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<mods:title id="52097524E09969758A7B9C1E184C2F3F">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<subSubSection id="A1986561AC1143330B83FEA58D0307C8" box="[151,870,366,393]" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="E93D36EAAC1143330B83FEA58D0307C8" blockId="72.[151,870,366,393]" box="[151,870,366,393]" pageId="72" pageNumber="73">
<heading id="B2758186AC1143330B83FEA58D0307C8" bold="true" box="[151,870,366,393]" fontSize="11" level="2" pageId="72" pageNumber="73" reason="4">
<emphasis id="DBF6EAF8AC1143330B83FEA58D0307C8" bold="true" box="[151,870,366,393]" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC1143330B83FEA58C9007C9" ID-CoL="DSMC" authority="Noskiewicz, 1939" authorityName="Noskiewicz" authorityYear="1939" box="[151,757,366,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi" status="stat. nov." subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC1143330B83FEA58F6407C8" bold="true" box="[151,257,366,393]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1143330A06FEA58FD707C8" bold="true" box="[274,434,366,393]" italics="true" pageId="72" pageNumber="73">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1143330AD7FEA58C7D07C8" bold="true" box="[451,536,366,393]" italics="true" pageId="72" pageNumber="73">pillichi</emphasis>
<bibRefCitation id="8D134B1BAC114333090BFEA58C9007C9" author="Noskiewicz, J." box="[543,757,366,392]" pageId="72" pageNumber="73" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz, 1939</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC11433309E8FEA48D0307C8" box="[764,870,367,393]" pageId="72" pageNumber="73">stat. rev.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330B83FE788D3E078A" box="[151,859,435,460]" pageId="72" pageNumber="73" type="reference_group">
<paragraph id="E93D36EAAC1143330B83FE788D3E078A" blockId="72.[151,859,435,460]" box="[151,859,435,460]" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC1143330B83FE788FBD078A" authorityName="Noskiewicz" authorityYear="1939" box="[151,472,435,459]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC1143330B83FE788E88078A" box="[151,237,435,459]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1143330BE8FE788FE4078A" box="[252,385,435,459]" italics="true" pageId="72" pageNumber="73">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1143330A84FE788FBD078A" box="[400,472,435,459]" italics="true" pageId="72" pageNumber="73">pillichi</emphasis>
</taxonomicName>
<materialsCitation id="59EA3CB7AC1143330ACAFE7F8D3E078A" box="[478,859,435,460]" collectionCode="UWCP" country="Hungary" location="Hungary" pageId="72" pageNumber="73" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="FF84FD63AC1143330ACAFE7F8F8B078A" box="[478,494,436,459]" count="1" pageId="72" pageNumber="73" type="female"></specimenCount>
<specimenCount id="FF84FD63AC1143330AF9FE7F8C66078A" box="[493,515,436,459]" count="1" pageId="72" pageNumber="73" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC1143330904FE7F8C09078D" box="[528,620,436,460]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
,
<collectionCode id="8F93AE2FAC1143330962FE788CDA078A" box="[630,703,435,459]" country="Poland" name="University of Wroclaw" pageId="72" pageNumber="73">UWCP</collectionCode>
, not examined]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330B83FE328A85036A" pageId="72" pageNumber="73" type="discussion">
<paragraph id="E93D36EAAC1143330B83FE328DEF053A" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330B83FE328F6E0452" bold="true" box="[151,267,505,531]" pageId="72" pageNumber="73">Remarks.</emphasis>
<taxonomicName id="2E824D69AC1143330A00FE328FA90452" authorityName="Noskiewicz" authorityYear="1939" box="[276,460,505,531]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330A00FE328FA90452" box="[276,460,505,531]" italics="true" pageId="72" pageNumber="73">Andrena pillichi</emphasis>
</taxonomicName>
was described from Simontornya in
<collectingCountry id="9195767AAC114333086DFE318DB90455" box="[889,988,506,532]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
, collected by F. Pillich who sent much material from this region to other workers, leading to the descriptions of multiple
<taxonomicName id="2E824D69AC1143330F44FDD68B6E0476" box="[1104,1291,541,567]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330F44FDD68ACA0476" box="[1104,1199,541,567]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
species
</taxonomicName>
(e.g.
<bibRefCitation id="8D134B1BAC1143330E5AFDD68EB7041A" author="Alfken, J. D." pageId="72" pageNumber="73" pagination="88 - 93" refId="ref87099" refString="Alfken, J. D. (1933) Beitrage zur Kenntnis palaarktischer Bienen (Hym. Apid.). 3. Beitrag. Mitteilungen der Deutschen Entomologischen Gesellschaft, 4, 88 - 93. https: // doi. org / 10.1002 / mmnd. 193319330105" type="journal article" year="1933">Alfken 1933</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1143330BF4FD8A8F7E041A" author="Alfken, J. D." box="[224,283,577,603]" pageId="72" pageNumber="73" pagination="377 - 383" refId="ref87148" refString="Alfken, J. D. (1936) Neue Andrena-Arten vom Wiener Becken, aus dem Burgenland und Ungarn. Veroffentlichungen aus dem Uberseemuseum Bremen, 1, 377 - 383." type="journal article" year="1936">1936</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1143330A3DFD8A8F97041A" author="Noskiewicz, J." box="[297,498,577,603]" pageId="72" pageNumber="73" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz 1939</bibRefCitation>
). Noskiewicz gave detailed identification criteria to allow separation of
<taxonomicName id="2E824D69AC1143330E38FD898BF9041A" authorityName="Noskiewicz" authorityYear="1939" box="[1324,1436,577,604]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330E38FD898BF9041A" box="[1324,1436,577,604]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
from
<taxonomicName id="2E824D69AC1143330BC0FDAD8F05043E" authorityName="Perkins" authorityYear="1914" box="[212,352,613,640]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC1143330BC0FDAD8F05043E" box="[212,352,613,640]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
; both species are bivoltine, but
<taxonomicName id="2E824D69AC11433309D6FDAD8D4A043E" authorityName="Noskiewicz" authorityYear="1939" box="[706,815,613,640]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC11433309D6FDAD8D4A043E" box="[706,815,613,640]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
is noticeably shinier on the clypeus, scutum, scutellum, and tergal discs. Detailed identification criteria are given by
<bibRefCitation id="8D134B1BAC1143330852FD428A7A04E2" author="Noskiewicz, J." box="[838,1055,649,675]" pageId="72" pageNumber="73" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz (1939)</bibRefCitation>
,
<bibRefCitation id="8D134B1BAC1143330F38FD428B6504E2" author="Osytshnjuk, A. Z." box="[1068,1280,649,675]" pageId="72" pageNumber="73" pagination="315 - 369" refId="ref89594" refString="Osytshnjuk, A. Z. (1978) The identification of the insects of the European part of the USSR. Vol. 3. Hymenoptera. Part one. Opred Faune SSSR, 119, 315 - 369. [in Russian]" type="journal article" year="1978">Osytshnjuk (1978)</bibRefCitation>
, and
<bibRefCitation id="8D134B1BAC1143330E29FD428FD90486" author="Schmid-Egger, C. &amp; Scheuchl, E." pageId="72" pageNumber="73" refId="ref90781" refString="Schmid-Egger, C. &amp; Scheuchl, E. (1997) Illustrierte Bestimmungstabellen der Wildbienen Deutschlands und Osterreichs und Berucksichtigung der Arten der Schweiz. Band III. Andrenidae. Velden, Eigenverlag, 180 pp." type="book" year="1997">Schmid-Egger &amp; Scheuchl (1997)</bibRefCitation>
.
<bibRefCitation id="8D134B1BAC1143330AD3FD668C190486" author="Warncke, K." box="[455,636,685,711]" pageId="72" pageNumber="73" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke (1967)</bibRefCitation>
considered
<taxonomicName id="2E824D69AC1143330812FD658D100486" authorityName="Noskiewicz" authorityYear="1939" box="[774,885,685,712]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330812FD658D100486" box="[774,885,685,712]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
to be a valid species, but
<bibRefCitation id="8D134B1BAC1143330F89FD668B3C0486" author="Dylewska, M." box="[1181,1369,685,711]" pageId="72" pageNumber="73" pagination="359 - 708" refId="ref88522" refString="Dylewska, M. (1987) Die Gattung Andrena Fabricius (Andrenidae, Apoidea) in Nord- und Mitteleuropa. Acta Zoologica Cracoviensia, 30, 359 - 708." type="journal article" year="1987">Dylewska (1987</bibRefCitation>
, as
<taxonomicName id="2E824D69AC1143330E90FD658EBA04AA" authorityName="Noskiewicz" authorityYear="1939" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pilichi">
<emphasis id="DBF6EAF8AC1143330E90FD658EBA04AA" italics="true" pageId="72" pageNumber="73">A. pilichi</emphasis>
</taxonomicName>
subsequent incorrect spelling) and
<bibRefCitation id="8D134B1BAC1143330979FD1A8D3B04AA" author="Schwarz, M. &amp; Gusenleitner, F. &amp; Westrich, P. &amp; Dathe, H. H." box="[621,862,721,748]" pageId="72" pageNumber="73" pagination="1 - 398" refId="ref90955" refString="Schwarz, M., Gusenleitner, F., Westrich, P. &amp; Dathe, H. H. (1996) Katalog der Bienen Osterreichs, Deutschlands und der Schweiz. (Hymenoptera, Apidae). Entomofauna, Supplement 8, 1 - 398." type="journal article" year="1996">
Schwarz
<emphasis id="DBF6EAF8AC11433309C0FD198D6304AA" box="[724,774,721,747]" italics="true" pageId="72" pageNumber="73">et al</emphasis>
. (1996)
</bibRefCitation>
synonymised it with
<taxonomicName id="2E824D69AC1143330F44FD198AB804AA" authorityName="Perkins" authorityYear="1914" box="[1104,1245,721,748]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC1143330F44FD198AB804AA" box="[1104,1245,721,748]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
. Dylewska (who revised the
<typeStatus id="36398848AC1143330A08FD3D8F290551" box="[284,332,758,784]" pageId="72" pageNumber="73">type</typeStatus>
series) argued that some of the distinguishing characters given by Noskiewicz were insufficient, such as the antennal ratio of the males, the identical genital capsule, and variation in the degree of polish on the clypeus and scutellum. Schwarz
<emphasis id="DBF6EAF8AC114333091CFCF58C5F0516" box="[520,570,829,855]" italics="true" pageId="72" pageNumber="73">et al</emphasis>
. also argued that, based on a study of these two species including specimens from the Warncke collection, they could not consider them distinct.
</paragraph>
<paragraph id="E93D36EAAC1143330BD3FC4E8A85036A" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
Fieldwork in
<collectingCountry id="9195767AAC1143330A4BFC4E8FA105DE" box="[351,452,901,927]" name="Romania" pageId="72" pageNumber="73">Romania</collectingCountry>
produced long series of
<emphasis id="DBF6EAF8AC11433309C8FC4D8D8105DE" box="[732,996,901,928]" italics="true" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC11433309C8FC4D8D0F05DE" authorityName="Perkins" authorityYear="1914" box="[732,874,901,928]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">A. alfkenella</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1143330866FC4D8D8105DE" box="[882,996,901,927]" pageId="72" pageNumber="73" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
specimens. From two sites around
<collectingRegion id="2B46F808AC1143330E60FC4C8BF905E3" box="[1396,1436,903,930]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
in north-eastern
<collectingCountry id="9195767AAC1143330A42FC628FDB0582" box="[342,446,937,963]" name="Romania" pageId="72" pageNumber="73">Romania</collectingCountry>
, Ciurea (open fallow agricultural fields) and Ursoaia (open woody steppe grassland), seven sequences of
<taxonomicName id="2E824D69AC1143330A66FC058C3105A6" box="[370,596,973,1000]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC1143330A66FC058C3105A6" box="[370,596,973,1000]" italics="true" pageId="72" pageNumber="73">A. alfkenella pillichi</emphasis>
</taxonomicName>
were generated which showed low average intraspecific variation of 0.17% (
<figureCitation id="71B92A6FAC1143330B8BFC3A8F77024A" box="[159,274,1009,1035]" captionStart="FIGURE 37" captionStartId="57.[151,250,1819,1843]" captionTargetBox="[151,1433,311,1794]" captionTargetId="figure-78@57.[151,1435,309,1796]" captionTargetPageId="57" captionText="FIGURE 37. Phylogenetic tree (maximum likelihood) of Andrena subgenus Micrandrena Ashmead, 1899 based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi="http://doi.org/10.5281/zenodo.13209387" httpUri="https://zenodo.org/record/13209387/files/figure.png" pageId="72" pageNumber="73">Figure 37</figureCitation>
). At the Ciurea site, a single Romanian specimen of
<taxonomicName id="2E824D69AC1143330872FC398D91024A" authorityName="Perkins" authorityYear="1914" box="[870,1012,1009,1036]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC1143330872FC398D91024A" box="[870,1012,1009,1036]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC11433308E8FC398A5D024D" box="[1020,1080,1010,1036]" pageId="72" pageNumber="73" sensu="stricto">s. str.</taxonomicNameLabel>
was found in sympatry with
<taxonomicName id="2E824D69AC1143330E90FC398F3B026E" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC1143330E90FC398F3B026E" italics="true" pageId="72" pageNumber="73">A. alfkenella pillichi</emphasis>
</taxonomicName>
. This specimen was separated from the
<taxonomicName id="2E824D69AC1143330827FBDD8A7E026E" box="[819,1051,1045,1072]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC1143330827FBDD8A7E026E" box="[819,1051,1045,1072]" italics="true" pageId="72" pageNumber="73">A. alfkenella pillichi</emphasis>
</taxonomicName>
sequences by an average genetic distance of 3.76% (range 3.653.95%). More broadly, when including
<taxonomicName id="2E824D69AC11433308B8FBF18A5C0212" authorityName="Perkins" authorityYear="1914" box="[940,1081,1081,1108]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC11433308B8FBF18A5C0212" box="[940,1081,1081,1108]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1143330F54FBF18A1E0215" box="[1088,1147,1082,1108]" pageId="72" pageNumber="73" sensu="stricto">s. str.</taxonomicNameLabel>
sequences from
<collectingCountry id="9195767AAC1143330E22FBF28BFD0212" box="[1334,1432,1081,1107]" name="Portugal" pageId="72" pageNumber="73">Portugal</collectingCountry>
,
<collectingCountry id="9195767AAC1143330B83FB968EBF0236" box="[151,218,1117,1143]" name="Spain" pageId="72" pageNumber="73">Spain</collectingCountry>
,
<collectingCountry id="9195767AAC1143330BF7FB968F290236" box="[227,332,1117,1143]" name="Germany" pageId="72" pageNumber="73">Germany</collectingCountry>
, and
<collectingCountry id="9195767AAC1143330A90FB968FB60236" box="[388,467,1117,1143]" name="Greece" pageId="72" pageNumber="73">Greece</collectingCountry>
(
<collectingRegion id="2B46F808AC1143330AF4FB968C440236" box="[480,545,1117,1143]" country="Greece" name="Kriti" pageId="72" pageNumber="73">Crete</collectingRegion>
, see above), the two taxa were separated by an average genetic distance of 4.08% (range 3.654.56%). Since the two taxa can be found in direct sympatry with consistent genetic differentiation and since their morphology is consistent with the criteria identified by Noskiewicz (with the caveat that some morphological variation is typical in members of the subgenus
<taxonomicName id="2E824D69AC114333086EFB028A6802A2" authorityName="Ashmead" authorityYear="1899" box="[890,1037,1225,1251]" pageId="72" pageNumber="73" rank="subGenus" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC114333086EFB028A6802A2" box="[890,1037,1225,1251]" italics="true" pageId="72" pageNumber="73">Micrandrena</emphasis>
</taxonomicName>
, particularly bivoltine species),
<taxonomicName id="2E824D69AC1143330E90FB018E830346" authorityName="WOOD" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi" status="stat. rev.">
<emphasis id="DBF6EAF8AC1143330E90FB018E830346" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC1143330BFAFB268F360346" bold="true" box="[238,339,1261,1287]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="C0C55783AC1143330BFAFB268F360346" box="[238,339,1261,1287]" pageId="72" pageNumber="73">stat. rev.</taxonomicNameLabel>
</emphasis>
is returned to species status. Revision of European material of
<emphasis id="DBF6EAF8AC1143330F0CFB258B7B0346" box="[1048,1310,1261,1288]" italics="true" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC1143330F0CFB258AC30346" authorityName="Perkins" authorityYear="1914" box="[1048,1190,1261,1288]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">A. alfkenella</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1143330FB9FB258B7B0346" box="[1197,1310,1261,1287]" pageId="72" pageNumber="73" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
is required to establish the precise range limits, though
<taxonomicName id="2E824D69AC1143330990FAD98C97036A" authorityName="WOOD" authorityYear="2024" box="[644,754,1297,1324]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330990FAD98C97036A" box="[644,754,1297,1324]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
appears to be a strongly Pannonian species.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330BD3FAFE8F57006B" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="E93D36EAAC1143330BD3FAFE8F57006B" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330BD3FAFE8FD4030E" bold="true" box="[199,433,1333,1359]" pageId="72" pageNumber="73">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC1143330AA3FAFE8BFD0313" box="[439,1432,1333,1362]" collectingDate="2009-07-22" collectorName="J. Halada" country="Austria" location="1 km E Poysdorf" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Niederoesterreich">
<emphasis id="DBF6EAF8AC1143330AA3FAFE8C5B030E" bold="true" box="[439,574,1333,1359]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330AA3FAFE8C5C030E" box="[439,569,1333,1359]" name="Austria" pageId="72" pageNumber="73">AUSTRIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC1143330951FAFE8D66030E" box="[581,771,1333,1359]" country="Austria" name="Niederoesterreich" pageId="72" pageNumber="73">Niederosterreich</collectingRegion>
,
<location id="EC5D6031AC114333081AFAFE8DAA030E" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC114333081AFAFE8DAA030E" box="[782,975,1333,1360]" country="Austria" name="1 km E Poysdorf" pageId="72" pageNumber="73" stateProvince="Niederoesterreich">
<quantity id="2E7A9B0FAC114333081AFAFE8D2D030E" box="[782,840,1333,1359]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="km" value="1.0">1 km</quantity>
E Poysdorf
</location>
,
<date id="9D3C102AAC11433308CFFAFE8A38030E" box="[987,1117,1333,1359]" pageId="72" pageNumber="73" value="2009-07-22">
<collectingDate id="8D78E9C2AC11433308CFFAFE8A38030E" box="[987,1117,1333,1359]" pageId="72" pageNumber="73" value="2009-07-22">22.vii.2009</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330F7DFAFE8AE9030E" box="[1129,1164,1333,1359]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330FDFFAFD8B53030E" box="[1227,1334,1333,1360]" pageId="72" pageNumber="73">J. Halada</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330B83FA928BFD0337" box="[151,1432,1369,1398]" collectingDate="2003-04-05" collectingDateMax="2003-04-10" collectingDateMin="2003-04-05" collectorName="Y. Budaschkin" country="Austria" county="Krim" location="Vodianja" municipality="Karadagh" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC1143330B83FA928F700332" bold="true" box="[151,277,1369,1395]" pageId="72" pageNumber="73">CRIMEA:</emphasis>
<collectingCounty id="005C4E66AC1143330A34FA928F3B0332" box="[288,350,1369,1395]" pageId="72" pageNumber="73">Krim</collectingCounty>
,
<collectingMunicipality id="0959AC90AC1143330A79FA928FBF0332" box="[365,474,1369,1395]" pageId="72" pageNumber="73">Karadagh</collectingMunicipality>
[Kara Dag],
<location id="EC5D6031AC1143330963FA928CBF0332" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330963FA928CBF0332" box="[631,730,1369,1395]" country="Austria" county="Krim" municipality="Karadagh" name="Vodianja" pageId="72" pageNumber="73">Vodianja</location>
balka,
<location id="EC5D6031AC1143330827FA928D150332" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330827FA928D150332" box="[819,880,1369,1395]" country="Austria" county="Krim" municipality="Karadagh" name="Wald" pageId="72" pageNumber="73">Wald</location>
,
<date id="9D3C102AAC114333086BFA928A760335" box="[895,1043,1369,1396]" pageId="72" pageNumber="73" value="2003-04-05" valueMax="2003-04-10" valueMin="2003-04-05">
<collectingDate id="8D78E9C2AC114333086BFA928A760335" box="[895,1043,1369,1396]" pageId="72" pageNumber="73" value="2003-04-05" valueMax="2003-04-10" valueMin="2003-04-05">510.iv.2003</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330F37FA928A200332" box="[1059,1093,1369,1395]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330F9EFA918B560332" box="[1162,1331,1369,1396]" pageId="72" pageNumber="73">Y. Budaschkin</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330B83FAB68DCA03DB" box="[151,943,1405,1434]" collectingDate="1941-08-17" collectorName="V. Zavadil" country="Czech Republic" location="Cejc" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Jihomoravsky">
<emphasis id="DBF6EAF8AC1143330B83FAB68F4603D6" bold="true" box="[151,291,1405,1431]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330B83FAB68F7B03D6" box="[151,286,1405,1431]" name="Czech Republic" pageId="72" pageNumber="73">CZECHIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC1143330A3FFAB68FE803D6" box="[299,397,1405,1431]" country="Czech Republic" name="Jihomoravsky" pageId="72" pageNumber="73">Moravia</collectingRegion>
,
<location id="EC5D6031AC1143330A8DFAB48FAA03DB" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330A8DFAB48FAA03DB" box="[409,463,1407,1434]" country="Czech Republic" name="Cejc" pageId="72" pageNumber="73" stateProvince="Jihomoravsky">Čejč</location>
,
<date id="9D3C102AAC1143330AC8FAB68C0303D6" box="[476,614,1405,1431]" pageId="72" pageNumber="73" value="1941-08-17">
<collectingDate id="8D78E9C2AC1143330AC8FAB68C0303D6" box="[476,614,1405,1431]" pageId="72" pageNumber="73" value="1941-08-17">17.viii.1941</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330966FAB68CF003D6" box="[626,661,1405,1431]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC11433309C1FAB58D2803D6" box="[725,845,1405,1432]" pageId="72" pageNumber="73">V. Zavadil</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC11433308A8FAB68F1403FF" collectingDate="1941-08-12" collectorName="V. Zavadil" country="Czech Republic" location="Cejkovice" pageId="72" pageNumber="73" specimenCount="1" specimenCount-male="1" stateProvince="Jihomoravsky">
<collectingRegion id="2B46F808AC11433308A8FAB68A7B03D6" box="[956,1054,1405,1431]" country="Czech Republic" name="Jihomoravsky" pageId="72" pageNumber="73">Moravia</collectingRegion>
,
<location id="EC5D6031AC1143330F3EFAB48AFA03DB" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330F3EFAB48AFA03DB" box="[1066,1183,1407,1434]" country="Czech Republic" name="Cejkovice" pageId="72" pageNumber="73" stateProvince="Jihomoravsky">Čejkovice</location>
,
<date id="9D3C102AAC1143330FBEFAB68B5103D6" box="[1194,1332,1405,1431]" pageId="72" pageNumber="73" value="1941-08-12">
<collectingDate id="8D78E9C2AC1143330FBEFAB68B5103D6" box="[1194,1332,1405,1431]" pageId="72" pageNumber="73" value="1941-08-12">12.viii.1941</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330E55FAB68B0003D6" box="[1345,1381,1405,1431]" count="1" pageId="72" pageNumber="73" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC1143330B83FA698F6A03FA" box="[151,271,1441,1468]" pageId="72" pageNumber="73">V. Zavadil</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330A69FA6A8BFD03FA" box="[381,1432,1441,1470]" collectingDate="2023-08-23" collectionCode="TJWC" collectorName="T. J. Wood" country="Romania" location="1 km NNW Ciurea" pageId="72" pageNumber="73" specimenCount="4" specimenCount-female="4" stateProvince="Iasi">
<emphasis id="DBF6EAF8AC1143330A69FA6A8C7703FA" bold="true" box="[381,530,1441,1467]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330A69FA6A8C6803FA" box="[381,525,1441,1467]" name="Romania" pageId="72" pageNumber="73">ROMANIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC114333090EFA688C2003FF" box="[538,581,1443,1470]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
,
<location id="EC5D6031AC1143330945FA6A8D4E03FA" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330945FA6A8D4E03FA" box="[593,811,1441,1468]" country="Romania" name="1 km NNW Ciurea" pageId="72" pageNumber="73" stateProvince="Iasi">
<quantity id="2E7A9B0FAC1143330945FA6A8CEE03FA" box="[593,651,1441,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="km" value="1.0">1 km</quantity>
NNW Ciurea
</location>
, fallow fields,
<date id="9D3C102AAC11433308C7FA6A8A3803FA" box="[979,1117,1441,1467]" pageId="72" pageNumber="73" value="2023-08-23">
<collectingDate id="8D78E9C2AC11433308C7FA6A8A3803FA" box="[979,1117,1441,1467]" pageId="72" pageNumber="73" value="2023-08-23">23.viii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330F7DFA6A8AEB03FA" box="[1129,1166,1441,1467]" count="4" pageId="72" pageNumber="73" type="female">4♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330FD8FA698B2503FA" box="[1228,1344,1441,1468]" pageId="72" pageNumber="73">T.J. Wood</collectorName>
,
<collectionCode id="8F93AE2FAC1143330E5FFA6A8BFD03FA" box="[1355,1432,1441,1467]" pageId="72" pageNumber="73">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330B83FA0C8A90039E" box="[151,1269,1477,1506]" collectingDate="2023-07-21" collectionCode="TJWC" collectorName="T. J. Wood" country="Romania" county="Ursoaia" elevation="500" location="Dumnezeu" municipality="Dealul" pageId="72" pageNumber="73" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Iasi">
<collectingRegion id="2B46F808AC1143330B83FA0C8EA703A3" box="[151,194,1479,1506]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
,
<collectingRegion id="2B46F808AC1143330BDBFA0C8E9F03A3" box="[207,250,1479,1506]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
,
<quantity id="2E7A9B0FAC1143330A12FA0E8F2B03A1" box="[262,334,1477,1504]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="72" pageNumber="73" unit="m" value="500.0">
<elevation id="62AFD1D9AC1143330A12FA0E8F2B03A1" box="[262,334,1477,1504]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="72" pageNumber="73" unit="m" value="500.0">500 m</elevation>
</quantity>
E of
<collectingCounty id="005C4E66AC1143330A9BFA0E8F8C039E" box="[399,489,1477,1503]" pageId="72" pageNumber="73">Ursoaia</collectingCounty>
,
<collectingMunicipality id="0959AC90AC1143330AE1FA0E8C25039E" box="[501,576,1477,1503]" pageId="72" pageNumber="73">Dealul</collectingMunicipality>
lui
<location id="EC5D6031AC114333097AFA0D8C8F03A1" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC114333097AFA0D8C8F03A1" box="[622,746,1478,1504]" country="Romania" county="Ursoaia" municipality="Dealul" name="Dumnezeu" pageId="72" pageNumber="73" stateProvince="Iasi">Dumnezeu</location>
,
<date id="9D3C102AAC11433309E2FA0E8D1D039E" box="[758,888,1477,1503]" pageId="72" pageNumber="73" value="2023-07-21">
<collectingDate id="8D78E9C2AC11433309E2FA0E8D1D039E" box="[758,888,1477,1503]" pageId="72" pageNumber="73" value="2023-07-21">21.vii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330891FA0E8DDD039E" box="[901,952,1477,1503]" count="10" pageId="72" pageNumber="73" type="male">10♂</specimenCount>
,
<specimenCount id="FF84FD63AC11433308D1FA0E8D8C03A1" box="[965,1001,1477,1504]" count="5" pageId="72" pageNumber="73" type="female">5♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330F3CFA0D8AF9039E" box="[1064,1180,1477,1504]" pageId="72" pageNumber="73">T.J. Wood</collectorName>
,
<collectionCode id="8F93AE2FAC1143330FBCFA0E8A90039E" box="[1192,1269,1477,1503]" pageId="72" pageNumber="73">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330E15FA0E8D260047" collectingDate="1972-07-28" collectionCode="TJWC" collectorName="M. Kocourek" country="Slovakia" location="Sturovo" pageId="72" pageNumber="73" specimenCount="9" specimenCount-female="9">
<emphasis id="DBF6EAF8AC1143330E15FA0E8BF9039E" bold="true" box="[1281,1436,1477,1503]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330E15FA0E8BF2039E" box="[1281,1431,1477,1503]" name="Slovakia" pageId="72" pageNumber="73">SLOVAKIA</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC1143330B83FA208E970047" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330B83FA208E970047" box="[151,242,1515,1542]" country="Slovakia" name="Sturovo" pageId="72" pageNumber="73">Štúrovo</location>
,
<date id="9D3C102AAC1143330A14FA228FE70042" box="[256,386,1513,1539]" pageId="72" pageNumber="73" value="1972-07-28">
<collectingDate id="8D78E9C2AC1143330A14FA228FE70042" box="[256,386,1513,1539]" pageId="72" pageNumber="73" value="1972-07-28">28.vii.1972</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330A84FA228FD10042" box="[400,436,1513,1539]" count="9" pageId="72" pageNumber="73" type="female">9♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330AE1FA218CEA0042" box="[501,655,1513,1540]" pageId="72" pageNumber="73">M. Kocourek</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FAC11433309EDFA208D260047" box="[761,835,1515,1542]" pageId="72" pageNumber="73">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330844FA228F48006B" collectingDate="1987-07-21" collectorName="Z. Padr" country="Slovakia" location="Remetske Hamre" municipality="Vihorlat" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="0959AC90AC1143330844FA228DCA0042" box="[848,943,1513,1539]" pageId="72" pageNumber="73">Vihorlat</collectingMunicipality>
,
<location id="EC5D6031AC11433308A8FA228AE70042" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC11433308A8FA228AE70042" box="[956,1154,1513,1539]" country="Slovakia" municipality="Vihorlat" name="Remetske Hamre" pageId="72" pageNumber="73">Remetské Hámre</location>
,
<date id="9D3C102AAC1143330F9BFA228B740042" box="[1167,1297,1513,1539]" pageId="72" pageNumber="73" value="1987-07-21">
<collectingDate id="8D78E9C2AC1143330F9BFA228B740042" box="[1167,1297,1513,1539]" pageId="72" pageNumber="73" value="1987-07-21">21.vii.1987</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330E0BFA228B270042" box="[1311,1346,1513,1539]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330E90FA218EA90066" pageId="72" pageNumber="73">Z. Pádr</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330BD3F9FA8A60009A" pageId="72" pageNumber="73" type="distribution">
<paragraph id="E93D36EAAC1143330BD3F9FA8A60009A" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330BD3F9FA8F04000A" bold="true" box="[199,353,1585,1611]" pageId="72" pageNumber="73">Distribution.</emphasis>
Not completely clear due to synonymisation, but across Pannonian Europe, including eastern
<collectingCountry id="9195767AAC1143330B83F99E8E89002E" box="[151,236,1621,1647]" name="Austria" pageId="72" pageNumber="73">Austria</collectingCountry>
,
<collectingCountry id="9195767AAC1143330A15F99E8F3B002E" box="[257,350,1621,1647]" name="Czech Republic" pageId="72" pageNumber="73">Czechia</collectingCountry>
,
<collectingCountry id="9195767AAC1143330A66F99D8FB00031" box="[370,469,1622,1648]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
, northern
<collectingCountry id="9195767AAC114333094CF99E8CC7002E" box="[600,674,1621,1647]" name="Serbia" pageId="72" pageNumber="73">Serbia</collectingCountry>
,
<collectingCountry id="9195767AAC11433309A3F99E8D79002E" box="[695,796,1621,1647]" name="Slovakia" pageId="72" pageNumber="73">Slovakia</collectingCountry>
,
<collectingCountry id="9195767AAC1143330824F99E8DFD002E" box="[816,920,1621,1647]" name="Romania" pageId="72" pageNumber="73">Romania</collectingCountry>
, and
<collectingCountry id="9195767AAC11433308F0F99E8A27002E" box="[996,1090,1621,1647]" name="Ukraine" pageId="72" pageNumber="73">Ukraine</collectingCountry>
. Warnckes distribution map (
<bibRefCitation id="8D134B1BAC1143330B8BF9B28F9100D2" author="Gusenleitner, F. &amp; Schwarz, M." box="[159,500,1657,1683]" pageId="72" pageNumber="73" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
) shows a predominantly Pannonian distribution focused on
<collectingCountry id="9195767AAC1143330F86F9B18A9000D5" box="[1170,1269,1658,1684]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
, with scattered records across
<collectingCountry id="9195767AAC1143330A29F9568FFF00F6" box="[317,410,1693,1719]" name="Ukraine" pageId="72" pageNumber="73">Ukraine</collectingCountry>
, one in the Caucasus, and some around the northern part of the Adriatic Sea (
<collectingCountry id="9195767AAC1143330FEDF9568B4A00F6" box="[1273,1327,1693,1719]" name="Italy" pageId="72" pageNumber="73">Italy</collectingCountry>
,
<collectingCountry id="9195767AAC1143330E2DF9568BF400F6" box="[1337,1425,1693,1719]" name="Croatia" pageId="72" pageNumber="73">Croatia</collectingCountry>
). The records from the Caucasus and Adriatic region must be revised for clarity.
</paragraph>
</subSubSection>
</treatment>
</document>