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<document id="A516B76B4AFAC36A623B03C5B6ECE054" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia &amp; Ahrens, Dirk" docDate="2017" docId="03CB6967FF8E38583805FB2E906BFD5A" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla mlilwaneensis AHRENS, BECKETT, EBERLE &amp; FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="333" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="331" updateTime="1738712560284" updateUser="ExternalLinkService">
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<mods:title id="0887F557ACB5BDDF72C8A90AFE460AE4">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="72CCBA9C4BE30A47F9A8393E4D1CF5C9">Eberle, Jonas</mods:namePart>
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<taxonomicName id="4C62A3F2FF8E385E3805FB2E9554FB7A" ID-CoL="4K845" authority="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityName="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="mlilwaneensis" status="sp. nov.">
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,
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,
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&amp;
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.
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.
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(
<figureCitation id="1359C4F4FF8E385E3F8FFB019453FB78" box="[1203,1333,1204,1230]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. AD, Pleophylla fasciatipennis Blanchard (RSA, George); EH, Pleophylla pilosa Boheman (RSA, Karkloof for.); IL, Pleophylla pseudopilosa sp. nov. (holotype); MP, Pleophylla nelshoogteensis sp. nov. (holotype); Q T, Pleophylla mlilwaneensis sp. nov. (holotype); UX, Pleophylla burundiensis sp. nov. (holotype); YAb, Pleophylla congoensis sp. nov. (holotype); AcAf, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
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2Q
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T
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, 6)
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urn:lsid:zoobank.org:act:
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<emphasis id="B9160463FF8E385E3815FAF0953AFAED" box="[809,1116,1350,1372]" italics="true" pageId="10" pageNumber="331">Type material examined:</emphasis>
<materialsCitation id="3B0AD22CFF8E385E3F56FAF194E1FA2E" collectingDate="1996-11-17" collectingDateMax="1996-11-18" collectingDateMin="1996-11-17" collectionCode="NME" collectorName="M. Hartmann" country="Swaziland" latitude="-22.0" location="Mlilwane Wildlife Sanctuary" longLatPrecision="925" longitude="31.183332" pageId="10" pageNumber="331" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<collectingCountry id="F37598E1FF8E385E3E2FFAF194EDFAEA" box="[1299,1419,1350,1372]" name="Swaziland" pageId="10" pageNumber="331">Swaziland</collectingCountry>
,
<location id="8EBD8EAAFF8E385E3815FAD295EBFACC" LSID="urn:lsid:plazi:treatment:03CB6967FF8E38583805FB2E906BFD5A:8EBD8EAAFF8E385E3815FAD295EBFACC" box="[809,1165,1381,1403]" country="Swaziland" latitude="-22.0" longLatPrecision="925" longitude="31.183332" name="Mlilwane Wildlife Sanctuary" pageId="10" pageNumber="331">Mlilwane Wildlife Sanctuary</location>
26
<emphasis id="B9160463FF8E385E3F83FAD195AFFACC" box="[1215,1225,1382,1402]" italics="true" pageId="10" pageNumber="331">°</emphasis>
29,
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22
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S
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,
<geoCoordinate id="EE56BEB6FF8E385E3E04FAD294F6FACC" box="[1336,1424,1376,1402]" degrees="31" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="331" precision="15" seconds="0" value="31.183332">
31
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11
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E
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800mNN,
<date id="FFDCFEB1FF8E385E38A2FA349520FA2F" box="[926,1094,1411,1433]" pageId="10" pageNumber="331" value="1996-11-17" valueMax="1996-11-18" valueMin="1996-11-17">
<collectingDate id="EF980759FF8E385E38A2FA349520FA2F" box="[926,1094,1411,1433]" pageId="10" pageNumber="331" value="1996-11-17" valueMax="1996-11-18" valueMin="1996-11-17">17.-18.XI.1996</collectingDate>
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, leg.
