228 lines
26 KiB
XML
228 lines
26 KiB
XML
<document id="28D6E4A34EC312775AE9F04232596DF0" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia & Ahrens, Dirk" docDate="2017" docId="03CB6967FF9138413B91FE2C9538FEAC" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla stalsi AHRENS, BECKETT, EBERLE & FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="342" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="342" updateTime="1738712560284" updateUser="ExternalLinkService">
|
||
<mods:mods id="F2C7B524409584A1F220976C8287DFFE" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1FA170C4BD2578A9CAA1324CE589AB4A">
|
||
<mods:title id="8E7D6CBF62E405013496F038E219DD73">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="99BAEF0C3676F57DFC23E4417BF3368B" type="personal">
|
||
<mods:role id="052BC1386F49F401E4C84CB6CFF4F5CD">
|
||
<mods:roleTerm id="C81C028A0DA81DE726FE64547E60B21B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D9B26BC1B9ED8443974A73A464A92D8C">Eberle, Jonas</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="91B36116E9E1E97FFD53673F3952EBA2" type="personal">
|
||
<mods:role id="C2C3616C2E712B05A8CD460095BCF1FA">
|
||
<mods:roleTerm id="6E969BBCD94629D981348072DB0B8388">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D7B2B631DEE7B164A4B338E1286AD6AD">Beckett, Marc</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="BFB21C8FF766CFE329B151EA4A57BEC1" type="personal">
|
||
<mods:role id="D4377DC984E05018BDDA477D23C501D5">
|
||
<mods:roleTerm id="A7E77ADFB42D884DACA3949D93359BD7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="182CEB7AA1927EBD4E9CAE513F208048">Ozgul-Siemund, Alev</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="AB9E86D0086E77EE021A6A9D064B843C" type="personal">
|
||
<mods:role id="483731C32A065F9B94FC006E09D7FA56">
|
||
<mods:roleTerm id="9BCFAD572509212F7695ABD2683832E1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B46DE2E5EC69F0F5201467E9C95F825E">Frings, Jujina</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="917C959F699ABB7DE0DA1E5FCDB3A908" type="personal">
|
||
<mods:role id="E9E6D35E1B646CDAA3A7226D4AB8762C">
|
||
<mods:roleTerm id="FEA318C8AFC385B9508CCBA1017D4A67">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1449BF3AAC94807726F98AF42426AC13">Fabrizi, Silvia</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="199DF5153806DCB2CA76E2E4912F7985" type="personal">
|
||
<mods:role id="14443350770A43F9C3C641A7244664D5">
|
||
<mods:roleTerm id="27F9DD76C5371EA2E784DAA7DDD04D60">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="70047B633EE81BE26A8242D6EB76BB42">Ahrens, Dirk</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="6DC6C67AB37FA4A96B76C193CD4F3956">text</mods:typeOfResource>
|
||
<mods:relatedItem id="829696201B10ECED60BFFE24EC8E8AAA" type="host">
|
||
<mods:titleInfo id="C65191A717F15B552844F011F9BEC050">
|
||
<mods:title id="8F41700DE1DF4B796ECABECB51EF0507">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2B59782D35BBC328C3338B827B718C32">
|
||
<mods:date id="9DD4E64A8A0E8F69F181F60883E971F4">2017</mods:date>
|
||
<mods:detail id="AFBAD609D5E2622230A58C21AAEB8293" type="series">
|
||
<mods:title id="44E7524BAC91F59A455738AD4D6BCF5D">Zool. J. Linn. Soc.</mods:title>
|
||
</mods:detail>
|
||
<mods:detail id="B28336814E98EF52FB98367F779AAFB3" type="volume">
|
||
<mods:number id="C7109F60949B37D03F8E00D2DC154CFF">180</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4F86C30F4721C16BADDB39AF307220B6" type="issue">
|
||
<mods:number id="280D2B378AAF442DAAA884B21893DBA5">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="525F75A285524958EDF6E30BDB4F062A" unit="page">
|
||
<mods:start id="1EE0F22E47C2FD2AFB0433A114504929">321</mods:start>
|
||
<mods:end id="30E9AF55519FA1DC27C418A87839B7C5">353</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="09E3C897BD7C2B15B0BBAA31897EA23D">
|
||
<mods:url id="D22814763EBE7D2ED4C80B62CD707273">https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="49C9A1DAA07E08131BA52E73AA70D2F6">journal article</mods:classification>
|
||
<mods:identifier id="26ABCF0461E3F99CB92BD1969950E99F" type="DOI">10.1111/zoj.12489</mods:identifier>
|
||
<mods:identifier id="A89D267A61C90FA4769A0562BD48BF15" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="D75D988048862638AA7D5B79471004C4" type="Zenodo-Dep">14807083</mods:identifier>
|
||
<mods:identifier id="B1A91A058432FAAE8EE5326B40FE4B20" type="ZooBank">3663E0A-2E17-4346-A388-6BEE53081814</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03CB6967FF9138413B91FE2C9538FEAC" LSID="urn:lsid:plazi:treatment:03CB6967FF9138413B91FE2C9538FEAC" httpUri="http://treatment.plazi.org/id/03CB6967FF9138413B91FE2C9538FEAC" lastPageNumber="342" pageId="21" pageNumber="342">
|
||
<subSubSection id="C3788BFAFF9138413B91FE2C902EFD9E" pageId="21" pageNumber="342" type="nomenclature">
|
||
<paragraph id="8BDDD871FF9138413B91FE2C93D0FE79" blockId="21.[173,774,407,464]" pageId="21" pageNumber="342">
|
||
<heading id="D0956F1DFF9138413B91FE2C93D0FE79" centered="true" fontSize="9" level="2" pageId="21" pageNumber="342" reason="2">
|
||
<taxonomicName id="4C62A3F2FF9138413B91FE2C9032FE78" ID-CoL="4K84G" authority="AHRENS, BECKETT, EBERLE & FABRIZI" authorityName="AHRENS, BECKETT, EBERLE & FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
|
||
<emphasis id="B9160463FF9138413B91FE2C90F9FE19" bold="true" box="[173,415,411,432]" italics="true" pageId="21" pageNumber="342">
|
||
<smallCapsWord id="8D3B4EADFF9138413B91FE2C9021FE19" baselines="427,427" box="[173,327,411,432]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="21" pageNumber="342">PLEOPHYLLA</smallCapsWord>
|
||
<smallCapsWord id="8D3B4EADFF9138413A6CFE2990F9FE19" baselines="427" box="[336,415,414,431]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="stalsi" pageId="21" pageNumber="342">STALSI</smallCapsWord>
|
||
</emphasis>
|
||
<smallCapsWord id="8D3B4EADFF9138413A94FE2E9362FE06" baselines="427,428" box="[424,516,409,433]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="21" pageNumber="342">AHRENS</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="8D3B4EADFF9138413929FE2E931AFE06" baselines="427,428" box="[533,636,409,433]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="21" pageNumber="342">BECKETT</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="8D3B4EADFF91384139B1FE2E9380FE19" baselines="427,427" box="[653,742,409,433]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Eberle" pageId="21" pageNumber="342">EBERLE</smallCapsWord>
|
||
&
|
||
<smallCapsWord id="8D3B4EADFF9138413BC0FE009032FE78" baselines="457,458" box="[252,340,439,463]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="21" pageNumber="342">FABRIZI</smallCapsWord>
|
||
</taxonomicName>
|
||
<emphasis id="B9160463FF9138413A62FE0A90A5FE66" bold="true" box="[350,451,437,464]" pageId="21" pageNumber="342">
|
||
<taxonomicNameLabel id="A225B918FF9138413A62FE0A90A5FE66" box="[350,451,437,464]" pageId="21" pageNumber="342" rank="species">
|
||
<smallCapsWord id="8D3B4EADFF9138413A62FE0A901CFE78" baselines="458" box="[350,378,445,462]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="21" pageNumber="342">SP</smallCapsWord>
|
||
.
