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<document id="3F75481165781C315798F8C9F9A21655" ID-CLB-Dataset="8900" ID-DOI="10.11646/zootaxa.4902.1.1" ID-GBIF-Dataset="6e5d2bf3-51e7-49e7-8c50-83ee03de0772" ID-ISSN="1175-5326" ID-Zenodo-Dep="4419088" ID-ZooBank="4019A40A-8F24-4B05-8615-B0E693017C77" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609837940543" checkinUser="plazi" docAuthor="Harzhauser, Mathias" docDate="2021" docId="02388C29FFB5AE12FF14836FE10AFC3D" docLanguage="en" docName="zootaxa.4902.1.1.pdf" docOrigin="Zootaxa 4902 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Gibbula vimontiae Monterosato 1884" docType="treatment" docVersion="10" lastPageNumber="30" masterDocId="FE01F451FFA9AE0FFF838535E067FF82" masterDocTitle="The Cainozoic to present-day record of Circum-Mediterranean, NE Atlantic and North Sea Cantharidinae and Trochinae (Trochoidea, Gastropoda) - a synopsis" masterLastPageNumber="81" masterPageNumber="1" pageNumber="29" updateTime="1698920037795" updateUser="ExternalLinkService">
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<mods:title id="5EB15F45D0734DE6BB84E28156B47B56">The Cainozoic to present-day record of Circum-Mediterranean, NE Atlantic and North Sea Cantharidinae and Trochinae (Trochoidea, Gastropoda) - a synopsis</mods:title>
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<mods:namePart id="6410C82F9E4E4DD39BBFC8AD5F7FA2AD">Harzhauser, Mathias</mods:namePart>
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<mods:date id="59331AD0F542C9504B1FEC09163311E1">2021</mods:date>
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<treatment id="02388C29FFB5AE12FF14836FE10AFC3D" ID-DOI="http://doi.org/10.5281/zenodo.4438947" ID-GBIF-Taxon="176581652" ID-Zenodo-Dep="4438947" LSID="urn:lsid:plazi:treatment:02388C29FFB5AE12FF14836FE10AFC3D" httpUri="http://treatment.plazi.org/id/02388C29FFB5AE12FF14836FE10AFC3D" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C28B6EB4FFB5AE13FF14836FE253F91B" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8A2E3D3FFFB5AE13FF14836FE235F9F6" blockId="28.[151,594,1626,1726]" box="[151,594,1626,1653]" pageId="28" pageNumber="29">
<heading id="D1668A53FFB5AE13FF14836FE235F9F6" bold="true" box="[151,594,1626,1653]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<taxonomicName id="4D9146BCFFB5AE13FF14836FE235F9F6" authority="Monterosato, 1884" authorityName="Monterosato" authorityYear="1884" box="[151,594,1626,1653]" class="Gastropoda" family="Trochidae" genus="Gibbula" kingdom="Animalia" order="Archaeogastropoda" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="vimontiae">
<emphasis id="B8E5E12DFFB5AE13FF14836FE235F9F6" bold="true" box="[151,594,1626,1653]" pageId="28" pageNumber="29">
<emphasis id="B8E5E12DFFB5AE13FF14836FE109F9F7" bold="true" box="[151,366,1626,1653]" italics="true" pageId="28" pageNumber="29">Gibbula vimontiae</emphasis>
Monterosato, 1884
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8A2E3D3FFFB5AE13FF44834AE253F91B" blockId="28.[151,594,1626,1726]" box="[199,564,1663,1689]" pageId="28" pageNumber="29">Present-day: Mediterranean Sea.</paragraph>
</subSubSection>
<subSubSection id="C28B6EB4FFB5AE13FF448396E288F85F" pageId="28" pageNumber="29" type="description">
<paragraph id="8A2E3D3FFFB5AE13FF448396E1A1F93C" blockId="28.[151,594,1626,1726]" box="[199,454,1699,1726]" pageId="28" pageNumber="29">(MolluscaBase 2020h)</paragraph>
<paragraph id="8A2E3D3FFFB5AE13FF14823AE2BBF8F3" blockId="28.[151,1436,1807,1905]" pageId="28" pageNumber="29">
<emphasis id="B8E5E12DFFB5AE13FF14823AE0BFF8AB" bold="true" box="[151,216,1807,1833]" pageId="28" pageNumber="29">Note:</emphasis>
<taxonomicName id="4D9146BCFFB5AE13FF5C823AE2B8F8AB" authority="Andreoli &amp; Marsigli, 1997" authorityName="Andreoli &amp; Marsigli" authorityYear="1997" box="[223,735,1807,1833]" class="Gastropoda" family="Trochidae" genus="Gibbula" kingdom="Animalia" order="Archaeogastropoda" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="bertarelli">
<emphasis id="B8E5E12DFFB5AE13FF5C823AE1CDF8AB" box="[223,426,1807,1833]" italics="true" pageId="28" pageNumber="29">Gibbula bertarelli</emphasis>
<bibRefCitation id="EE0040CEFFB5AE13FE32823AE2B8F8AB" author="Andreoli, G. &amp; Marsigli, S." box="[433,735,1807,1833]" pageId="28" pageNumber="29" pagination="19 - 21" refId="ref36582" refString="Andreoli, G. &amp; Marsigli, S. (1997) A new species of Trochus from the Tuscan Pliocene. World Shells, 20, 19 - 21." type="journal article" year="1997">Andreoli &amp; Marsigli, 1997</bibRefCitation>
</taxonomicName>
, from the Zanclean of the Mediterraean Sea (
<bibRefCitation id="EE0040CEFFB5AE13FB55823AE53CF8AB" author="Chirli, C." box="[1238,1371,1807,1833]" pageId="28" pageNumber="29" pagination="1 - 113" refId="ref39828" refString="Chirli, C. (2004) Archaeogastropoda. Malacofauna Pliocenica Toscana, 4, 1 - 113." type="journal article" year="2004">Chirli 2004</bibRefCitation>
;
<bibRefCitation id="EE0040CEFFB5AE13FAE5823AE174F8CF" author="Forli, M. &amp; Quaggiotto, E. &amp; Germana, A." pageId="28" pageNumber="29" pagination="68 - 74" refId="ref43329" refString="Forli, M., Quaggiotto, E. &amp; Germana, A. (2014) Ridescrizione di Gibbula anodosula Sacco, 1896, specie valida del Pliocene italiano (Gastropoda: Trochidae). Bollotino Malacologico, 50, 68 - 74." type="journal article" year="2014">
Forli
<emphasis id="B8E5E12DFFB5AE13FF148206E0AEF8CF" box="[151,201,1843,1869]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
. 2014
</bibRefCitation>
), is excluded from
<taxonomicName id="4D9146BCFFB5AE13FE688206E220F8CF" authorityName="Risso" authorityYear="1826" box="[491,583,1843,1869]" class="Gastropoda" family="Trochidae" genus="Gibbula" kingdom="Animalia" order="Archaeogastropoda" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">
<emphasis id="B8E5E12DFFB5AE13FE688206E220F8CF" box="[491,583,1843,1869]" italics="true" pageId="28" pageNumber="29">Gibbula</emphasis>
</taxonomicName>
and is currently placed in the
<taxonomicName id="4D9146BCFFB5AE13FC1F8206E471F8CF" authorityName="Rafinesque" authorityYear="1815" box="[924,1046,1843,1869]" class="Gastropoda" family="Turbinidae" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Mollusca" rank="family">Turbinidae</taxonomicName>
genus
<taxonomicName id="4D9146BCFFB5AE13FBE58206E5FBF8CF" authority="Magne, 1941" authorityName="Magne" authorityYear="1941" box="[1126,1436,1843,1869]" class="Gastropoda" family="Turbinidae" genus="Heteroninella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trochida" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">
<emphasis id="B8E5E12DFFB5AE13FBE58206E566F8CF" box="[1126,1281,1843,1869]" italics="true" pageId="28" pageNumber="29">Heteroninella</emphasis>
<bibRefCitation id="EE0040CEFFB5AE13FA8B8206E5FBF8CF" author="Magne, A." box="[1288,1436,1843,1869]" pageId="28" pageNumber="29" pagination="28 - 36" refId="ref48415" refString="Magne, A. (1941) Un nouveau point fossilifere du Calcaire a Asteries. Proces-verbaux de la Societe linneenne de Bordeaux, 92, 28 - 36." type="journal article" year="1941">Magne, 1941</bibRefCitation>
</taxonomicName>
(Landau &amp; Lozouet 2002;
<bibRefCitation id="EE0040CEFFB5AE13FE418262E2B7F8F3" author="Landau, B. M. &amp; Mulder, H." box="[450,720,1879,1905]" pageId="28" pageNumber="29" pagination="25 - 57" refId="ref47230" refString="Landau, B. M. &amp; Mulder, H. (2020) Additions and corrections to the Gastropod fauna of the Pliocene of Estepona, southwestern Spain. Basteria, 84 (- 3), 25 - 57." type="journal article" year="2020">Landau &amp; Mulder 2020</bibRefCitation>
).
</paragraph>
<paragraph id="8A2E3D3FFFB5AE13FF1482ABE267F83B" blockId="28.[151,751,1950,2013]" box="[151,512,1950,1977]" pageId="28" pageNumber="29">
<emphasis id="B8E5E12DFFB5AE13FF1482ABE267F83B" bold="true" box="[151,512,1950,1977]" pageId="28" pageNumber="29">
Genus
<taxonomicName id="4D9146BCFFB5AE13FF6D82ABE1EDF83B" authority="Harzhauser, 2021" authorityName="Harzhauser" authorityYear="2021" box="[238,394,1950,1977]" class="Gastropoda" family="Trochidae" genus="Gibbuliculus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B8E5E12DFFB5AE13FF6D82ABE1EDF83B" bold="true" box="[238,394,1950,1977]" italics="true" pageId="28" pageNumber="29">Gibbuliculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3D65C56FFB5AE13FE1182AAE267F83B" box="[402,512,1951,1977]" pageId="28" pageNumber="29" rank="genus">nov. gen.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8A2E3D3FFFB5AE13FF1482F6E288F85F" blockId="28.[151,751,1950,2013]" box="[151,751,1987,2013]" pageId="28" pageNumber="29">
= “
<emphasis id="B8E5E12DFFB5AE13FF3982F6E14AF85F" box="[186,301,1987,2013]" italics="true" pageId="28" pageNumber="29">Colliculus</emphasis>
” sensu auctores non Monterosato, 1888
</paragraph>
</subSubSection>
<subSubSection id="C28B6EB4FFB4AE12FF1485A2E08CFF57" pageId="29" pageNumber="30" type="type_taxon">
<paragraph id="8A2E3D3FFFB4AE12FF1485A2E08CFF57" blockId="29.[151,1437,150,852]" pageId="29" pageNumber="30">
<materialsCitation id="3AF93762FFB4AE12FF1485A2E080FF57" ID-GBIF-Occurrence="3015412307" collectorName="Tortonian, NW" country="France" location="upper Miocene" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<emphasis id="B8E5E12DFFB4AE12FF1485A2E15FFF32" bold="true" box="[151,312,150,177]" pageId="29" pageNumber="30">
<typeStatus id="552A839DFFB4AE12FF1485A2E0B5FF33" box="[151,210,151,177]" pageId="29" pageNumber="30">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4D9146BCFFB4AE12FEC785A2E47BFF33" authority="Landau, Van Dingenen &amp; Ceulemans, 2017" authorityName="Landau, Van Dingenen &amp; Ceulemans" authorityYear="2017" box="[324,1052,151,177]" class="Gastropoda" family="Trochidae" genus="Gibbula" kingdom="Animalia" order="Archaeogastropoda" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="brebioni">
<emphasis id="B8E5E12DFFB4AE12FEC785A2E26DFF33" box="[324,522,151,177]" italics="true" pageId="29" pageNumber="30">Gibbula brebioni</emphasis>
<bibRefCitation id="EE0040CEFFB4AE12FD9485A2E47BFF33" author="Landau, B. M. &amp; van Dingenen, F. &amp; Ceulemans, L." box="[535,1052,151,177]" pageId="29" pageNumber="30" pagination="75 - 166" refId="ref47316" refString="Landau, B. M., van Dingenen, F. &amp; Ceulemans, L. (2017) The upper Miocene gastropods of northwestern France, 1. Patellogastropoda and Vetigastropoda. Cainozoic Research, 17 (2), 75 - 166." type="journal article" year="2017">Landau, Van Dingenen &amp; Ceulemans, 2017</bibRefCitation>
</taxonomicName>
;
<location id="8F4E6BE4FFB4AE12FBAE85A2E4B9FF33" LSID="urn:lsid:plazi:treatment:02388C29FFB5AE12FF14836FE10AFC3D:8F4E6BE4FFB4AE12FBAE85A2E4B9FF33" box="[1069,1246,151,177]" country="France" name="upper Miocene" pageId="29" pageNumber="30">upper Miocene</location>
,
<collectorName id="276458E9FFB4AE12FB6D85A2E5FBFF33" box="[1262,1436,151,177]" pageId="29" pageNumber="30">Tortonian, NW</collectorName>
<collectingCountry id="F2867DAFFFB4AE12FF14858EE080FF57" box="[151,231,187,213]" name="France" pageId="29" pageNumber="30">France</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C28B6EB4FFB4AE12FF4485E8E34FFE07" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8A2E3D3FFFB4AE12FF4485E8E34FFE07" blockId="29.[151,1437,150,852]" pageId="29" pageNumber="30">
<emphasis id="B8E5E12DFFB4AE12FF4485E8E125FF75" bold="true" box="[199,322,221,247]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
Small trochiform shells with conical spire. Protoconch smooth, paucispiral. Spire whorls flat to moderately convex, rarely convex; shallow to weakly incised suture. Weak to moderately prominent spiral sculpture, usually of fine, close-set spiral cords; axial sculpture weak, restricted to prosocline growth lines. Rounded basal angulation in most species; base weakly convex. Weak to faint columellar swelling present. Umbilicus moderately narrow to distinctly narrowed to chink by inner lip.
</paragraph>
</subSubSection>
<subSubSection id="C28B6EB4FFB4AE12FF4484BAE3B9FDB5" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8A2E3D3FFFB4AE12FF4484BAE398FE2B" blockId="29.[151,1437,150,852]" box="[199,1023,399,425]" pageId="29" pageNumber="30">
<emphasis id="B8E5E12DFFB4AE12FF4484BAE129FE2B" bold="true" box="[199,334,399,425]" pageId="29" pageNumber="30">Etymology.</emphasis>
A combination of
<taxonomicName id="4D9146BCFFB4AE12FD9D84BAE21DFE2B" authorityName="Risso" authorityYear="1826" box="[542,634,399,425]" class="Gastropoda" family="Trochidae" genus="Gibbula" kingdom="Animalia" order="Archaeogastropoda" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B8E5E12DFFB4AE12FD9D84BAE21DFE2B" box="[542,634,399,425]" italics="true" pageId="29" pageNumber="30">Gibbula</emphasis>
</taxonomicName>
and
<emphasis id="B8E5E12DFFB4AE12FD3384BAE344FE2B" box="[688,803,399,425]" italics="true" pageId="29" pageNumber="30">Colliculus</emphasis>
; gender masculine.
</paragraph>
<paragraph id="8A2E3D3FFFB4AE12FF448487E3B9FDB5" blockId="29.[151,1437,150,852]" pageId="29" pageNumber="30">
<emphasis id="B8E5E12DFFB4AE12FF448487E21BFE4F" bold="true" box="[199,636,434,461]" pageId="29" pageNumber="30">Stratigraphic and geographic range.</emphasis>
First species appeared in the Rupelian in the Proto-Mediterranean Sea and during the Egerian (late Oligocene) in the Central Paratethys. During the Miocene,
<taxonomicName id="4D9146BCFFB4AE12FB0684E3E571FE72" authority="Harzhauser, 2021" authorityName="Harzhauser" authorityYear="2021" box="[1157,1302,470,496]" class="Gastropoda" family="Trochidae" genus="Gibbuliculus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="29" pageNumber="29" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B8E5E12DFFB4AE12FB0684E3E571FE72" box="[1157,1302,470,496]" italics="true" pageId="29" pageNumber="30">Gibbuliculus</emphasis>
</taxonomicName>
is recorded from the NE Atlantic, the Proto-Mediterranean Sea and the entire Paratethys and reached the North Sea during the Pliocene. Last occurrences are known from the Mediterranean Pleistocene.
</paragraph>
</subSubSection>
<subSubSection id="C28B6EB4FFB4AE12FF448775E10AFC3D" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8A2E3D3FFFB4AE12FF448775E390FCD6" blockId="29.[151,1437,150,852]" pageId="29" pageNumber="30">
<emphasis id="B8E5E12DFFB4AE12FF448775E12BFDD8" bold="true" box="[199,332,576,602]" pageId="29" pageNumber="30">Discussion.</emphasis>
Several small sized fossil species with moderately high conical shells, flat to moderately convex whorls, frequently with weak basal angulation, a moderately wide to distinctly narrowed umbilicus and weak to moderately prominent spiral sculpture have been traditionally placed in
<emphasis id="B8E5E12DFFB4AE12FC3087B2E441FD23" box="[947,1062,647,673]" italics="true" pageId="29" pageNumber="30">Colliculus</emphasis>
Monterosato, 1888 (e.g.
<bibRefCitation id="EE0040CEFFB4AE12FABF87BDE0B5FD47" author="Boettger, O." pageId="29" pageNumber="30" pagination="101 - 244" refId="ref38086" refString="Boettger, O. (1907) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Sz ˆ renyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenb ʾ rgischen Vereins f ʾ r Naturwissenschaften zu Hermannstadt, 55, 101 - 244. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1907">Boettger 1907</bibRefCitation>
;
<bibRefCitation id="EE0040CEFFB4AE12FF5E879EE198FD47" author="Cossmann, M. &amp; Peyrot, A." box="[221,511,683,709]" pageId="29" pageNumber="30" pagination="5 - 180" refId="ref40873" refString="Cossmann, M. &amp; Peyrot, A. (1917) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 70, 5 - 180. [https: // www. biodiversitylibrary. org / item / 103369 # page / 11 / mode / 1 up]" type="journal article" year="1917">Cossmann &amp; Peyrot 1917</bibRefCitation>
;
<bibRefCitation id="EE0040CEFFB4AE12FD88879EE2E0FD47" author="Iljina, L. B." box="[523,647,683,709]" pageId="29" pageNumber="30" pagination="1 - 149" refId="ref45768" refString="Iljina, L. B. (1993) Handbook for identification of marine Middle Miocene gastropods of Southwestern Eurasia. Trudy Paleontologicheskogo instituta, 255, 1 - 149." type="journal article" year="1993">Iljina 1993</bibRefCitation>
;
<bibRefCitation id="EE0040CEFFB4AE12FD11879EE3CDFD47" author="Marquet, R. &amp; Landau, B. M." box="[658,938,683,709]" pageId="29" pageNumber="30" pagination="13 - 49" refId="ref48755" refString="Marquet, R. &amp; Landau, B. M. (2006) The gastropod fauna of the Luchtbal Sand Member (Lillo Formation, Zanclean, Early Pliocene) of the Antwerp region (Belgium). Cainozoic Research, 5 (1 - 2), 13 - 49." type="journal article" year="2006">Marquet &amp; Landau 2006</bibRefCitation>
; Ceulemans
<emphasis id="B8E5E12DFFB4AE12FBB4879EE40FFD47" box="[1079,1128,683,709]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. 2016;
<bibRefCitation id="EE0040CEFFB4AE12FB3F879EE5F6FD47" author="Landau, B. M. &amp; van Dingenen, F. &amp; Ceulemans, L." box="[1212,1425,683,709]" pageId="29" pageNumber="30" pagination="75 - 166" refId="ref47316" refString="Landau, B. M., van Dingenen, F. &amp; Ceulemans, L. (2017) The upper Miocene gastropods of northwestern France, 1. Patellogastropoda and Vetigastropoda. Cainozoic Research, 17 (2), 75 - 166." type="journal article" year="2017">
Landau
<emphasis id="B8E5E12DFFB4AE12FA95879EE520FD47" box="[1302,1351,683,709]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. 2017
</bibRefCitation>
). In the molecular analysis of
<bibRefCitation id="EE0040CEFFB4AE12FE5D87FAE2DFFD6A" author="Barco, A. &amp; Evans, J. &amp; Schembri, P. J. &amp; Taviani, M. &amp; Oliverio, M." box="[478,696,718,745]" pageId="29" pageNumber="30" pagination="785 - 793" refId="ref37548" refString="Barco, A., Evans, J., Schembri, P. J., Taviani, M. &amp; Oliverio, M. (2013) Testing the applicability of DNA barcoding for Mediterranean species of top-shells (Gastropoda, Trochidae, Gibbula s. l.). Marine Biology Research, 9 (8), 785 - 793. https: // doi. org / 10.1080 / 17451000.2013.765585" type="journal article" year="2013">
Barco
<emphasis id="B8E5E12DFFB4AE12FDAA87FAE23BFD6A" box="[553,604,718,744]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. (2013)
</bibRefCitation>
, however,
<taxonomicName id="4D9146BCFFB4AE12FCB687FBE4BCFD6A" authority="Payraudeau, 1826" authorityName="Payraudeau" authorityYear="1826" box="[821,1243,718,744]" class="Gastropoda" family="Trochidae" genus="Trochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="adansonii">
<emphasis id="B8E5E12DFFB4AE12FCB687FBE462FD6A" box="[821,1029,718,744]" italics="true" pageId="29" pageNumber="30">Trochus adansonii</emphasis>
Payraudeau, 1826
</taxonomicName>
, the
<typeStatus id="552A839DFFB4AE12FA9187FAE525FD6B" box="[1298,1346,719,745]" pageId="29" pageNumber="30">type</typeStatus>
species of
<emphasis id="B8E5E12DFFB4AE12FF3587C7E14EFC8E" box="[182,297,754,780]" italics="true" pageId="29" pageNumber="30">Colliculus</emphasis>
, nests within
<taxonomicName id="4D9146BCFFB4AE12FE4587C7E233FC8E" box="[454,596,754,780]" class="Gastropoda" family="Trochidae" genus="Steromphala" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B8E5E12DFFB4AE12FE4587C7E233FC8E" box="[454,596,754,780]" italics="true" pageId="29" pageNumber="30">Steromphala</emphasis>
</taxonomicName>
species and
<emphasis id="B8E5E12DFFB4AE12FD6087C7E331FC8E" box="[739,854,754,780]" italics="true" pageId="29" pageNumber="30">Colliculus</emphasis>
is treated as synonym of
<taxonomicName id="4D9146BCFFB4AE12FBF487C7E562FC8E" box="[1143,1285,754,780]" class="Gastropoda" family="Trochidae" genus="Steromphala" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B8E5E12DFFB4AE12FBF487C7E562FC8E" box="[1143,1285,754,780]" italics="true" pageId="29" pageNumber="30">Steromphala</emphasis>
</taxonomicName>
(see Uribe
<emphasis id="B8E5E12DFFB4AE12FA0B87C7E0CAFCAD" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. 2017a). In terms of shell shape, these species are far from
<taxonomicName id="4D9146BCFFB4AE12FCD28620E3B8FCAD" box="[849,991,789,815]" class="Gastropoda" family="Trochidae" genus="Steromphala" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B8E5E12DFFB4AE12FCD28620E3B8FCAD" box="[849,991,789,815]" italics="true" pageId="29" pageNumber="30">Steromphala</emphasis>
</taxonomicName>
species as defined by
<bibRefCitation id="EE0040CEFFB4AE12FB5E8620E082FCD1" author="Affenzeller, S. &amp; Haar, N. &amp; Steiner, G." pageId="29" pageNumber="30" pagination="789 - 812" refId="ref36294" refString="Affenzeller, S., Haar, N. &amp; Steiner, G. (2017) Revision of the genus complex Gibbula: an integrative approach to delineating the Eastern Mediterranean genera Gibbula Risso, 1826, Steromphala Gray, 1847, and Phorcus Risso, 1826 using DNA-barcoding and geometric morphometrics (Vetigastropoda, Trochoidea). Organisms Diversity &amp; Evolution, 17 (4), 789 - 812. https: // doi. org / 10.1007 / s 13127 - 017 - 0343 - 5" type="journal article" year="2017">
Affenzeller
<emphasis id="B8E5E12DFFB4AE12FAE08623E5F2FCAD" box="[1379,1429,789,815]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. (2017)
</bibRefCitation>
. Similarly, the high trochiform shell excludes a placement in
<taxonomicName id="4D9146BCFFB4AE12FC17860CE397FCD1" authorityName="Risso" authorityYear="1826" box="[916,1008,825,851]" class="Gastropoda" family="Trochidae" genus="Gibbula" kingdom="Animalia" order="Archaeogastropoda" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B8E5E12DFFB4AE12FC17860CE397FCD1" box="[916,1008,825,851]" italics="true" pageId="29" pageNumber="30">Gibbula</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8A2E3D3FFFB4AE12FF148690E10AFC3D" blockId="29.[151,365,933,959]" box="[151,365,933,959]" pageId="29" pageNumber="30">
<emphasis id="B8E5E12DFFB4AE12FF148690E10AFC3D" bold="true" box="[151,365,933,959]" pageId="29" pageNumber="30">Included species:</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>