<collectorName id="2697BDA7FF8E385E3FBFFA339446FA2E" box="[1155,1312,1411,1433]" pageId="10" pageNumber="331">M. Hartmann</collectorName>
” (
<collectionCode id="ED7340B4FF8E385E3E06FA34941AFA2E" box="[1338,1404,1411,1432]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="10" pageNumber="331">NME</collectionCode>
).
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</paragraph>
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<emphasis id="B9160463FF8E385E3815FA7692D7FA60" box="[809,945,1473,1494]" italics="true" pageId="10" pageNumber="331">Description:</emphasis>
Body length:
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, elytral length:
<quantity id="4C9A7594FF8E385E3815FA6892E4FA42" box="[809,898,1503,1525]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="331" unit="mm" value="6.8">6.8 mm</quantity>
, maximum width: 4.0 mm. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.74. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum moderately large. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.44. Metatibia, ratio of width/length: 1/3.71. Metatarsi dorsally smooth; first metatarsomere slightly shorter than subsequent one. Aedeagus:
<figureCitation id="1359C4F4FF8838583A37FE2A90E6FE04" box="[267,384,413,435]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. AD, Pleophylla fasciatipennis Blanchard (RSA, George); EH, Pleophylla pilosa Boheman (RSA, Karkloof for.); IL, Pleophylla pseudopilosa sp. nov. (holotype); MP, Pleophylla nelshoogteensis sp. nov. (holotype); Q T, Pleophylla mlilwaneensis sp. nov. (holotype); UX, Pleophylla burundiensis sp. nov. (holotype); YAb, Pleophylla congoensis sp. nov. (holotype); AcAf, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">
Fig. 2Q
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S
</figureCitation>
. Habitus:
<figureCitation id="1359C4F4FF8838583AC9FE2A932AFE04" box="[501,588,413,435]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. AD, Pleophylla fasciatipennis Blanchard (RSA, George); EH, Pleophylla pilosa Boheman (RSA, Karkloof for.); IL, Pleophylla pseudopilosa sp. nov. (holotype); MP, Pleophylla nelshoogteensis sp. nov. (holotype); Q T, Pleophylla mlilwaneensis sp. nov. (holotype); UX, Pleophylla burundiensis sp. nov. (holotype); YAb, Pleophylla congoensis sp. nov. (holotype); AcAf, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="12" pageNumber="333">Fig. 2T</figureCitation>
.
</paragraph>
<caption id="DF1D88F9FF8F385F3B9AF9029473F888" ID-DOI="http://doi.org/10.5281/zenodo.14807095" ID-Zenodo-Dep="14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="11" pageNumber="332" startId="11.[166,245,1717,1736]" targetBox="[237,1338,218,1653]" targetPageId="11" targetType="figure">
<paragraph id="8BDDD871FF8F385F3B9AF9029473F888" blockId="11.[166,1444,1717,1854]" pageId="11" pageNumber="332">
<emphasis id="B9160463FF8F385F3B9AF9029076F97F" bold="true" box="[166,272,1717,1737]" pageId="11" pageNumber="332">Figure 5.</emphasis>
Female genitalia of
<taxonomicName id="4C62A3F2FF8F385F3AC2F9019381F97F" box="[510,743,1718,1737]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B9160463FF8F385F3AC2F901930AF97F" box="[510,620,1718,1737]" italics="true" pageId="11" pageNumber="332">Pleophylla</emphasis>
species. A
</taxonomicName>
,
<taxonomicName id="4C62A3F2FF8F385F39CBF9019565F97F" box="[759,1027,1718,1737]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="fasciatipennis">
<emphasis id="B9160463FF8F385F39CBF9019565F97F" box="[759,1027,1718,1737]" italics="true" pageId="11" pageNumber="332">Pleophylla fasciatipennis</emphasis>
</taxonomicName>
; B,
<taxonomicName id="4C62A3F2FF8F385F3F05F9019597F97F" box="[1081,1265,1718,1737]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B9160463FF8F385F3F05F9019597F97F" box="[1081,1265,1718,1737]" italics="true" pageId="11" pageNumber="332">Pleophylla pilosa</emphasis>
</taxonomicName>
; C,
<emphasis id="B9160463FF8F385F3E1BF90290C0F950" bold="true" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3E1BF902905BF953" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="pseudopilosa" status="sp. nov.">
<emphasis id="B9160463FF8F385F3E1BF902905BF953" bold="true" italics="true" pageId="11" pageNumber="332">Pleophylla pseudopilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F3A75F96490C0F950" box="[329,422,1747,1766]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; D,
<emphasis id="B9160463FF8F385F3AE6F965921BF950" bold="true" box="[474,893,1746,1766]" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3AE6F9659273F953" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[474,789,1746,1765]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="nelshoogteensis" status="sp. nov.">
<emphasis id="B9160463FF8F385F3AE6F9659273F953" bold="true" box="[474,789,1746,1765]" italics="true" pageId="11" pageNumber="332">Pleophylla nelshoogteensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F381CF964921BF950" box="[800,893,1747,1766]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E,
<emphasis id="B9160463FF8F385F388CF965945FF950" bold="true" box="[944,1337,1746,1766]" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F388CF96595B6F953" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[944,1232,1746,1765]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="burundiensis" status="sp. nov.">
<emphasis id="B9160463FF8F385F388CF96595B6F953" bold="true" box="[944,1232,1746,1765]" italics="true" pageId="11" pageNumber="332">Pleophylla burundiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F3FE0F964945FF950" box="[1244,1337,1747,1766]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; F,
<emphasis id="B9160463FF8F385F3E56F9659363F8B5" bold="true" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3E56F96590C4F8B5" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="settentrionalis" status="sp. nov.">
<emphasis id="B9160463FF8F385F3E56F96590C4F8B5" bold="true" italics="true" pageId="11" pageNumber="332">Pleophylla settentrionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F3A96F9479363F8B5" box="[426,517,1776,1795]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; G,
<taxonomicName id="4C62A3F2FF8F385F3908F9479270F8B5" authorityName="Burmeister" authorityYear="1855" box="[564,790,1776,1795]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B9160463FF8F385F3908F9479270F8B5" box="[564,790,1776,1795]" italics="true" pageId="11" pageNumber="332">Pleophylla ferruginea</emphasis>
</taxonomicName>
; H,
<emphasis id="B9160463FF8F385F387AF94795EDF8B5" bold="true" box="[838,1163,1776,1795]" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F387AF947954EF8B5" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[838,1064,1776,1795]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="murzini" status="sp. nov.">
<emphasis id="B9160463FF8F385F387AF947954EF8B5" bold="true" box="[838,1064,1776,1795]" italics="true" pageId="11" pageNumber="332">Pleophylla murzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F3F0CF94795EDF8B5" box="[1072,1163,1776,1795]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; I,
<taxonomicName id="4C62A3F2FF8F385F3F8DF94794FBF8B5" authorityName="Burmeister" authorityYear="1855" box="[1201,1437,1776,1795]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="navicularis">
<emphasis id="B9160463FF8F385F3F8DF94794FBF8B5" box="[1201,1437,1776,1795]" italics="true" pageId="11" pageNumber="332">Pleophylla navicularis</emphasis>
</taxonomicName>
; J,
<taxonomicName id="4C62A3F2FF8F385F3BFFF8BA90F4F896" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[195,402,1805,1824]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="silvatica">
<emphasis id="B9160463FF8F385F3BFFF8BA90F4F896" box="[195,402,1805,1824]" italics="true" pageId="11" pageNumber="332">Pleophylla silvatica</emphasis>
</taxonomicName>
; K,
<emphasis id="B9160463FF8F385F3AF8F8BA922CF897" bold="true" box="[452,842,1805,1825]" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3AF8F8BA9383F896" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[452,741,1805,1824]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="transkeiensis" status="sp. nov.">
<emphasis id="B9160463FF8F385F3AF8F8BA9383F896" bold="true" box="[452,741,1805,1824]" italics="true" pageId="11" pageNumber="332">Pleophylla transkeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F39D3F8B9922CF897" box="[751,842,1806,1825]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; L,
<emphasis id="B9160463FF8F385F3845F8BA95D5F897" bold="true" box="[889,1203,1805,1825]" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3845F8BA9528F896" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[889,1102,1805,1824]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="ruthae" status="sp. nov.">
<emphasis id="B9160463FF8F385F3845F8BA9528F896" bold="true" box="[889,1102,1805,1824]" italics="true" pageId="11" pageNumber="332">Pleophylla ruthae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F3F64F8B995D5F897" box="[1112,1203,1806,1825]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; M,
<emphasis id="B9160463FF8F385F3FD5F8BA903CF888" bold="true" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3FD5F8BA919EF88B" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="warnockae" status="sp. nov.">
<emphasis id="B9160463FF8F385F3FD5F8BA919EF88B" bold="true" italics="true" pageId="11" pageNumber="332">Pleophylla warnockae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F3A3CF89C903CF888" box="[256,346,1835,1854]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; N,
<emphasis id="B9160463FF8F385F3AB4F89D93BAF888" bold="true" box="[392,732,1834,1854]" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3AB4F89D931DF88B" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[392,635,1834,1853]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="harrisoni" status="sp. nov.">
<emphasis id="B9160463FF8F385F3AB4F89D931DF88B" bold="true" box="[392,635,1834,1853]" italics="true" pageId="11" pageNumber="332">Pleophylla harrisoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F39BEF89C93BAF888" box="[642,732,1835,1854]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; O,
<emphasis id="B9160463FF8F385F3835F89D9548F888" bold="true" box="[777,1070,1834,1854]" pageId="11" pageNumber="332">
<taxonomicName id="4C62A3F2FF8F385F3835F89D92AAF88B" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[777,972,1834,1853]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
<emphasis id="B9160463FF8F385F3835F89D92AAF88B" bold="true" box="[777,972,1834,1853]" italics="true" pageId="11" pageNumber="332">Pleophylla stalsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8F385F38E8F89C9548F888" box="[980,1070,1835,1854]" pageId="11" pageNumber="332" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDDD871FF8838583BACFE669048FDC9" blockId="12.[144,759,464,639]" pageId="12" pageNumber="333">
<emphasis id="B9160463FF8838583BACFE66932EFE53" box="[144,584,464,486]" italics="true" pageId="12" pageNumber="333">
Diagnosis:
<taxonomicName id="4C62A3F2FF8838583A29FE67932EFE53" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[277,584,464,485]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="mlilwaneensis" status="sp. nov.">Pleophylla mlilwaneensis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A225B918FF883858395DFE6693ADFE50" box="[609,715,465,486]" pageId="12" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
is rather similar to
<taxonomicName id="4C62A3F2FF8838583A5BFE589353FDB2" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[359,565,495,516]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="333" phylum="Arthropoda" rank="species" species="nelshoogteensis" status="sp. nov.">
<emphasis id="B9160463FF8838583A5BFE589353FDB2" box="[359,565,495,516]" italics="true" pageId="12" pageNumber="333">P. nelshoogteensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF883858397FFE5893C6FDB2" box="[579,672,495,516]" pageId="12" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
in the shape of parameres. It differs from the latter in having the distal basomedial tooth in the apical third of the left paramere (rather than having both in basal half).
</paragraph>
<paragraph id="8BDDD871FF8838583BACFD2E906BFD5A" blockId="12.[144,758,665,748]" pageId="12" pageNumber="333">
<emphasis id="B9160463FF8838583BACFD2E906BFD18" box="[144,269,665,686]" italics="true" pageId="12" pageNumber="333">Etymology:</emphasis>
The new species is named after its
<typeStatus id="54D966D3FF88385839F9FD2E9390FD18" box="[709,758,665,686]" pageId="12" pageNumber="333">type</typeStatus>
locality, the Mlilwane Wildlife Sanctuary in
<collectingCountry id="F37598E1FF8838583BACFD61906FFD5A" box="[144,265,726,748]" name="Swaziland" pageId="12" pageNumber="333">Swaziland</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>