|
||
<smallCapsWord id="8D3B4EADFF9138413AB7FE0A90DDFE78" baselines="458" box="[395,443,445,462]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="21" pageNumber="342">NOV</smallCapsWord>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1359C4F4FF9138413AEAFE00933EFE79" box="[470,600,437,463]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">
|
||
<smallCapsWord id="8D3B4EADFF9138413AEAFE00936DFE78" baselines="457,458" box="[470,523,439,463]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="21" pageNumber="342">FIGS</smallCapsWord>
|
||
4Q
|
||
<emphasis id="B9160463FF9138413905FE0E9321FE79" box="[569,583,441,463]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||
T
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="1359C4F4FF9138413955FE0293EBFE79" box="[617,653,437,463]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="21" pageNumber="342">5O</figureCitation>
|
||
, 6)
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF9138413B9FFE43902EFD9E" blockId="21.[163,778,500,552]" pageId="21" pageNumber="342">
|
||
<uri id="FFF3D473FF9138413B9FFE43902EFD9E" pageId="21" pageNumber="342">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFC4E2A4FF9138413A97FE43902EFD9E" pageId="21" pageNumber="342">0F7D7614-7CFD-44C3-9AAC-398093991B09</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3788BFAFF9138413B9AFDF29538FEAC" pageId="21" pageNumber="342" type="materials_examined">
|
||
<paragraph id="8BDDD871FF9138413B9AFDF29382FD65" blockId="21.[166,782,580,724]" pageId="21" pageNumber="342">
|
||
<emphasis id="B9160463FF9138413B9AFDF290AFFDEF" box="[166,457,580,602]" italics="true" pageId="21" pageNumber="342">Type material examined:</emphasis>
|
||
<materialsCitation id="3B0AD22CFF9138413AE4FDF390FFFD65" collectingDate="2005-11-12" collectionCode="SANC" collectorName="W. Breytenbach" country="South Africa" elevation="1639" latitude="-24.933332" location="Graskop" longLatPrecision="503" longitude="30.47" municipality="Pilgrimsrest" pageId="21" pageNumber="342" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<typeStatus id="54D966D3FF9138413AE4FDF39327FDEC" box="[472,577,580,602]" pageId="21" pageNumber="342" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCount id="9D6413F8FF9138413973FDF09339FDEF" box="[591,607,583,601]" count="1" pageId="21" pageNumber="342" type="male">♂</specimenCount>
|
||
“
|
||
<collectingCountry id="F37598E1FF9138413948FDF3926CFDEC" box="[628,778,580,602]" name="South Africa" pageId="21" pageNumber="342">South Africa</collectingCountry>
|
||
: MPU
|
||
<location id="8EBD8EAAFF9138413BCAFDD49030FDCF" LSID="urn:lsid:plazi:treatment:03CB6967FF9138413B91FE2C9538FEAC:8EBD8EAAFF9138413BCAFDD49030FDCF" box="[246,342,611,633]" country="South Africa" latitude="-24.933332" longLatPrecision="503" longitude="30.47" municipality="Pilgrimsrest" name="Graskop" pageId="21" pageNumber="342">Graskop</location>
|
||
<quantity id="4C9A7594FF9138413A56FDD590C3FDCF" box="[362,421,610,633]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="342" unit="km" value="5.0">5 km</quantity>
|
||
on
|
||
<collectingMunicipality id="6BB9420BFF9138413AD5FDD49310FDCF" box="[489,630,611,633]" pageId="21" pageNumber="342">Pilgrimsrest</collectingMunicipality>
|
||
rd
|
||
<geoCoordinate id="EE56BEB6FF9138413984FDD4926AFDCE" box="[696,780,606,632]" degrees="24" direction="south" minutes="56" orientation="latitude" pageId="21" pageNumber="342" precision="15" seconds="0" value="-24.933332">
|
||
24
|
||
<emphasis id="B9160463FF91384139EFFDD393BBFDCE" box="[723,733,612,632]" italics="true" pageId="21" pageNumber="342">°</emphasis>
|
||
56
|
||
<emphasis id="B9160463FF91384139C4FDE9939BFDDB" bold="true" box="[760,765,606,621]" italics="true" pageId="21" pageNumber="342">
|
||
<superScript id="7C177539FF91384139C4FDE9939BFDDB" attach="right" box="[760,765,606,621]" fontSize="6" pageId="21" pageNumber="342">0</superScript>
|
||
</emphasis>
|
||
S
|
||
</geoCoordinate>
|
||
<geoCoordinate id="EE56BEB6FF9138413B9AFD369193FD20" box="[166,245,641,662]" degrees="30.47" direction="east" orientation="longitude" pageId="21" pageNumber="342" precision="555" value="30.47">30.47E</geoCoordinate>
|
||
<date id="FFDCFEB1FF9138413A26FD3690F2FD21" box="[282,404,641,663]" pageId="21" pageNumber="342" value="2005-11-12">
|
||
<collectingDate id="EF980759FF9138413A26FD3690F2FD21" box="[282,404,641,663]" pageId="21" pageNumber="342" value="2005-11-12">12.XI.2005</collectingDate>
|
||
</date>
|
||
<quantity id="4C9A7594FF9138413A85FD369368FD21" box="[441,526,641,663]" metricMagnitude="3" metricUnit="m" metricValue="1.639" pageId="21" pageNumber="342" unit="m" value="1639.0">
|
||
<elevation id="004F3F42FF9138413A85FD369368FD21" box="[441,526,641,663]" metricMagnitude="3" metricUnit="m" metricValue="1.639" pageId="21" pageNumber="342" unit="m" value="1639.0">1639 m</elevation>
|
||
</quantity>
|
||
<collectorName id="2697BDA7FF913841390EFD359263FD21" box="[562,773,641,663]" pageId="21" pageNumber="342">W. Breytenbach</collectorName>
|
||
/ Collected from
|
||
<taxonomicName id="4C62A3F2FF9138413A6EFD179083FD03" box="[338,485,672,693]" class="Magnoliopsida" family="Proteaceae" genus="Protea" kingdom="Plantae" order="Proteales" pageId="21" pageNumber="342" phylum="Tracheophyta" rank="species" species="caffra">
|
||
<emphasis id="B9160463FF9138413A6EFD179083FD03" box="[338,485,672,693]" italics="true" pageId="21" pageNumber="342">Protea caffra</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C62A3F2FF9138413AC8FD179314FD03" box="[500,626,672,693]" class="Magnoliopsida" family="Proteaceae" kingdom="Plantae" order="Proteales" pageId="21" pageNumber="342" phylum="Tracheophyta" rank="family">Proteaceae</taxonomicName>
|
||
) many leaves were beaten” (
|
||
<collectionCode id="ED7340B4FF9138413A7BFD0990F5FD65" box="[327,403,702,723]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="21" pageNumber="342" type="Museum">SANC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54D966D3FF9138413A9FFD08937FFD62" box="[419,537,703,724]" pageId="21" pageNumber="342" type="paratype">Paratypes</typeStatus>
|
||
: see Appendix S1.
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF9138413B9AFD5893FAFA75" blockId="21.[166,781,750,1475]" pageId="21" pageNumber="342">
|
||
<emphasis id="B9160463FF9138413B9AFD589049FCB2" box="[166,303,751,772]" italics="true" pageId="21" pageNumber="342">Description:</emphasis>
|
||
Body length:
|
||
<quantity id="4C9A7594FF9138413AD2FD599321FCB2" box="[494,583,750,772]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="21" pageNumber="342" unit="mm" value="9.9">9.9 mm</quantity>
|
||
, elytral length:
|
||
<quantity id="4C9A7594FF9138413B9AFCBA9199FC94" box="[166,255,781,803]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="21" pageNumber="342" unit="mm" value="6.7">6.7 mm</quantity>
|
||
, maximum width:
|
||
<quantity id="4C9A7594FF9138413AD3FCBB932FFC94" box="[495,585,780,802]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="21" pageNumber="342" unit="mm" value="5.1">5.1 mm</quantity>
|
||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.64. Pronotum apparently glabrous (with only a few erect setae anteriorly, otherwise with only minute setae); unicoloured; basal margin of pronotum complete; larger punctures absent from pronotum. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra sparse (fewer than ten setae per interval); sutural interval in apical declivity of elytra distinctly elevated but convex; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity short, less than 1/4 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/ 1.38. Metatibia, ratio of width/length: 1/3.94. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||
<figureCitation id="1359C4F4FF91384139A8FA39926FFA12" box="[660,777,1422,1444]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">
|
||
Fig. 4Q
|
||
<emphasis id="B9160463FF91384139D6FA269390FA15" box="[746,758,1425,1443]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||
S
|
||
</figureCitation>
|
||
. Habitus:
|
||
<figureCitation id="1359C4F4FF9138413A2CFA1A9000FA74" box="[272,358,1453,1475]" captionStart="Figure 4" captionStartId="9.[166,245,1676,1695]" captionTargetBox="[189,1421,218,1624]" captionTargetId="figure-153@9.[179,1427,195,1653]" captionTargetPageId="9" captionText="Figure 4. A–D, Pleophylla mpumalanga sp. nov. (holotype); E–H, Pleophylla charlyi sp. nov. (holotype); I–L, Pleophylla lizleri sp. nov. (holotype); M–P, Pleophylla taitaensis sp. nov. (holotype); Q–T, Pleophylla stalsi sp. nov. (holotype); U–X, Pleophylla wakkerstroomensis sp. nov. (holotype); Y–Ab, Pleophylla tongaatsana Peringuey (RSA, Sjonajona); Ac–Af, Pleophylla kruegeri sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="21" pageNumber="342">Fig. 4T</figureCitation>
|
||
. Female genitalia:
|
||
<figureCitation id="1359C4F4FF9138413903FA1A93FEFA75" box="[575,664,1453,1475]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="21" pageNumber="342">Fig. 5O</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF9138413B9AFA6B90AFF9FA" blockId="21.[166,781,1499,1613]" pageId="21" pageNumber="342">
|
||
<emphasis id="B9160463FF9138413B9AFA6B908AFA46" box="[166,492,1499,1521]" italics="true" pageId="21" pageNumber="342">
|
||
Diagnosis:
|
||
<taxonomicName id="4C62A3F2FF9138413A10FA6C908AFA46" authorityName="Eberle & Beckett & Ozgul-Siemund & Frings & Fabrizi & Ahrens" authorityYear="2017" box="[300,492,1499,1520]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="342" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">Pleophylla stalsi</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A225B918FF9138413ACBFA6C9334FA46" box="[503,594,1499,1520]" pageId="21" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from all other
|
||
<taxonomicName id="4C62A3F2FF9138413BD3FA4D90A4F9A6" box="[239,450,1530,1552]" pageId="21" pageNumber="342">
|
||
<emphasis id="B9160463FF9138413BD3FA4D9001F9B9" box="[239,359,1530,1551]" italics="true" pageId="21" pageNumber="342">Pleophylla</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the short white pilosity on the dorsal surface, and the almost absent erect pilosity on the pronotum.
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF9138413B9AF9D49372F8DB" blockId="21.[166,781,1635,1901]" pageId="21" pageNumber="342">
|
||
<emphasis id="B9160463FF9138413B9AF9D4907CF9CE" box="[166,282,1635,1656]" italics="true" pageId="21" pageNumber="342">Variation:</emphasis>
|
||
Body length:
|
||
<quantity id="4C9A7594FF9138413AF9F9D4933CF9CE" box="[453,602,1635,1656]" metricMagnitude="-2" metricUnit="m" metricValue="1.015" metricValueMax="1.04" metricValueMin="0.99" pageId="21" pageNumber="342" unit="mm" value="10.15" valueMax="10.4" valueMin="9.9">
|
||
9.9
|
||
<emphasis id="B9160463FF9138413ADBF9D19095F9CE" box="[487,499,1638,1656]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||
10.4 mm
|
||
</quantity>
|
||
, elytral length:
|
||
<quantity id="4C9A7594FF9138413B9AF9369048F921" box="[166,302,1665,1687]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="21" pageNumber="342" unit="mm" value="6.65" valueMax="6.7" valueMin="6.6">
|
||
6.6
|
||
<emphasis id="B9160463FF9138413BF4F93391B2F920" box="[200,212,1668,1686]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||
6.7 mm
|
||
</quantity>
|
||
, maximum width:
|
||
<quantity id="4C9A7594FF9138413926F93693C4F921" box="[538,674,1665,1687]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="21" pageNumber="342" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">
|
||
4.6
|
||
<emphasis id="B9160463FF9138413900F933932EF920" box="[572,584,1668,1686]" italics="true" pageId="21" pageNumber="342">–</emphasis>
|
||
5.1 mm
|
||
</quantity>
|
||
. Density of dark spots may vary, often body reddish brown and completely without spots. Female: antennal club of female straight, composed of five antennomeres, as long as the remaining antennomeres combined; sixth antennomere shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.5.
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF9138413801FF709538FEAC" blockId="21.[829,1444,199,282]" pageId="21" pageNumber="342">
|
||
<emphasis id="B9160463FF9138413801FF7092DDFF6A" box="[829,955,199,220]" italics="true" pageId="21" pageNumber="342">Etymology:</emphasis>
|
||
This new species is dedicated to Riaan Stals (
|
||
<collectionCode id="ED7340B4FF91384138B2FF5192B1FF4D" box="[910,983,230,251]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="21" pageNumber="342" type="Museum">SANC</collectionCode>
|
||
) in gratitude for his support of our research in
|
||
<collectingCountry id="F37598E1FF91384138FAFEB3953CFEAC" box="[966,1114,260,282]" name="South Africa" pageId="21" pageNumber="342">South Africa</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |