854,1902,1926]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="typus">
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<emphasis id="86E8EADAFF86457C7FE6F88B082491F7" box="[701,854,1902,1926]" italics="true" pageId="42" pageNumber="43">
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Otomys
|
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<typeStatus id="6B27886AFF86457C7E40F88A082491F7" box="[795,854,1903,1926]" pageId="42" pageNumber="43">typus</typeStatus>
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</emphasis>
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</taxonomicName>
|
||
species complex, differing from all other members by its distinctly larger and more robust skull with short and broad nasal bones, broad, angular and thickened zygomatic arches, very well developed supraorbital and parietal ridges and relatively very narrow interorbital region (referred to “Sp. C” above; see
|
||
<figureCitation id="2CA72A4DFF86457C7CACF83F094C9183" box="[503,574,2010,2034]" captionStart="FIGURE 5" captionStartId="24.[151,249,1851,1874]" captionTargetBox="[333,1169,219,1800]" captionTargetId="figure-139@24.[312,1271,193,1835]" captionTargetPageId="24" captionText="FIGURE 5. Principal components analysis (PCA) based on nine craniometric variables indicating scatter of individuals on PC1 and PC2 (a) and PC2 and PC3 (b) from six pre-defined taxon-groups of Ethiopian Otomys in relation to existing type specimens of fortior and degeni and new holotypes designated for new species (see Taxonomic Summary). Open squares = O. typus s.s., open triangles = O. fortior; open diamonds = O. yaldeni sp. nov. (= Sp. A); closed squares = O. helleri (= Sp. B); open circles = O. cheesmani sp. nov. (= Sp.C); closed diamonds = O. simiensis sp. nov. (= Sp. D); cross = degeni. Type specimens indicated; normal font indicates existing holotypes; bold font represents newly designated holotypes (see Taxonomic Summary)." figureDoi="http://doi.org/10.5281/zenodo.5280596" httpUri="https://zenodo.org/record/5280596/files/figure.png" pageId="42" pageNumber="43">Figs 5</figureCitation>
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||
,
|
||
<figureCitation id="2CA72A4DFF86457C7F11F83F09289183" box="[586,602,2010,2034]" captionStart="FIGURE 6" captionStartId="26.[151,249,1343,1366]" captionTargetBox="[190,1380,299,1307]" captionTargetId="figure-393@26.[151,1436,279,1318]" captionTargetPageId="26" captionText="FIGURE 6. Canonical variates analysis (plot of first two canonical variates) of six defined taxon-groups of Ethiopian Otomys based on nine craniometric variables. Position of individuals form “unknown” sample from Oromiya, Jimma (asterisks) predicted a posteriori. Open squares = O. typus s.s. (Mt Guna); Closed squares = O. fortior; open circles = O. helleri (= “Sp. B”); closed circles = O. yaldeni sp. nov. (= “Sp. A”); open triangles = O. cheesmani sp. nov. (= “Sp. C”); closed triangles = O. simiensis sp. nov. (= “Sp. D”)." figureDoi="http://doi.org/10.5281/zenodo.5280598" httpUri="https://zenodo.org/record/5280598/files/figure.png" pageId="42" pageNumber="43">6</figureCitation>
|
||
,
|
||
<figureCitation id="2CA72A4DFF86457C7F3DF83F09F09183" box="[614,642,2010,2034]" captionStart="FIGURE 17" captionStartId="48.[151,249,1971,1994]" captionTargetBox="[196,1367,183,1936]" captionTargetId="figure-35@48.[196,1384,177,1946]" captionTargetPageId="48" captionText="FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594)." figureDoi="http://doi.org/10.5281/zenodo.5280622" httpUri="https://zenodo.org/record/5280622/files/figure.png" pageId="42" pageNumber="43">17</figureCitation>
|
||
and
|
||
<tableCitation id="F91E0373FF86457C7FE2F83F086A9183" box="[697,792,2010,2034]" captionStart="TABLE 7" captionStartId="29.[151,234,1317,1340]" captionText="TABLE 7. ANOVA of selected cranial variables for 12 proposed taxa of O. typus s.l. Homogeneous groups represent Tukey’s" pageId="42" pageNumber="43">Tables 7</tableCitation>
|
||
,
|
||
<tableCitation id="F91E0373FF86457C7E7FF83F08469183" box="[804,820,2010,2034]" captionStart="TABLE 9" captionStartId="52.[151,234,1198,1221]" captionText="TABLE 9. Summary statistics for taxa recognised by this study." pageId="42" pageNumber="43">9</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B42336C8FF87457D7D9EFF7D09879785" blockId="43.[151,1437,151,1472]" pageId="43" pageNumber="44">
|
||
<materialsCitation id="04F43C95FF87457D7D9EFF7D0E4196DE" box="[197,1331,151,176]" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 28.1" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="86E8EADAFF87457D7D9EFF7D0A3196DE" box="[197,323,152,176]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">
|
||
<emphasis id="86E8EADAFF87457D7D9EFF7D0A4F96C1" bold="true" box="[197,317,152,176]" pageId="43" pageNumber="44">
|
||
<typeStatus id="6B27886AFF87457D7D9EFF7D0A4F96C1" box="[197,317,152,176]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="E43A9EB3FF87457D7C0EFF7D0A9996DE" box="[341,491,151,175]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 28.1</specimenCode>
|
||
.11.117 (adult female, dry skin and skull, collector’s number 6040)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7813FF7D08EC96A5" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 28.1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7813FF7D0BBA96A5" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 28.1</specimenCode>
|
||
.11.174 (adult male; dry skin and skull, collector’s number 6078)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7EF2FF590A789686" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 28.1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7EF2FF590F4496A5" box="[937,1078,188,212]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 28.1</specimenCode>
|
||
.11.175 (subadult male, dry skin and skull)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7C4DFF050FDD9686" box="[278,1199,223,247]" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 28.1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7C4DFF050AD69686" box="[278,420,223,247]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 28.1</specimenCode>
|
||
.11.176 (subadult male, dry skin and skull, collector’s number 6366)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D79E7FF050841976D" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 28.1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D79E7FF050E3B9686" box="[1212,1353,223,247]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 28.1</specimenCode>
|
||
.11.177 (adult female; dry skin and skull, collector’s number 6040)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7E64FEE10AC6974E" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 28.1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7E64FEE108BE976D" box="[831,972,260,284]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 28.1</specimenCode>
|
||
.11.178 (adult female, dry skin and skull, collector’s number 6070)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7C9BFECD0E4E974E" box="[448,1340,295,319]" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 28.1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7C9BFECD093E974E" box="[448,588,295,319]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 28.1</specimenCode>
|
||
.11.179 (adult female, dry skin and skull, collector’s number 6165)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7813FECD08C09715" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 1937.2" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7813FECD0B979715" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 1937.2</specimenCode>
|
||
.24.82 (adult male, dry skin and skull, collector’s number 7383)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7EE5FEA9094897F6" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 1937.2" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7EE5FEA90F1B9715" box="[958,1129,332,356]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 1937.2</specimenCode>
|
||
.24.83 (adult male, dry skin and skull, collector’s number 7387)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7F11FE950BA997DD" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 1937.2" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7F11FE95098A97F6" box="[586,760,367,391]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 1937.2</specimenCode>
|
||
.24.85 (adult male, dry skin and skull, collector’s number 7404)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="04F43C95FF87457D7DB2FE710E1097DD" box="[233,1378,404,428]" collectionCode="BMNH" pageId="43" pageNumber="44" specimenCode="BMNH 1937.2" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7DB2FE710AE797DD" box="[233,405,404,428]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 1937.2</specimenCode>
|
||
.24.86 (adult female, dry skin and skull)—all 11 from the type locality, collected by
|
||
</materialsCitation>
|
||
<materialsCitation id="04F43C95FF87457D7837FE71095397BE" collectionCode="R" location="Cheesman" pageId="43" pageNumber="44" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="D28DAE0DFF87457D7837FE710EF397DA" box="[1388,1409,404,427]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="44">R</collectionCode>
|
||
.E.
|
||
<location id="B1436013FF87457D7DCCFE520A7B97BE" LSID="urn:lsid:plazi:treatment:3C3587DEFF86457D7DCCF9910B9B93B1:B1436013FF87457D7DCCFE520A7B97BE" box="[151,265,439,463]" name="Cheesman" pageId="43" pageNumber="44">Cheesman</location>
|
||
between 1926 and 1937
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="04F43C95FF87457D7F75FE5D09839785" collectingDate="1968-08-19" collectionCode="BMNH" collectorName="Great Abbai Expedition" country="Ethiopia" elevation="2500" latitude="10.333333" location="Gojam" longLatPrecision="1298" longitude="37.766666" municipality="Debra Marcos" pageId="43" pageNumber="44" specimenCode="BMNH 70.657" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="E43A9EB3FF87457D7F75FE5D09AA97BE" box="[558,728,439,463]" collectionCode="BMNH" pageId="43" pageNumber="44">BMNH 70.657</specimenCode>
|
||
(
|
||
<collectingCountry id="CC8B7658FF87457D7FB2FE52083E97BE" box="[745,844,439,463]" name="Ethiopia" pageId="43" pageNumber="44">Ethiopia</collectingCountry>
|
||
,
|
||
<location id="B1436013FF87457D7E01FE5208DB97BE" LSID="urn:lsid:plazi:treatment:3C3587DEFF86457D7DCCF9910B9B93B1:B1436013FF87457D7E01FE5208DB97BE" box="[858,937,439,463]" country="Ethiopia" latitude="10.333333" longLatPrecision="1298" longitude="37.766666" municipality="Debra Marcos" name="Gojam" pageId="43" pageNumber="44">Gojam</location>
|
||
,
|
||
<collectingMunicipality id="5447ACB2FF87457D7EECFE520F2A97BE" box="[951,1112,439,463]" pageId="43" pageNumber="44">Debra Marcos</collectingMunicipality>
|
||
,
|
||
<quantity id="73649B2DFF87457D793EFE520FCC97BE" box="[1125,1214,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="m" value="2500.0">
|
||
<elevation id="3FB1D1FBFF87457D793EFE520FCC97BE" box="[1125,1214,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="m" value="2500.0">2500 m</elevation>
|
||
</quantity>
|
||
;
|
||
<geoCoordinate id="D1A8500FFF87457D7990FE520E5E97BE" box="[1227,1324,439,464]" degrees="10" direction="north" minutes="20" orientation="latitude" pageId="43" pageNumber="44" precision="925" value="10.333333">10°20’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D1A8500FFF87457D7862FE520EE597BE" box="[1337,1431,439,464]" degrees="37" direction="east" minutes="46" orientation="longitude" pageId="43" pageNumber="44" precision="925" value="37.766666">37°46’E</geoCoordinate>
|
||
; collected by
|
||
<collectorName id="1969531EFF87457D7C7AFE39095A9785" box="[289,552,476,500]" pageId="43" pageNumber="44">Great Abbai Expedition</collectorName>
|
||
,
|
||
<date id="C0221008FF87457D7F69FE39099A9785" box="[562,744,476,500]" pageId="43" pageNumber="44" value="1968-08-19">
|
||
<collectingDate id="D066E9E0FF87457D7F69FE39099A9785" box="[562,744,476,500]" pageId="43" pageNumber="44" value="1968-08-19">19 August 1968</collectingDate>
|
||
</date>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FC866543FF87457D7D9EFE1A09A495D5" pageId="43" pageNumber="44" type="description">
|
||
<paragraph id="B42336C8FF87457D7D9EFE1A08D694D6" blockId="43.[151,1437,151,1472]" pageId="43" pageNumber="44">
|
||
<emphasis id="86E8EADAFF87457D7D9EFE1A0A259469" bold="true" box="[197,343,511,536]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Description.</emphasis>
|
||
The dorsal pelage of
|
||
<taxonomicName id="739C4D4BFF87457D7F09FDE5099E9469" authorityName="Taylor & Lavrenchenko & Carleton & Verheyen & Bennett & Oosthuizen & Maree" authorityYear="2011" box="[594,748,512,536]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="cheesmani">
|
||
<emphasis id="86E8EADAFF87457D7F09FDE509149469" box="[594,614,512,536]" italics="true" pageId="43" pageNumber="44">O</emphasis>
|
||
.
|
||
<emphasis id="86E8EADAFF87457D7F2DFDE5099E9469" box="[630,748,512,536]" italics="true" pageId="43" pageNumber="44">cheesmani</emphasis>
|
||
</taxonomicName>
|
||
is bright brown with a reddish shade. Ventral pelage is pale yellowish-grey, the individual hairs yellow tipped and grey at the base. The ears are blackish, their inner surface covered with short rufous hairs. The dorsal surface of the forefeet and hindfeet is dark grey; the claws are grey. The tail is relatively short (49.3% of HB). The caudal hairs are blackish on the upper surface and pale yellowish on the lower surface; however, the tail does not appear distinctly bicoloured.
|
||
</paragraph>
|
||
<paragraph id="B42336C8FF87457D7D9EFD5109A495D5" blockId="43.[151,1437,151,1472]" pageId="43" pageNumber="44">
|
||
The robust skull of
|
||
<taxonomicName id="739C4D4BFF87457D7CC5FD51094794BD" authorityName="Taylor & Lavrenchenko & Carleton & Verheyen & Bennett & Oosthuizen & Maree" authorityYear="2011" box="[414,565,692,716]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="cheesmani">
|
||
<emphasis id="86E8EADAFF87457D7CC5FD510AC094BD" box="[414,434,692,716]" italics="true" pageId="43" pageNumber="44">O</emphasis>
|
||
.
|
||
<emphasis id="86E8EADAFF87457D7CE4FD51094794BD" box="[447,565,692,716]" italics="true" pageId="43" pageNumber="44">cheesmani</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="2CA72A4DFF87457D7F1EFD5109E694BD" box="[581,660,692,716]" captionStart="FIGURE 17" captionStartId="48.[151,249,1971,1994]" captionTargetBox="[196,1367,183,1936]" captionTargetId="figure-35@48.[196,1384,177,1946]" captionTargetPageId="48" captionText="FIGURE 17(a–c). Photographs comparing dorsal, ventral and lateral views of the skulls of holotype of Otomys yaldeni (ZMMU 162594)." figureDoi="http://doi.org/10.5281/zenodo.5280622" httpUri="https://zenodo.org/record/5280622/files/figure.png" pageId="43" pageNumber="44">Fig. 17</figureCitation>
|
||
) is reflected in the large means recorded for most cranial dimensions, which exceed those of all other representatives of the
|
||
<taxonomicName id="739C4D4BFF87457D7E50FD3D08199481" authorityName="sensu Bohmann" authorityYear="1952" box="[779,875,728,752]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="typus">
|
||
<emphasis id="86E8EADAFF87457D7E50FD3D086D9481" box="[779,799,728,752]" italics="true" pageId="43" pageNumber="44">O</emphasis>
|
||
.
|
||
<emphasis id="86E8EADAFF87457D7E6BFD3C08199481" box="[816,875,729,752]" italics="true" pageId="43" pageNumber="44">typus</emphasis>
|
||
</taxonomicName>
|
||
complex except for interorbital width (
|
||
<tableCitation id="F91E0373FF87457D786FFD320EFC949E" box="[1332,1422,727,751]" captionStart="TABLE 9" captionStartId="52.[151,234,1198,1221]" captionText="TABLE 9. Summary statistics for taxa recognised by this study." pageId="43" pageNumber="44">Table 9</tableCitation>
|
||
). Although the observed ranges of certain external and cranial variables overlap, four cranial variables do not overlap in size: GLS (
|
||
<quantity id="73649B2DFF87457D7C6FFCFA0AAB9546" box="[308,473,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="4.575" metricValueMax="4.82" metricValueMin="4.33" pageId="43" pageNumber="44" unit="mm" value="45.75" valueMax="48.2" valueMin="43.3">43.3–48.2 mm</quantity>
|
||
versus
|
||
<quantity id="73649B2DFF87457D7F6AFCFA09AB9546" box="[561,729,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="3.88" metricValueMax="4.33" metricValueMin="3.4299999999999997" pageId="43" pageNumber="44" unit="mm" value="38.8" valueMax="43.3" valueMin="34.3">34.3–43.3 mm</quantity>
|
||
); MXTL (11.0–12.0 mm versus
|
||
<quantity id="73649B2DFF87457D7913FCFA0F909546" box="[1096,1250,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" metricValueMax="10.5" metricValueMin="8.1" pageId="43" pageNumber="44" unit="mm" value="9.3" valueMax="10.5" valueMin="8.1">8.1–10.5 mm</quantity>
|
||
); NAW (8.0–9.0 mm versus 5.8–8.0 mm); PL (
|
||
<quantity id="73649B2DFF87457D7CB8FCA109F4952A" box="[483,646,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" metricValueMax="2.62" metricValueMin="2.36" pageId="43" pageNumber="44" unit="mm" value="24.9" valueMax="26.2" valueMin="23.6">23.6–26.2 mm</quantity>
|
||
versus
|
||
<quantity id="73649B2DFF87457D7F80FCA1080C952D" box="[731,894,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="2.0149999999999997" metricValueMax="2.36" metricValueMin="1.67" pageId="43" pageNumber="44" unit="mm" value="20.15" valueMax="23.6" valueMin="16.7">16.7–23.6 mm</quantity>
|
||
) (
|
||
<tableCitation id="F91E0373FF87457D7ECFFCA10898952D" box="[916,1002,836,860]" captionStart="TABLE 9" captionStartId="52.[151,234,1198,1221]" captionText="TABLE 9. Summary statistics for taxa recognised by this study." pageId="43" pageNumber="44">Table 9</tableCitation>
|
||
). The anterior face of the lower incisor has two well defined and equally deep grooves. The upper third molar has 8 or 9 complete laminae (
|
||
<quantity id="73649B2DFF87457D785FFC820E5D950E" box="[1284,1327,871,895]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="43" pageNumber="44" unit="in" value="9.0">9 in</quantity>
|
||
the
|
||
<typeStatus id="6B27886AFF87457D783AFC820BBB95D5" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
) and the lower first molar 4 complete laminae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FC866543FF87457D7D9EFC4A0E6F95B6" box="[197,1309,943,968]" pageId="43" pageNumber="44" type="distribution">
|
||
<paragraph id="B42336C8FF87457D7D9EFC4A0E6F95B6" blockId="43.[151,1437,151,1472]" box="[197,1309,943,968]" pageId="43" pageNumber="44">
|
||
<emphasis id="86E8EADAFF87457D7D9EFC4A0A2C95B9" bold="true" box="[197,350,943,968]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Distribution.</emphasis>
|
||
Known only from two localities in northwestern
|
||
<collectingCountry id="CC8B7658FF87457D7ED3FC4A089B95B6" box="[904,1001,943,967]" name="Ethiopia" pageId="43" pageNumber="44">Ethiopia</collectingCountry>
|
||
,
|
||
<quantity id="73649B2DFF87457D7EAFFC4A0FFD95B6" box="[1012,1167,943,967]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="43" pageNumber="44" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100–2500 m</quantity>
|
||
(see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FC866543FF87457D7D9EFC310E51920D" pageId="43" pageNumber="44" type="biology_ecology">
|
||
<paragraph id="B42336C8FF87457D7D9EFC310E51920D" blockId="43.[151,1437,151,1472]" pageId="43" pageNumber="44">
|
||
<emphasis id="86E8EADAFF87457D7D9EFC310A55959D" bold="true" box="[197,295,980,1004]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Ecology.</emphasis>
|
||
The type locality currently represents a true agricultural landscape, presumably much transformed from the time when Cheesman collected the series. The
|
||
<typeStatus id="6B27886AFF87457D7E4DFC120807927E" box="[790,885,1015,1039]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="6B27886AFF87457D7EF6FC1D0F679261" box="[941,1045,1016,1040]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="739C4D4BFF87457D7966FC1D0FA49261" authorityName="Taylor & Lavrenchenko & Carleton & Verheyen & Bennett & Oosthuizen & Maree" authorityYear="2011" box="[1085,1238,1016,1040]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="cheesmani">
|
||
<emphasis id="86E8EADAFF87457D7966FC1D0F239261" box="[1085,1105,1016,1040]" italics="true" pageId="43" pageNumber="44">O</emphasis>
|
||
.
|
||
<emphasis id="86E8EADAFF87457D793BFC1D0FA49261" box="[1120,1238,1016,1040]" italics="true" pageId="43" pageNumber="44">cheesmani</emphasis>
|
||
</taxonomicName>
|
||
were supposedly collected in an intrazonal wetland habitat. The specimen from Debra Marcos (BMNH 70.657) was captured in low
|
||
<taxonomicName id="739C4D4BFF87457D7DCCFBA50B919229" box="[151,227,1088,1112]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="86E8EADAFF87457D7DCCFBA50B919229" box="[151,227,1088,1112]" italics="true" pageId="43" pageNumber="44">Acacia</emphasis>
|
||
</taxonomicName>
|
||
scrub.
|
||
<taxonomicName id="739C4D4BFF87457D7C6BFBA509709229" authorityName="Taylor & Lavrenchenko & Carleton & Verheyen & Bennett & Oosthuizen & Maree" authorityYear="2011" box="[304,514,1088,1112]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="cheesmani">
|
||
<emphasis id="86E8EADAFF87457D7C6BFBA509709229" box="[304,514,1088,1112]" italics="true" pageId="43" pageNumber="44">Otomys cheesmani</emphasis>
|
||
</taxonomicName>
|
||
occupies lower elevations (
|
||
<quantity id="73649B2DFF87457D7E6CFBA508A29229" box="[823,976,1088,1112]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="43" pageNumber="44" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100–2500 m</quantity>
|
||
) than other Ethiopian
|
||
<taxonomicName id="739C4D4BFF87457D7993FBA50E6F9229" authorityName="F. Cuvier" authorityYear="1824" box="[1224,1309,1088,1112]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="86E8EADAFF87457D7993FBA50E6F9229" box="[1224,1309,1088,1112]" italics="true" pageId="43" pageNumber="44">Otomys</emphasis>
|
||
</taxonomicName>
|
||
(excluding the rainforest dweller
|
||
<taxonomicName id="739C4D4BFF87457D7CD6FB810A8A920D" authorityName="Thomas" authorityYear="1906" box="[397,504,1124,1148]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="fortior">
|
||
<emphasis id="86E8EADAFF87457D7CD6FB810A8A920D" box="[397,504,1124,1148]" italics="true" pageId="43" pageNumber="44">O. fortior</emphasis>
|
||
</taxonomicName>
|
||
). Nevertheless, habitat requirements of this new species remain unclear.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FC866543FF87457D7D9EFB6D0B9B93B1" pageId="43" pageNumber="44" type="etymology">
|
||
<paragraph id="B42336C8FF87457D7D9EFB6D0B9B93B1" blockId="43.[151,1437,151,1472]" pageId="43" pageNumber="44">
|
||
<emphasis id="86E8EADAFF87457D7D9EFB6D0A3E92D1" bold="true" box="[197,332,1160,1184]" inLineHeading="true" pageId="43" pageNumber="44" reason="1">Etymology.</emphasis>
|
||
We selected the specific epithet to recognize Robert E. Cheesman (1878–1962), who collected the
|
||
<typeStatus id="6B27886AFF87457D7DCCFB490B8B92B5" box="[151,249,1196,1220]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
and topotypic series of this distinctive species of
|
||
<taxonomicName id="739C4D4BFF87457D7E7AFB49080492B5" authorityName="F. Cuvier" authorityYear="1824" box="[801,886,1196,1220]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="86E8EADAFF87457D7E7AFB49080492B5" box="[801,886,1196,1220]" italics="true" pageId="43" pageNumber="44">Otomys</emphasis>
|
||
</taxonomicName>
|
||
. Cheesman was a British military and diplomatic officer in the Middle East, Arabian Peninsula, and nearby Africa, and like many other professionals of this era, he developed broad interests in the geography and natural history of the lands where he worked. During his service as Consul in North-West
|
||
<collectingCountry id="CC8B7658FF87457D7CCEFAFD097A9341" box="[405,520,1304,1328]" name="Ethiopia" pageId="43" pageNumber="44">Abyssinia</collectingCountry>
|
||
, Cheesman not only explored the source of the
|
||
<collectingRegion id="7658F82AFF87457D797BFAFD0FFF9341" box="[1056,1165,1304,1328]" country="Sudan" name="An Nil al Azraq" pageId="43" pageNumber="44">Blue Nile</collectingRegion>
|
||
and surveyed its course from Lake Tana to
|
||
<collectingCountry id="CC8B7658FF87457D7C37FAD90AA99325" box="[364,475,1340,1364]" name="Sudan" pageId="43" pageNumber="44">the Sudan</collectingCountry>
|
||
border (
|
||
<emphasis id="86E8EADAFF87457D7F60FAD90F009325" box="[571,1138,1340,1364]" italics="true" pageId="43" pageNumber="44">
|
||
Lake Tana and the
|
||
<collectingRegion id="7658F82AFF87457D7E54FAD908089325" box="[783,890,1340,1364]" country="Sudan" name="An Nil al Azraq" pageId="43" pageNumber="44">Blue Nile</collectingRegion>
|
||
: An Abyssinian Quest
|
||
</emphasis>
|
||
by R. E. Cheesman, 1936; Macmillan: London, 400 pp.), but also found time to collect bird and mammal specimens for the British Museum. The species name combines the surname Cheesman and genitive singular case-ending “i” indicative of masculine gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |