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<document id="12A03869BCCE748593E10B93050DC516" ID-DOI="10.11646/zootaxa.5483.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13209307" ID-ZooBank="AF0272DB-5588-411D-9EAE-DED4785BF170" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1722786113536" checkinUser="plazi" docAuthor="WOOD, T. J." docDate="2024" docId="612B87FCAC77434B0B83FA948AAD05E4" docLanguage="en" docName="zootaxa.5483.1.1.pdf" docOrigin="Zootaxa 5483 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5483.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Andrena (Chrysandrena) fulvago" docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="9D12FF84AC59437B0B14FFCB8E650641" masterDocTitle="Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)" masterLastPageNumber="150" masterPageNumber="1" pageNumber="47" updateTime="1723562696088" updateUser="guilherme">
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<mods:title id="766A02B2ED07EBF350EAE62EA8A3946C">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<treatment id="612B87FCAC77434B0B83FA948AAD05E4" LSID="urn:lsid:plazi:treatment:612B87FCAC77434B0B83FA948AAD05E4" httpUri="http://treatment.plazi.org/id/612B87FCAC77434B0B83FA948AAD05E4" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="A1986561AC7743550B83FA948CBA0338" box="[151,735,1375,1402]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="E93D36EAAC7743550B83FA948CBA0338" blockId="46.[151,735,1375,1402]" box="[151,735,1375,1402]" pageId="46" pageNumber="47">
<heading id="B2758186AC7743550B83FA948CBA0338" bold="true" box="[151,735,1375,1402]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
<taxonomicName id="2E824D69AC7743550B83FA948CBA0338" ID-CoL="DRPZ" authority="(Christ, 1791)" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[151,735,1375,1402]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fulvago" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550B83FA948CBA0338" bold="true" box="[151,735,1375,1402]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550B83FA948F64033B" bold="true" box="[151,257,1375,1402]" italics="true" pageId="46" pageNumber="47">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7743550A06FA948FDA033B" bold="true" box="[274,447,1375,1402]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7743550AC4FA948C4E033B" bold="true" box="[464,555,1375,1402]" italics="true" pageId="46" pageNumber="47">fulvago</emphasis>
(
<bibRefCitation id="8D134B1BAC774355092FFA948CB20338" author="Christ, J. L." box="[571,727,1375,1401]" pageId="46" pageNumber="47" refId="ref88151" refString="Christ, J. L. (1791) Naturgeschichte, Klassification und Nomenclatur der Insekten vom Bienen, Wespen und Ameisengeschlecht. Herrmann, Frankfurt, Main, 535 pp. https: // doi. org / 10.5962 / bhl. title. 87724" type="book" year="1791">Christ, 1791</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="A1986561AC7743550B83FA6E8A0F03BB" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="E93D36EAAC7743550B83FA6E8CC003FD" blockId="46.[151,1130,1444,1530]" box="[151,677,1444,1469]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="C99211C4AC7743550B83FA6E8CC003FD" box="[151,677,1444,1469]" pageId="46" pageNumber="47">
<treatmentCitation id="682310FBAC7743550B83FA6E8FAB03FD" author="Christ, J. L." box="[151,462,1444,1468]" page="189" pageId="46" pageNumber="47" year="1791">
<taxonomicName id="2E824D69AC7743550B83FA6E8FAB03FD" authority="Christ, 1791: 189" authorityName="Christ" authorityPageNumber="189" authorityYear="1791" box="[151,462,1444,1468]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7743550B83FA6E8F7203FD" box="[151,279,1444,1468]" italics="true" pageId="46" pageNumber="47">Apis fulvago</emphasis>
<materialsCitation id="59EA3CB7AC7743550A09FA6F8FAB03FD" box="[285,462,1444,1468]" country="Germany" location="Germany" pageId="46" pageNumber="47" specimenCount="1">
<bibRefCitation id="8D134B1BAC7743550A09FA6F8FAB03FD" author="Christ, J. L." box="[285,462,1444,1468]" pageId="46" pageNumber="47" refId="ref88151" refString="Christ, J. L. (1791) Naturgeschichte, Klassification und Nomenclatur der Insekten vom Bienen, Wespen und Ameisengeschlecht. Herrmann, Frankfurt, Main, 535 pp. https: // doi. org / 10.5962 / bhl. title. 87724" type="book" year="1791">Christ, 1791: 189</bibRefCitation>
</materialsCitation>
</taxonomicName>
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[
<collectingCountry id="9195767AAC7743550ACEFA6F8C5903FD" box="[474,572,1444,1468]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
,
<typeStatus id="36398848AC7743550952FA6E8C1703FC" box="[582,626,1445,1469]" pageId="46" pageNumber="47">type</typeStatus>
lost]
</treatmentCitationGroup>
</paragraph>
<paragraph id="E93D36EAAC7743550B83FA088DFB039A" blockId="46.[151,1130,1444,1530]" box="[151,926,1475,1499]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="C99211C4AC7743550B83FA088DFB039A" box="[151,926,1475,1499]" pageId="46" pageNumber="47">
<treatmentCitation id="682310FBAC7743550B83FA088C6A039A" author="Smith, F." box="[151,527,1475,1499]" page="59" pageId="46" pageNumber="47" year="1849">
<taxonomicName id="2E824D69AC7743550B83FA088C6A039A" authority="Smith, 1849: 59" authorityName="Smith" authorityPageNumber="59" authorityYear="1849" box="[151,527,1475,1499]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="DBF6EAF8AC7743550B83FA088F06039A" box="[151,355,1475,1499]" italics="true" pageId="46" pageNumber="47">Andrena constrictus</emphasis>
<materialsCitation id="59EA3CB7AC7743550A7DFA088C6A039A" box="[361,527,1475,1499]" collectionCode="OUMNH" country="United Kingdom" location="Scotland" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC7743550A7DFA088C6A039A" author="Smith, F." box="[361,527,1475,1499]" pageId="46" pageNumber="47" refId="ref91052" refString="Smith, F. (1849) Descriptions of New Species of British Bees. Zoologist, 7, LVII - LX." type="journal volume" year="1849">Smith, 1849: 59</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC774355090DFA088C49039B" box="[537,556,1475,1498]" count="1" pageId="46" pageNumber="47" type="male">♂</specimenCount>
[
<collectingCountry id="9195767AAC774355092CFA088CF3039A" box="[568,662,1475,1499]" name="United Kingdom" pageId="46" pageNumber="47">Scotland</collectingCountry>
, OUMNH, not examined]
</treatmentCitationGroup>
</paragraph>
<paragraph id="E93D36EAAC7743550B83FA2A8A0F03BB" blockId="46.[151,1130,1444,1530]" box="[151,1130,1505,1530]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="C99211C4AC7743550B83FA2A8A0F03BB" box="[151,1130,1505,1530]" pageId="46" pageNumber="47">
<treatmentCitation id="682310FBAC7743550B83FA2A8C4D03B8" author="Morawitz, F." box="[151,552,1505,1529]" page="68" pageId="46" pageNumber="47" year="1886">
<taxonomicName id="2E824D69AC7743550B83FA2A8C4D03B8" ID-CoL="DRCJ" authority="Morawitz, 1886: 68" authorityName="Morawitz" authorityPageNumber="68" authorityYear="1886" box="[151,552,1505,1529]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550B83FA2A8F3D03B8" box="[151,344,1505,1529]" italics="true" pageId="46" pageNumber="47">Andrena colonialis</emphasis>
<materialsCitation id="59EA3CB7AC7743550A4AFA2A8C4D03B8" box="[350,552,1505,1529]" collectionCode="ZISP" country="Azerbaijan" location="Azerbaijan" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC7743550A4AFA2A8C4D03B8" author="Morawitz, F." box="[350,552,1505,1529]" pageId="46" pageNumber="47" pagination="57 - 81" refId="ref89436" refString="Morawitz, F. (1886) Neue transcaucasische Apidae. Horae Societatis Entomologicae Rossicae, 20, 57 - 81." type="journal article" year="1886">Morawitz, 1886: 68</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC7743550927FA298C2603B8" box="[563,579,1506,1529]" count="1" pageId="46" pageNumber="47" type="female">♀</specimenCount>
<specimenCount id="FF84FD63AC7743550957FA298C3C03B8" box="[579,601,1506,1529]" count="1" pageId="46" pageNumber="47" type="male">♂</specimenCount>
[
<collectingCountry id="9195767AAC7743550971FA2A8CBF03B8" box="[613,730,1505,1529]" name="Azerbaijan" pageId="46" pageNumber="47">Azerbaijan</collectingCountry>
, ZISP, photograph examined]
<emphasis id="DBF6EAF8AC7743550F19FA298A0F03BB" bold="true" box="[1037,1130,1506,1530]" pageId="46" pageNumber="47">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC77434B0B83F9E38B17048C" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="discussion">
<paragraph id="E93D36EAAC7743550B83F9E38C6E015B" blockId="46.[151,1437,1576,2035]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550B83F9E38F6E0003" bold="true" box="[151,267,1576,1602]" pageId="46" pageNumber="47">Remarks.</emphasis>
<taxonomicName id="2E824D69AC7743550A00F9E38FB50003" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[276,464,1576,1602]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7743550A00F9E38FB50003" box="[276,464,1576,1602]" italics="true" pageId="46" pageNumber="47">Andrena fulvago</emphasis>
</taxonomicName>
is one of the oldest described
<taxonomicName id="2E824D69AC7743550825F9E38D8B0003" box="[817,1006,1576,1602]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550825F9E38DF50003" box="[817,912,1576,1602]" italics="true" pageId="46" pageNumber="47">Andrena</emphasis>
species
</taxonomicName>
, and the concept has remained robust for more than two centuries despite the lack of
<typeStatus id="36398848AC77435509A5F9878C840027" box="[689,737,1612,1638]" pageId="46" pageNumber="47">type</typeStatus>
material. It is the
<typeStatus id="36398848AC77435508A6F9878D870027" box="[946,994,1612,1638]" pageId="46" pageNumber="47">type</typeStatus>
species of the subgenus
<taxonomicName id="2E824D69AC7743550FE9F9878F5C00CB" authority="Hedicke, 1933" authorityName="Hedicke" authorityYear="1933" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550FE9F9878BF90027" box="[1277,1436,1612,1638]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
Hedicke, 1933
</taxonomicName>
that is found across the Palaearctic and which was confirmed as distinct from the Palaearctic subgenus
<taxonomicName id="2E824D69AC7743550B83F95F8FA500EF" authority="Hedicke, 1933" authorityName="Hedicke" authorityYear="1933" box="[151,448,1684,1710]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC7743550B83F95F8F7700EF" box="[151,274,1684,1710]" italics="true" pageId="46" pageNumber="47">Euandrena</emphasis>
Hedicke, 1933
</taxonomicName>
by
<bibRefCitation id="8D134B1BAC7743550AF9F95F8C8300EF" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[493,742,1684,1710]" pageId="46" pageNumber="47" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty
<emphasis id="DBF6EAF8AC774355095CF95E8C1F00EF" box="[584,634,1684,1710]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. (2022b)
</bibRefCitation>
. The two subgenera are very similar, but can be most clearly separated in the female sex by the presence of plumose scopal hairs in female
<taxonomicName id="2E824D69AC77435508E1F9738AF10093" authorityName="Hedicke" authorityYear="1933" box="[1013,1172,1720,1746]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC77435508E1F9738AF10093" box="[1013,1172,1720,1746]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
as an adaptation for the collection of
<taxonomicName id="2E824D69AC7743550A38F9178FAC00B7" box="[300,457,1756,1782]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="subFamily" subFamily="Cichorioideae">Cichorioideae</taxonomicName>
(
<taxonomicName id="2E824D69AC7743550ACCF9178C3200B7" box="[472,599,1756,1782]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) pollen, all known species of
<taxonomicName id="2E824D69AC77435508B3F9178A2300B7" authorityName="Hedicke" authorityYear="1933" box="[935,1094,1756,1782]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC77435508B3F9178A2300B7" box="[935,1094,1756,1782]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
specialising on this subfamily (
<bibRefCitation id="8D134B1BAC7743550B8AF8CB8F20015B" author="Westrich, P." box="[158,325,1792,1818]" pageId="46" pageNumber="47" refId="ref91897" refString="Westrich, P. (1989) Die wildbienen Baden-Wurttembergs. Eugen Ulmer, Stuttgart, 972 pp." type="book" year="1989">Westrich 1989</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7743550A45F8CB8F86015B" author="Wood, T. J." box="[337,483,1792,1818]" pageId="46" pageNumber="47" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
; c).
</paragraph>
<paragraph id="E93D36EAAC77434B0BD3F8EF8F7F06B8" blockId="46.[151,1437,1576,2035]" lastBlockId="48.[151,1437,151,933]" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
The majority of the important
<taxonomicName id="2E824D69AC7743550902F8EF8CA9017F" box="[534,716,1828,1854]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550902F8EF8C10017F" box="[534,629,1828,1854]" italics="true" pageId="46" pageNumber="47">Andrena</emphasis>
species
</taxonomicName>
described by Ferdinand Morawitz have recently been illustrated, including the designation of
<typeStatus id="36398848AC7743550AC9F8838C2A0123" box="[477,591,1864,1890]" pageId="46" pageNumber="47" type="lectotype">lectotypes</typeStatus>
where appropriate (
<bibRefCitation id="8D134B1BAC7743550827F8838A530123" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[819,1078,1864,1890]" pageId="46" pageNumber="47" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC77435508AEF8828D890123" box="[954,1004,1864,1890]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. 2021
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7743550F57F8838AEB0123" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." box="[1091,1166,1864,1890]" pageId="46" pageNumber="47" pagination="105 - 176" refId="ref87433" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2022 b) The bees of the genus Andrena Fabricius, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. Zookeys, 1120, 105 - 176. https: // doi. org / 10.3897 / zookeys. 1120.90206" type="journal article" year="2022">2022b</bibRefCitation>
). This includes the type material for
<taxonomicName id="2E824D69AC7743550A3DF8A78FDF01C7" authorityName="Morawitz" authorityYear="1886" box="[297,442,1900,1926]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550A3DF8A78FDF01C7" box="[297,442,1900,1926]" italics="true" pageId="46" pageNumber="47">A. colonialis</emphasis>
</taxonomicName>
which was described from the Caucasus (
<bibRefCitation id="8D134B1BAC77435508BEF8A78AD001C7" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[938,1205,1900,1926]" pageId="46" pageNumber="47" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC7743550F27F8A68A0D01C7" box="[1075,1128,1900,1926]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. 2021
</bibRefCitation>
). The identity of
<taxonomicName id="2E824D69AC7743550E90F8A78F6001EB" authorityName="Morawitz" authorityYear="1886" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550E90F8A78F6001EB" italics="true" pageId="46" pageNumber="47">A. colonialis</emphasis>
</taxonomicName>
has been obscure due to its differential treatment in the literature.
<bibRefCitation id="8D134B1BAC7743550F12F85B8B1401EB" author="Gusenleitner, F. &amp; Schwarz, M." box="[1030,1393,1936,1962]" pageId="46" pageNumber="47" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
list
<taxonomicName id="2E824D69AC7743550B83F87F8F40018F" authorityName="Morawitz" authorityYear="1886" box="[151,293,1972,1998]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550B83F87F8F40018F" box="[151,293,1972,1998]" italics="true" pageId="46" pageNumber="47">A. colonialis</emphasis>
</taxonomicName>
as a
<taxonomicName id="2E824D69AC7743550A76F87F8C64018F" authorityName="Hedicke" authorityYear="1933" box="[354,513,1972,1998]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550A76F87F8C64018F" box="[354,513,1972,1998]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
, but
<bibRefCitation id="8D134B1BAC7743550928F87F8D35018F" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[572,848,1972,1998]" pageId="46" pageNumber="47" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC77435509D0F87E8C93018F" box="[708,758,1972,1998]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. (2008)
</bibRefCitation>
list it as a
<taxonomicName id="2E824D69AC77435508DAF87F8A2C018F" authorityName="Hedicke" authorityYear="1933" box="[974,1097,1972,1998]" pageId="46" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC77435508DAF87F8A2C018F" box="[974,1097,1972,1998]" italics="true" pageId="46" pageNumber="47">Euandrena</emphasis>
</taxonomicName>
, and do not include it in their revision of
<taxonomicName id="2E824D69AC7743550A0CF8138FDC01B3" authorityName="Hedicke" authorityYear="1933" box="[280,441,2008,2034]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550A0CF8138FDC01B3" box="[280,441,2008,2034]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D134B1BAC7743550AD3F8138CAB01B2" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Cierzniak, T." box="[455,718,2008,2035]" pageId="46" pageNumber="47" refId="ref89759" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Cierzniak, T. (2005) Andreninae of the Central and Eastern Palaearctic. Part 1. Polish Entomological Monographs II. Polish Entomological Society, Poznan, 235 pp." type="book" year="2005">
Osytshnjuk
<emphasis id="DBF6EAF8AC7743550945F8128CE601B3" box="[593,643,2008,2034]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. 2005
</bibRefCitation>
). This is surprising, because the female
<typeStatus id="36398848AC7743550F85F8138A9C01B3" box="[1169,1273,2008,2034]" pageId="46" pageNumber="47" type="lectotype">lectotype</typeStatus>
designated by
<bibRefCitation id="8D134B1BAC7643540B89F9358FCC0159" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[157,425,1790,1816]" pageId="47" pageNumber="48" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC7643540A36F9348F370159" box="[290,338,1790,1816]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2008)
</bibRefCitation>
and illustrated by
<bibRefCitation id="8D134B1BAC7643540966F9358DC10158" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[626,932,1790,1817]" pageId="47" pageNumber="48" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC76435409E0F9348D460159" box="[756,803,1790,1816]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2021: 15)
</bibRefCitation>
is clearly a
<taxonomicName id="2E824D69AC7643540F31F9358AA10159" authorityName="Hedicke" authorityYear="1933" box="[1061,1220,1790,1816]" pageId="47" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7643540F31F9358AA10159" box="[1061,1220,1790,1816]" italics="true" pageId="47" pageNumber="48">Chrysandrena</emphasis>
</taxonomicName>
, as it has obviously plumose tibial scopal hairs. Structurally, there is no difference between
<taxonomicName id="2E824D69AC76435408C3F8E98A02017D" authorityName="Morawitz" authorityYear="1886" box="[983,1127,1826,1852]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC76435408C3F8E98A02017D" box="[983,1127,1826,1852]" italics="true" pageId="47" pageNumber="48">A. colonialis</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC7643540FB0F8E98B7F017D" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[1188,1306,1826,1852]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7643540FB0F8E98B7F017D" box="[1188,1306,1826,1852]" italics="true" pageId="47" pageNumber="48">A. fulvago</emphasis>
</taxonomicName>
; both have narrow facial foveae that narrow further ventrally, plumose tibial scopae, hind tibiae that are lightened orange, pale pubescence over the remaining parts of the body, a light terminal fringe, and the clypeus, scutum, and terga display a polished integument with distinct isolated punctures. The only major difference is that the colouration of the tergal margins is lighter and more extensive in
<taxonomicName id="2E824D69AC7643540976F8798C95018D" authorityName="Morawitz" authorityYear="1886" box="[610,752,1970,1996]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7643540976F8798C95018D" box="[610,752,1970,1996]" italics="true" pageId="47" pageNumber="48">A. colonialis</emphasis>
</taxonomicName>
. However, this is variable within
<taxonomicName id="2E824D69AC7643540F79F8798A85018D" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[1133,1248,1970,1996]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7643540F79F8798A85018D" box="[1133,1248,1970,1996]" italics="true" pageId="47" pageNumber="48">A. fulvago</emphasis>
</taxonomicName>
across its range, and is considered to represent only variation. Furthermore,
<bibRefCitation id="8D134B1BAC7643540838F81D8A5E01B1" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[812,1083,2006,2032]" pageId="47" pageNumber="48" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC76435408A6F81C8D8601B1" box="[946,995,2006,2032]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2008)
</bibRefCitation>
report that
<taxonomicName id="2E824D69AC7643540FA8F81D8B2C01B1" authorityName="Morawitz" authorityYear="1886" box="[1212,1353,2006,2032]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7643540FA8F81D8B2C01B1" box="[1212,1353,2006,2032]" italics="true" pageId="47" pageNumber="48">A. colonialis</emphasis>
</taxonomicName>
flies in July and August and is a specialist of
<taxonomicName id="2E824D69AC69434B092AFF5C8CD906F0" box="[574,700,151,177]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
, foraging predominantly on
<taxonomicName id="2E824D69AC69434B08EFFF5C8AAE06F0" box="[1019,1227,151,177]" class="Magnoliopsida" family="Asteraceae" genus="Cichorium" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="intybus">
<emphasis id="DBF6EAF8AC69434B08EFFF5C8AAE06F0" box="[1019,1227,151,177]" italics="true" pageId="48" pageNumber="49">Cichorium intybus</emphasis>
</taxonomicName>
. This is consistent with the flight period and dietary niche of
<taxonomicName id="2E824D69AC69434B0997FF708C9C0694" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[643,761,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC69434B0997FF708C9C0694" box="[643,761,187,213]" italics="true" pageId="48" pageNumber="49">A. fulvago</emphasis>
</taxonomicName>
; oligolecty on
<taxonomicName id="2E824D69AC69434B08B3FF708A470694" box="[935,1058,187,213]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
is currently unknown in the true
<taxonomicName id="2E824D69AC69434B0B83FF148F7706B8" authorityName="Hedicke" authorityYear="1933" box="[151,274,223,249]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0B83FF148F7706B8" box="[151,274,223,249]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="BDFD6662AC7643540B83F9A48C280091" ID-DOI="http://doi.org/10.5281/zenodo.13209373" ID-Zenodo-Dep="13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1647,1671]" targetBox="[151,1436,181,1624]" targetPageId="47" targetType="figure">
<paragraph id="E93D36EAAC7643540B83F9A48C280091" blockId="47.[151,1437,1647,1744]" pageId="47" pageNumber="48">
<emphasis id="DBF6EAF8AC7643540B83F9A48F4500C6" bold="true" box="[151,288,1647,1671]" pageId="47" pageNumber="48">FIGURE 30.</emphasis>
<taxonomicName id="2E824D69AC7643540A31F9BB8DBE00C6" authority="Tadauchi, Miyanaga &amp; Dawut, 2005" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[293,987,1647,1672]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="almas" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7643540A31F9BB8F1E00C9" box="[293,379,1648,1672]" italics="true" pageId="47" pageNumber="48">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7643540A9EF9BB8C7F00C9" box="[394,538,1648,1672]" italics="true" pageId="47" pageNumber="48">Chrysandrena</emphasis>
)
<emphasis id="DBF6EAF8AC764354093CF9BB8C0000C9" box="[552,613,1648,1672]" italics="true" pageId="47" pageNumber="48">almas</emphasis>
<bibRefCitation id="8D134B1BAC764354097EF9BB8DBE00C6" author="Tadauchi, O. &amp; Miyanaga, R. &amp; Dawut, A." box="[618,987,1647,1672]" pageId="47" pageNumber="48" pagination="9 - 17" refId="ref91507" refString="Tadauchi, O., Miyanaga, R. &amp; Dawut, A. (2005) A new species belonging to the subgenus Euandrena of the genus Andrena from Xinjiang Uygur, China with notes on nest structure (Hymenoptera, Andrenidae). Esakia, 45, 9 - 17. https: // doi. org / 10.5109 / 2699" type="journal article" year="2005">Tadauchi, Miyanaga &amp; Dawut, 2005</bibRefCitation>
</taxonomicName>
female from Kazakhstan. A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC69434B0BD3FEC88A250420" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
This association between
<taxonomicName id="2E824D69AC69434B0AFBFEC88CEB075C" authorityName="Hedicke" authorityYear="1933" box="[495,654,259,285]" pageId="48" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC69434B0AFBFEC88CEB075C" box="[495,654,259,285]" italics="true" pageId="48" pageNumber="49">Chrysandrena</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC69434B09DCFEC88D26075C" box="[712,835,259,285]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
can be reinforced by the new transfer made here of
<taxonomicName id="2E824D69AC69434B0B83FEEC8CBA0700" authority="Tadauchi, Miyanaga &amp; Dawut, 2005" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[151,735,294,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="almas">
<emphasis id="DBF6EAF8AC69434B0B83FEEC8F250700" box="[151,320,295,321]" italics="true" pageId="48" pageNumber="49">Andrena almas</emphasis>
<bibRefCitation id="8D134B1BAC69434B0A53FEEC8CBA0700" author="Tadauchi, O. &amp; Miyanaga, R. &amp; Dawut, A." box="[327,735,294,321]" pageId="48" pageNumber="49" pagination="9 - 17" refId="ref91507" refString="Tadauchi, O., Miyanaga, R. &amp; Dawut, A. (2005) A new species belonging to the subgenus Euandrena of the genus Andrena from Xinjiang Uygur, China with notes on nest structure (Hymenoptera, Andrenidae). Esakia, 45, 9 - 17. https: // doi. org / 10.5109 / 2699" type="journal article" year="2005">Tadauchi, Miyanaga &amp; Dawut, 2005</bibRefCitation>
</taxonomicName>
from
<taxonomicName id="2E824D69AC69434B0837FEEC8DFB0700" authorityName="Hedicke" authorityYear="1933" box="[803,926,295,321]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0837FEEC8DFB0700" box="[803,926,295,321]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
to
<taxonomicName id="2E824D69AC69434B08D6FEEC8A040700" authorityName="Hedicke" authorityYear="1933" box="[962,1121,295,321]" pageId="48" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC69434B08D6FEEC8A040700" box="[962,1121,295,321]" italics="true" pageId="48" pageNumber="49">Chrysandrena</emphasis>
</taxonomicName>
. This species was described from
<collectingRegion id="2B46F808AC69434B0BC0FE808F530724" box="[212,310,331,357]" country="China" name="Xinjiang Uygur" pageId="48" pageNumber="49">Xinjiang</collectingRegion>
in western
<collectingCountry id="9195767AAC69434B0AA3FE808F9F0724" box="[439,506,331,357]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
where it flies in August and is associated with
<taxonomicName id="2E824D69AC69434B0F12FE808B6C0724" box="[1030,1289,331,357]" class="Magnoliopsida" family="Asteraceae" genus="Chondrilla" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="brevirostris">
<emphasis id="DBF6EAF8AC69434B0F12FE808B6C0724" box="[1030,1289,331,357]" italics="true" pageId="48" pageNumber="49">Chondrilla brevirostris</emphasis>
</taxonomicName>
(
<taxonomicName id="2E824D69AC69434B0E02FE808BF00724" box="[1302,1429,331,357]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) (
<bibRefCitation id="8D134B1BAC69434B0B8AFEA48FF107C8" author="Tadauchi, O. &amp; Miyanaga, R. &amp; Dawut, A." box="[158,404,366,393]" pageId="48" pageNumber="49" pagination="9 - 17" refId="ref91507" refString="Tadauchi, O., Miyanaga, R. &amp; Dawut, A. (2005) A new species belonging to the subgenus Euandrena of the genus Andrena from Xinjiang Uygur, China with notes on nest structure (Hymenoptera, Andrenidae). Esakia, 45, 9 - 17. https: // doi. org / 10.5109 / 2699" type="journal article" year="2005">
Tadauchi
<emphasis id="DBF6EAF8AC69434B0A05FEA48F2307C8" box="[273,326,367,393]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2005
</bibRefCitation>
). In the description, the authors clearly mention the plumose scopal hairs, but concluded on a placement in the
<taxonomicName id="2E824D69AC69434B0A8CFE588C7607EC" authorityName="Hedicke" authorityYear="1933" box="[408,531,403,429]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0A8CFE588C7607EC" box="[408,531,403,429]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
. Confusingly,
<taxonomicName id="2E824D69AC69434B09A8FE588D4507EC" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[700,800,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="almas">
<emphasis id="DBF6EAF8AC69434B09A8FE588D4507EC" box="[700,800,403,429]" italics="true" pageId="48" pageNumber="49">A. almas</emphasis>
</taxonomicName>
was later not included in a subsequent revision of East Asian
<taxonomicName id="2E824D69AC69434B0BF7FE7C8F3B0790" authorityName="Hedicke" authorityYear="1933" box="[227,350,439,465]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0BF7FE7C8F3B0790" box="[227,350,439,465]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D134B1BAC69434B0A64FE7C8C0D0790" author="Xu, H. - L. &amp; Tadauchi, O." box="[368,616,439,465]" pageId="48" pageNumber="49" pagination="77 - 90" refId="ref92855" refString="Xu, H. - L. &amp; Tadauchi, O. (2012) A revision of the subgenus Euandrena of the genus Andrena of Eastern Asia (Hymenoptera: Apoidea: Andrenidae). Esakia, 52, 77 - 90. https: // doi. org / 10.5109 / 25400" type="journal article" year="2012">Xu &amp; Tadauchi 2012</bibRefCitation>
) for unspecified reasons even though its description was mentioned in the introduction of this work. The combination of plumose scopal hairs, summer flight period, and association with
<taxonomicName id="2E824D69AC69434B0BC5FE348F290458" box="[209,332,511,537]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
supports movement from
<taxonomicName id="2E824D69AC69434B0968FE348C920458" authorityName="Hedicke" authorityYear="1933" box="[636,759,511,537]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0968FE348C920458" box="[636,759,511,537]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
to
<emphasis id="DBF6EAF8AC69434B080BFE348DA30458" box="[799,966,511,537]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="2E824D69AC69434B080BFE348DA70458" authorityName="Hedicke" authorityYear="1933" box="[799,962,511,537]" pageId="48" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">Chrysandrena</taxonomicName>
.
</emphasis>
Examination of additional material from
<collectingCountry id="9195767AAC69434B0B83FDE88F7F047C" box="[151,282,547,573]" name="Kazakhstan" pageId="48" pageNumber="49">Kazakhstan</collectingCountry>
(
<figureCitation id="71B92A6FAC69434B0A24FDE88F8E047C" box="[304,491,547,573]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30A–D</figureCitation>
),
<collectingCountry id="9195767AAC69434B0911FDE88CEB047C" box="[517,654,547,573]" name="Afghanistan" pageId="48" pageNumber="49">Afghanistan</collectingCountry>
(
<figureCitation id="71B92A6FAC69434B09B0FDE88D32047C" box="[676,855,547,573]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30E–F</figureCitation>
), and
<collectingCountry id="9195767AAC69434B08BDFDE88D91047C" box="[937,1012,547,573]" name="Russia" pageId="48" pageNumber="49">Russia</collectingCountry>
(Siberia) substantially expands the geographic range, and the species may be present in other dry parts of Central Asia.
</paragraph>
<paragraph id="E93D36EAAC69434B0BD3FDA08B17048C" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
Separately,
<bibRefCitation id="8D134B1BAC69434B0A58FDA08F8704C4" author="Smith, F." box="[332,482,619,645]" pageId="48" pageNumber="49" refId="ref91052" refString="Smith, F. (1849) Descriptions of New Species of British Bees. Zoologist, 7, LVII - LX." type="journal volume" year="1849">Smith (1849)</bibRefCitation>
described
<taxonomicName id="2E824D69AC69434B0974FDA08C9804C5" authorityName="Smith" authorityYear="1849" box="[608,765,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="DBF6EAF8AC69434B0974FDA08C9804C5" box="[608,765,619,645]" italics="true" pageId="48" pageNumber="49">A. constrictus</emphasis>
</taxonomicName>
[recte
<taxonomicName id="2E824D69AC69434B085BFDA08D8504C5" authorityName="Smith" authorityYear="1849" box="[847,992,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="constricta">
<emphasis id="DBF6EAF8AC69434B085BFDA08D8504C5" box="[847,992,619,645]" italics="true" pageId="48" pageNumber="49">A. constricta</emphasis>
</taxonomicName>
] from
<collectingCountry id="9195767AAC69434B0F26FDA08AFD04C4" box="[1074,1176,619,645]" name="United Kingdom" pageId="48" pageNumber="49">Scotland</collectingCountry>
. This species has been listed as a synonym of
<taxonomicName id="2E824D69AC69434B0A80FD448C6204E8" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[404,519,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC69434B0A80FD448C6204E8" box="[404,519,655,681]" italics="true" pageId="48" pageNumber="49">A. fulvago</emphasis>
</taxonomicName>
for a long time (e.g.
<bibRefCitation id="8D134B1BAC69434B09F9FD448DF704E8" author="Warncke, K." box="[749,914,655,681]" pageId="48" pageNumber="49" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke 1967</bibRefCitation>
). However,
<taxonomicName id="2E824D69AC69434B0F03FD448AEC04E8" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[1047,1161,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC69434B0F03FD448AEC04E8" box="[1047,1161,655,681]" italics="true" pageId="48" pageNumber="49">A. fulvago</emphasis>
</taxonomicName>
has never been recorded from
<collectingCountry id="9195767AAC69434B0BC1FD788F52048C" box="[213,311,691,717]" name="United Kingdom" pageId="48" pageNumber="49">Scotland</collectingCountry>
(
<bibRefCitation id="8D134B1BAC69434B0A51FD788C24048C" author="Else, G. R. &amp; Edwards, M." box="[325,577,691,717]" pageId="48" pageNumber="49" refId="ref88553" refString="Else, G. R. &amp; Edwards, M. (2018) Handbook of the bees of the British Isles. The Ray Society, London, 775 pp." type="book" year="2018">Else &amp; Edwards 2018</bibRefCitation>
).
<typeStatus id="36398848AC69434B0947FD788CEE048C" box="[595,651,691,717]" pageId="48" pageNumber="49">Type</typeStatus>
inspection is necessary to clarify the true identity of
<taxonomicName id="2E824D69AC69434B0FC8FD788B0E048D" authorityName="Smith" authorityYear="1849" box="[1244,1387,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="constricta">
<emphasis id="DBF6EAF8AC69434B0FC8FD788B0E048D" box="[1244,1387,691,717]" italics="true" pageId="48" pageNumber="49">A. constricta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC69434B0BD3FD1D8AAD05E4" pageId="48" pageNumber="49" type="distribution">
<paragraph id="E93D36EAAC69434B0BD3FD1D8D540554" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
<emphasis id="DBF6EAF8AC69434B0BD3FD1D8F0404B1" bold="true" box="[199,353,726,752]" pageId="48" pageNumber="49">Distribution.</emphasis>
Palaearctic, from
<collectingCountry id="9195767AAC69434B093BFD1C8C1504B0" box="[559,624,727,753]" name="Georgia" pageId="48" pageNumber="49">Iberia</collectingCountry>
across Europe and northern
<collectingCountry id="9195767AAC69434B08A7FD1C8A6604B0" box="[947,1027,727,753]" name="Turkey" pageId="48" pageNumber="49">Turkey</collectingCountry>
to the Caucasus and Eastern Siberia (
<bibRefCitation id="8D134B1BAC69434B0B8BFD308F920554" author="Gusenleitner, F. &amp; Schwarz, M." box="[159,503,763,789]" pageId="48" pageNumber="49" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC69434B0916FD308D400554" author="Proshchalykin M. Y. &amp; Astafurova Y. V. &amp; Sidorov D. A." box="[514,805,763,789]" pageId="48" pageNumber="49" pagination="263 - 276" refId="ref90418" refString="Proshchalykin M. Y., Astafurova Y. V. &amp; Sidorov D. A. (2017) Family Andrenidae. In: Lelej, A. S., Proshchalykin, M. Y. &amp; Loktionov, V. M. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute Russian Academy of Sciences, Supplement 6, 263 - 276." type="journal article" year="2017">
Proshchalykin
<emphasis id="DBF6EAF8AC69434B09BDFD308CBF0554" box="[681,730,763,789]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
<paragraph id="E93D36EAAC69434B0BD3FCD58AAD05E4" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
<emphasis id="DBF6EAF8AC69434B0BD3FCD58F990579" bold="true" box="[199,508,798,824]" pageId="48" pageNumber="49">Other material examined</emphasis>
(
<taxonomicName id="2E824D69AC69434B0904FCD48CD80578" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[528,701,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="almas">
<emphasis id="DBF6EAF8AC69434B0904FCD48CD80578" box="[528,701,799,825]" italics="true" pageId="48" pageNumber="49">Andrena almas</emphasis>
</taxonomicName>
)
<materialsCitation id="59EA3CB7AC69434B09C5FCD58BFD051C" collectingDate="1962-08-07" collectorName="Baloutchah, W &amp; K. Lindberg &amp; No A &amp; Entomology Collection &amp; Lund Museum" country="Afghanistan" location="AFGHANISTAN" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC69434B09C5FCD58DCD0579" bold="true" box="[721,936,798,824]" pageId="48" pageNumber="49">
<collectingCountry id="9195767AAC69434B09C5FCD58DC70579" box="[721,930,798,824]" name="Afghanistan" pageId="48" pageNumber="49">AFGHANISTAN</collectingCountry>
:
</emphasis>
<collectorName id="4477533CAC69434B08A6FCD48A380578" box="[946,1117,799,825]" pageId="48" pageNumber="49">Baloutchah, W</collectorName>
Tagab Echnan,
<date id="9D3C102AAC69434B0E08FCD48BFD0578" box="[1308,1432,799,825]" pageId="48" pageNumber="49" value="1962-08-07">
<collectingDate id="8D78E9C2AC69434B0E08FCD48BFD0578" box="[1308,1432,799,825]" pageId="48" pageNumber="49" value="1962-08-07">7.viii.1962</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC69434B0B83FC888EDF051C" box="[151,186,835,861]" count="1" pageId="48" pageNumber="49" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC69434B0A15FC888FF5051C" box="[257,400,835,861]" pageId="48" pageNumber="49">K. Lindberg</collectorName>
, OÖLM (
<collectorName id="4477533CAC69434B0910FC888C20051C" box="[516,581,835,861]" pageId="48" pageNumber="49">No A</collectorName>
1075, ex. coll.
<collectorName id="4477533CAC69434B0816FC888A75051C" box="[770,1040,835,861]" pageId="48" pageNumber="49">Entomology Collection</collectorName>
,
<collectorName id="4477533CAC69434B0F0AFC888AAE051C" box="[1054,1227,835,861]" pageId="48" pageNumber="49">Lund Museum</collectorName>
;
<figureCitation id="71B92A6FAC69434B0FCEFC888BF5051C" box="[1242,1424,835,861]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30A–D</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC69434B0B83FCAD8AAC05C0" box="[151,1225,870,899]" collectingDate="1991-08-07" collectorName="M. Snizek" country="Kazakhstan" location="Alma-Ata" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Almaty Oblysy">
<emphasis id="DBF6EAF8AC69434B0B83FCAD8F0305C1" bold="true" box="[151,358,870,896]" pageId="48" pageNumber="49">
<collectingCountry id="9195767AAC69434B0B83FCAD8F0405C1" box="[151,353,870,896]" name="Kazakhstan" pageId="48" pageNumber="49">KAZAKHSTAN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC69434B0A7FFCAC8FBE05C0" LSID="urn:lsid:plazi:treatment:612B87FCAC77434B0B83FA948AAD05E4:EC5D6031AC69434B0A7FFCAC8FBE05C0" box="[363,475,871,897]" country="Kazakhstan" name="Alma-Ata" pageId="48" pageNumber="49" stateProvince="Almaty Oblysy">Alma-Ata</location>
[
<collectingRegion id="2B46F808AC69434B0AF3FCAC8C2605C0" box="[487,579,871,897]" country="Kazakhstan" name="Almaty Oblysy" pageId="48" pageNumber="49">Almaty</collectingRegion>
],
<date id="9D3C102AAC69434B0940FCAC8CB505C0" box="[596,720,871,897]" pageId="48" pageNumber="49" value="1991-08-07">
<collectingDate id="8D78E9C2AC69434B0940FCAC8CB505C0" box="[596,720,871,897]" pageId="48" pageNumber="49" value="1991-08-07">7.viii.1991</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC69434B09CEFCAC8C9805C0" box="[730,765,871,897]" count="1" pageId="48" pageNumber="49" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC69434B082CFCAC8DC805C0" box="[824,941,871,897]" pageId="48" pageNumber="49">M. Snižek</collectorName>
, OÖLM (
<figureCitation id="71B92A6FAC69434B0F02FCAC8AA405C0" box="[1046,1217,871,897]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30E–F</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC69434B0FC7FCAD8AA105E4" collectingDate="2010-08-03" collectionCode="NHMUK" collectorName="Michez &amp; DeMeule" country="Russia" elevation="282" location="Cherepanovo" municipality="Eastern Siberia" pageId="48" pageNumber="49" specimenCount="5" specimenCount-female="5">
<emphasis id="DBF6EAF8AC69434B0FC7FCAD8B2605C1" bold="true" box="[1235,1347,870,896]" pageId="48" pageNumber="49">
<collectingCountry id="9195767AAC69434B0FC7FCAD8B5B05C1" box="[1235,1342,870,896]" name="Russia" pageId="48" pageNumber="49">RUSSIA</collectingCountry>
:
</emphasis>
<collectingMunicipality id="0959AC90AC69434B0E5CFCAC8E8C05E4" pageId="48" pageNumber="49">Eastern Siberia</collectingMunicipality>
,
<location id="EC5D6031AC69434B0BE1FC408FE805E4" LSID="urn:lsid:plazi:treatment:612B87FCAC77434B0B83FA948AAD05E4:EC5D6031AC69434B0BE1FC408FE805E4" box="[245,397,907,933]" country="Russia" municipality="Eastern Siberia" name="Cherepanovo" pageId="48" pageNumber="49">Cherepanovo</location>
, train station,
<quantity id="2E7A9B0FAC69434B093FFC408C1105E4" box="[555,628,907,933]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="48" pageNumber="49" unit="m" value="282.0">
<elevation id="62AFD1D9AC69434B093FFC408C1105E4" box="[555,628,907,933]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="48" pageNumber="49" unit="m" value="282.0">282 m</elevation>
</quantity>
,
<date id="9D3C102AAC69434B0994FC408C9E05E4" box="[640,763,907,933]" pageId="48" pageNumber="49" value="2010-08-03">
<collectingDate id="8D78E9C2AC69434B0994FC408C9E05E4" box="[640,763,907,933]" pageId="48" pageNumber="49" value="2010-08-03">3.viii.2010</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC69434B0813FC418D4E05E4" box="[775,811,906,933]" count="5" pageId="48" pageNumber="49" type="female">5♀</specimenCount>
, leg.
<collectorName id="4477533CAC69434B087CFC408DD905E4" box="[872,956,907,933]" pageId="48" pageNumber="49">Michez</collectorName>
&amp;
<collectorName id="4477533CAC69434B08F4FC408A2E05E4" box="[992,1099,907,933]" pageId="48" pageNumber="49">DeMeule</collectorName>
,
<collectionCode id="8F93AE2FAC69434B0F42FC408AA105E4" box="[1110,1220,907,933]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="48" pageNumber="49" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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