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<document id="0FD52BB9A2072273B8D922DF2D3FA481" ID-CLB-Dataset="26691" ID-DOI="http://dx.doi.org/10.3897/evolsyst.2.30778" ID-GBIF-Dataset="00106674-20d9-4d70-bb0f-8f000fb311f2" ID-Pensoft-Pub="2535-0730-2-169" ID-ZooBank="C0E89FEC8BE54DE9803D784FF6727BA0" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2535-0730-2-169" ModsDocOrigin="Evolutionary Systematics 2 (2)" ModsDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" checkinTime="1558621757476" checkinUser="pensoft" docAuthor="Framenau, Volker W. &amp; Baehr, Barbara C." docDate="2018" docId="C83CAEFD1667C94D0D5D77B5EF44039A" docLanguage="en" docName="EvolutSyst 2(2): 169-241" docOrigin="Evolutionary Systematics 2 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.2.30778" docTitle="Artoria extraordinaria Framenau &amp; Baehr, 2018, sp. n." docType="treatment" docUuid="9701DAC2-672B-4B8F-BC2A-E37F465485C1" docUuidSource="ZooBank" docVersion="6" lastPageNumber="169" masterDocId="FFE8FFB72743554D943BFFB83D324B46" masterDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" masterLastPageNumber="241" masterPageNumber="169" pageNumber="169" updateTime="1701381204780" updateUser="ExternalLinkService">
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<mods:title id="6C5D9838F3D035639AA6E4E3E6D9E4F5">The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)</mods:title>
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<mods:namePart id="41152D6E53402984FFF2E1F50905D693">Framenau, Volker W.</mods:namePart>
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<mods:namePart id="ADE509E6742E136CBB17DEC08461AA65">Baehr, Barbara C.</mods:namePart>
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<mods:title id="E15381469D0AFD1AF737B34DDE1F7242">Evolutionary Systematics</mods:title>
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<mods:date id="4929E410A84CD2EB597ED5738548A60A">2018</mods:date>
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<treatment id="C83CAEFD1667C94D0D5D77B5EF44039A" ID-GBIF-Taxon="157251891" LSID="urn:lsid:zoobank.org:act:9701DAC2-672B-4B8F-BC2A-E37F465485C1" httpUri="http://treatment.plazi.org/id/C83CAEFD1667C94D0D5D77B5EF44039A" lastPageNumber="169" pageId="0" pageNumber="169">
<subSubSection id="E0D05A42198B4F4D72794294A500D179" pageId="0" pageNumber="169" type="nomenclature">
<paragraph id="2175D8DE8368F8193105104535ACCADF" pageId="0" pageNumber="169">
<taxonomicName id="6DDF008E5E7A31E9C2E909EBE006C91D" ID-CoL="H2BM" LSID="http://zoobank.org/9701DAC2-672B-4B8F-BC2A-E37F465485C1" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria extraordinaria" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="extraordinaria">Artoria extraordinaria</taxonomicName>
<taxonomicNameLabel id="E32C2AB521246573902BB110EA1849FF" pageId="0" pageNumber="169">sp. n.</taxonomicNameLabel>
Figs 2E, 15, 18
<normalizedToken id="6AF96137FAB4BB2B9490F48150656457" originalValue="AH">A-H</normalizedToken>
, 47I Curious Forest Runner
</paragraph>
</subSubSection>
<subSubSection id="2E2EECF16B7658ABCD7F1DFCA83599EE" pageId="0" pageNumber="169" type="materials_examined">
<paragraph id="02B1241F6BB925507392E217718E1011" pageId="0" pageNumber="169">Material examined.</paragraph>
<paragraph id="A781A2B6FD2CCCEF8115BEF95E43C2C9" pageId="0" pageNumber="169">
Holotype male, Macquarie Road, 70 m S from junction with Milo Road, Buckenbowra State Forest (
<geoCoordinate id="58C3AF2D6DDF12D6D3E2B31416976205" direction="south" orientation="latitude" precision="925" value="-35.633335">35°38'S</geoCoordinate>
,
<geoCoordinate id="58A978F7F924B9BE50B7B334CBA6D093" direction="east" orientation="longitude" precision="925" value="149.88333">149°53'E</geoCoordinate>
, New South Wales, AUSTRALIA], 16 Mar 1999, R. Harris, H.M. Smith, leaf litter sample (AM KS128074). Paratype: 1 female, same data as holotype (AM KS68638).
</paragraph>
</subSubSection>
<subSubSection id="03890B44C5584A40076C8DE79E15B1F5" pageId="0" pageNumber="169" type="materials_examined">
<paragraph id="350A14A3F36E8A9206B997F2BAC33B30" pageId="0" pageNumber="169">Other material examined.</paragraph>
<paragraph id="DA2FD781379ABCD8C499689B82CABC1C" pageId="0" pageNumber="169">
10 males in 9 records (all NSW). AUSTRALIA: New South Wales: 1 male, 30 km SE of Braidwood, 250 m along Corn Trail Road from junction with Highway 54,
<geoCoordinate id="20E13A79F827D321A8153E7E34CA47FA" direction="south" orientation="latitude" precision="15" value="-35.561943">
35°33
<normalizedToken id="5C79937CE75E31A4F89C44F67850B2D1" originalValue="43”">'43&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="B7605B91615CCCA064847A98DA8910A5" direction="east" orientation="longitude" precision="15" value="150.0089">
150°00
<normalizedToken id="FEA939B0FB5D463617518C9A711F19D5" originalValue="32”">'32&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS68635); 1 male, Deua National Park, Dampier Mt Fire Trail, 3.5 km E of junction with Minuma Range Fire Trail,
<geoCoordinate id="FFFBBA5A8E0E2182066CE2365E9E2EDD" direction="south" orientation="latitude" precision="15" value="-35.986946">
35°59
<normalizedToken id="50FDEA387BAA31E0A593D797A2AD71E2" originalValue="13”">'13&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="9CD5078CD15D6C2A05FF208B921B99F2" direction="east" orientation="longitude" precision="15" value="149.7111">
149°42
<normalizedToken id="B3AAB924D9A38624F53E5B75E8C0370E" originalValue="40”">'40&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS68637); 1 male, Deua National Park, Minuma Range Fire Trail, approx.. 1.5 km ENE of apex of Dampier Mt,
<geoCoordinate id="0700382E16BB0E2D5FD4D8838D76C873" direction="south" orientation="latitude" precision="15" value="-35.98611">
35°59
<normalizedToken id="3539296004896D8324428C00B33F545D" originalValue="10”">'10&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="5458099F4CD33BF3C136F4D02F93FE79" direction="east" orientation="longitude" precision="15" value="149.6561">
149°39
<normalizedToken id="B4B82A761A8290A254CFE091F2DA7F0D" originalValue="22”">'22&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS64397); 2 males, Enfield State Forest, Dodds Fire Trail, 1 km from Enfield Road,
<geoCoordinate id="5F86D4EDA1E978EDA92C5DD6A8B80B0E" direction="south" orientation="latitude" precision="15" value="-31.396389">
31°23
<normalizedToken id="351F4CA4AA16C2FB8937C632B09B37F0" originalValue="47”">'47&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="47452E89D6AFD164132B325DF680619F" direction="east" orientation="longitude" precision="15" value="151.88306">
151°52
<normalizedToken id="16ED68478921842578849B91F5731FE5" originalValue="59”">'59&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS39804); 1 male, Enfield State Forest, Dodds Fire Trail, about 3 km from Enfield Road,
<geoCoordinate id="79D10F432C8C5EDA808CE02275B8BC88" direction="south" orientation="latitude" precision="15" value="-31.404722">
31°24
<normalizedToken id="2CF1359C8633A2B0F0BCD3C8E6B48C56" originalValue="17”">'17&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="C18E83A9B454E3F875A0B936EF36B5B0" direction="east" orientation="longitude" precision="15" value="151.88333">
151°53
<normalizedToken id="E15491CF796A8D939BE09CFC3C8588D6" originalValue="00”">'00&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS39803); 1 male, Monga State Forest, Lookout Road,
<geoCoordinate id="07E11EA9027D8D1ABF91E334C4A2023B" direction="south" orientation="latitude" precision="15" value="-35.57972">
35°34
<normalizedToken id="DFC96E6679B921363C9CA7CD7744B2D0" originalValue="47”">'47&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="C341AC978DBE226C17E0BD4922343E53" direction="east" orientation="longitude" precision="15" value="149.905">
149°54
<normalizedToken id="F9F9A7F73BADFD03B7ABEC9A78E39C45" originalValue="18”">'18&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS64399); 1 male, Monga State Forest, Northern Fire Trail, 1.85 km N from junction with McKeuns Road,
<geoCoordinate id="4578E37D3D263896FAA81A105A6375FD" direction="south" orientation="latitude" precision="15" value="-35.53611">
35°32
<normalizedToken id="3B23413DDE9DAE937EF08B9B6734A2A4" originalValue="10”">'10&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="5F190DC1EAAC2D03C48AEA45D08FD80A" direction="east" orientation="longitude" precision="15" value="149.88805">
149°53
<normalizedToken id="5FEDBF73AD513F71334124CCFC2866B5" originalValue="17”">'17&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS68634); 1 male, Monga State Forest, W side of Turtle Creek Fire Trail, 1.7 km from junction with Boundary Fire Trail,
<geoCoordinate id="0D29EC21DF29FAB66D54354917944018" direction="south" orientation="latitude" precision="15" value="-35.63889">
35°38
<normalizedToken id="276AA514971EF26D91E944B0A9085292" originalValue="20”">'20&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="26004C018CCF34B874587C78295E84A6" direction="east" orientation="longitude" precision="15" value="149.93112">
149°55
<normalizedToken id="325547DC66763D7F02DBC2E6319DDAC1" originalValue="52”">'52&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS64398); 1 male, Putty State Forest; approx. 80 m from Hunter Main Trail and just past '3
<normalizedToken id="0471E7CB04A69F29B2D8AC8AD4271218" originalValue="Ways">Ways'</normalizedToken>
,
<geoCoordinate id="3A5A0F0F0D0CB318D37BE65BDC64B643" direction="south" orientation="latitude" precision="15" value="-32.75722">
32°45
<normalizedToken id="F944B9C51B527ECF44C2CF8EB9F6A704" originalValue="26”">'26&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="1F9FA13A26EBBF7D0DF0CEA4FC23BCC6" direction="east" orientation="longitude" precision="15" value="150.5625">
150°33
<normalizedToken id="FEA86DF0127B7483E0076A02612C9376" originalValue="45”">'45&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS128874);
</paragraph>
</subSubSection>
<subSubSection id="AD6629FB729CF58795EE7EF3ADACE54A" pageId="0" pageNumber="169" type="etymology">
<paragraph id="149CED93EF494C5E50684736DA121438" pageId="0" pageNumber="169">Etymology.</paragraph>
<paragraph id="CA4CE1C1FE38FD002FB3E6094833D71D" pageId="0" pageNumber="169">
The specific epithet is an adjective in apposition derived from Latin (
<taxonomicName id="D0C405A23533A9D31F3038BDB0919736" lsidName="extraordinaria" pageId="0" pageNumber="169" rank="species" species="extraordinaria">extraordinaria</taxonomicName>
- unusual) and refers to the unusual shape of the basoembolic apophysis of the male pedipalp.
</paragraph>
</subSubSection>
<subSubSection id="0F36C5F5EECAF97B3CA8849C022821CC" pageId="0" pageNumber="169" type="diagnosis">
<paragraph id="9A823C99D196A901127D998EED93451B" pageId="0" pageNumber="169">Diagnosis.</paragraph>
<paragraph id="E3E1A8C0E160487FBF5792EBDFE83E7C" pageId="0" pageNumber="169">
The male pedipalp of
<taxonomicName id="BA5053BC8046AB6CCF05142245AC8D28" lsidName="A. extraordinaria" pageId="0" pageNumber="169" rank="species" species="extraordinaria">A. extraordinaria</taxonomicName>
sp. n. is highly unusual within the genus owing to the shape of the basoembolic apophysis, which forms an almost 360-degree circle (Fig. 47I). The female epigyne is poorly defined with two almost transparent lateral lobes (Fig. 18G).
</paragraph>
</subSubSection>
<subSubSection id="C4491DEB8124AD20E8BBFD7468E3A8F8" pageId="0" pageNumber="169" type="description">
<paragraph id="BBCF345A48AB6811F22A2A5C877ED0A8" pageId="0" pageNumber="169">Description.</paragraph>
<paragraph id="002F8BD772CBE799D02003A90C35BAFF" pageId="0" pageNumber="169">Male (based on holotype, AM KS1128074).</paragraph>
<paragraph id="AAC1F289F973B3A36E8907E3290A18F9" pageId="0" pageNumber="169">Total length 3.6.</paragraph>
<paragraph id="E8B4498CA9B5909A82B1A8F845B2C787" pageId="0" pageNumber="169">Prosoma. Length 2.1, width 1.4; carapace light reddish-brown with dark radial pattern and black V-shaped pattern between cephalic and thoracic region; indistinct and irregular broad lighter central and marginal band (Fig. 18A); sternum pale, with darker margin (Fig. 18B).</paragraph>
<paragraph id="7DC9BDC2AE5E409AA5F99E9867C278EC" pageId="0" pageNumber="169">Eyes (Fig. 2E). Diameter of AME: 0.10; ALE: 0.09; PME: 0.30; PLE: 0.20.</paragraph>
<paragraph id="BC4F1AFE2F043030D852C96249DBA826" pageId="0" pageNumber="169">Anterior eye row. Strongly procurved, distance between AME/ALE at least twice AME/AME.</paragraph>
<paragraph id="A748767B08801BB9BF5583EFC6BA8372" pageId="0" pageNumber="169">Chelicerae. Dark brown.</paragraph>
<paragraph id="33CAADB0E3E9FA4437257EA47D50BAE2" pageId="0" pageNumber="169">Labium. Dark brown, with lighter anterior rim (Fig. 18B)</paragraph>
<paragraph id="DE93766B30F2BB1C09F88364D2BA6006" pageId="0" pageNumber="169">Pedipalp coxae. Dark brown, with lighter anterior rim (Fig. 18B).</paragraph>
<paragraph id="0ACCFD4EC8CDA2D4E356BE10B73D43AD" pageId="0" pageNumber="169">Legs. Yellow brown with darker annulations; metatarsi and tarsi darker, less annulated (Fig. 18A).</paragraph>
<paragraph id="DF064F983E7754F2D40D66D8D32B7EAF" pageId="0" pageNumber="169">Opisthosoma. Length 1.5, width 1.1; dark grey with pale anterior cardiac mark and light irregular pattern (Fig. 18A). Venter pale with darker pattern (Fig. 18B); spinnerets pale.</paragraph>
<paragraph id="95640D44217CD72CFBDDEB6A00C212DE" pageId="0" pageNumber="169">Pedipalps. Tibia as long as broad; cymbium tip with cluster of macrosetae (Fig. 18E, F); dorsal scopula patch present; tegular apophysis spatulate with two lateral appendices, basally narrowed to 2/3 width, retrolateral tip pointed inwards not reaching margin of cymbium (Fig. 18E); palea about 1 1/2 times as long as wide; basoembolic apophysis small, rounded; embolus elongated coiled with long thin tip, reaching beyond terminal apophysis; terminal apophysis small with pointed tip (Fig. 47I).</paragraph>
<paragraph id="7404207CB82D1B352DD2F3E937FFD0F6" pageId="0" pageNumber="169">Female (based on AM KS68638).</paragraph>
<paragraph id="E28F775DF64DDC23A453E0C1515FF0AF" pageId="0" pageNumber="169">Total length 3.8.</paragraph>
<paragraph id="5208FD17B530DC6619E89E4C449D07DF" pageId="0" pageNumber="169">Prosoma. Length 2.0, width 1.4; carapace and sternum colouration as male (Fig. 18C).</paragraph>
<paragraph id="EBC84F58724DD154DBB3EB49C85B4DB0" pageId="0" pageNumber="169">Eyes. Diameter of AME 0.08, ALE 0.09, PME 0.30, PLE 0.21.</paragraph>
<paragraph id="3CAEED0F19488946AA9A6F2977C856E7" pageId="0" pageNumber="169">Anterior eye row. Strongly procurved, distance between AME/ALE at least twice AME/AME.</paragraph>
<paragraph id="518DDB849E6A195480293D32E331B847" pageId="0" pageNumber="169">Opisthosoma. Length 1.8, width 1.2; otherwise as male, but opisthosoma pattern more obscure (Fig. 18C, D).</paragraph>
<paragraph id="F9BBFEC36E931F918C26A171308C3BFC" pageId="0" pageNumber="169">Epigyne. About as long as wide, poorly sclerotised at posterior tips, atrium lighter (Fig. 18G); spermathecal heads ellipsoid in inverted v-shaped position about 1/2 short diameter apart, spermathecal stalks attached ventrally, globular at base (Fig. 18H).</paragraph>
</subSubSection>
<subSubSection id="24ECA25BF99D66F2FFDA548630F79728" pageId="0" pageNumber="169" type="life history and habitat preferences">
<paragraph id="9128EF6DDC2F79B77C0A733E1026CC75" pageId="0" pageNumber="169">Life history and habitat preferences.</paragraph>
<paragraph id="3E0B4D1A6E11EF6F36AED70302CBA8CD" pageId="0" pageNumber="169">
Based on locality data,
<taxonomicName id="5A2BE836B9E183952D4CAC2698EF0710" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria extraordinaria" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="extraordinaria">Artoria extraordinaria</taxonomicName>
sp. n. is a forest species. Detailed habitat data were not available on locality labels of the specimens. Most specimens were collected between February and April, with one male recorded from June, suggesting
<taxonomicName id="67DFC3B9AD6F4296CBEED5034EB57515" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria extraordinaria" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="extraordinaria">Artoria extraordinaria</taxonomicName>
sp. n. to be autumn to winter mature.
</paragraph>
</subSubSection>
<subSubSection id="6952C12D84155AECD0916D5127AF9971" pageId="0" pageNumber="169" type="distribution">
<paragraph id="A9AE4E7D55367D3CB16FEC66EB1D53B1" pageId="0" pageNumber="169">Distribution.</paragraph>
<paragraph id="32085775E5530467C24DC2DE14D5F900" pageId="0" pageNumber="169">This species was found some distance from the coast at scattered localities in the NSW North Coast (NNC), Sydney Basin (SYB) and South Easter Corner (SEC) IBRA bioregions (Fig. 15).</paragraph>
<caption id="11F9E44F782712272B865DC45BE05E12" pageId="0" pageNumber="169">
<paragraph id="5B66AF5965ADD1D85BA15044CF853C76" pageId="0" pageNumber="169">
Figure 18.
<taxonomicName id="7A29974263DC076C2F6C0C0C3F211E30" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria extraordinaria" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="extraordinaria">Artoria extraordinaria</taxonomicName>
sp. n., male holotype (AM KS128074), female paratype (AM KS68638): A, male habitus, dorsal view; B, male habitus, ventral view; C, female habitus, dorsal view; D, female habitus, ventral view; E, male pedipalp, ventral view; F, male pedipalp, retrolateral view; G, epigyne, ventral view; H, epigyne, dorsal view. Scale bars: habitus 1.0 mm; pedipalp, epigyne 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>