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<document id="8F34A26E48C91A60F978B37CF6F0EF51" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia &amp; Ahrens, Dirk" docDate="2017" docId="03CB6967FF8C385E3803FB9990E4FBCF" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla pseudopilosa AHRENS, BECKETT, EBERLE &amp; FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="331" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="329" updateTime="1738712560284" updateUser="ExternalLinkService">
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<mods:title id="726012F04228E2EB9242FB2DF47138C1">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="9528A021E1E87686DBF3AC27419F0E8E">Eberle, Jonas</mods:namePart>
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<mods:namePart id="55342AAB9BCAA254FDB5C9CCEC8FE1B5">Beckett, Marc</mods:namePart>
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<mods:namePart id="16BECADFC19C51E097CC6E8535BB9B41">Ozgul-Siemund, Alev</mods:namePart>
</mods:name>
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<mods:namePart id="1FF7BE7586024506574347E0E5F36A12">Frings, Jujina</mods:namePart>
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<mods:namePart id="1D356A6ED5ADA99D52B69492413A05AE">Fabrizi, Silvia</mods:namePart>
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<mods:date id="171EE584564B45E3107FA56DB0F00601">2017</mods:date>
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<mods:number id="2AF4EEA7EE73D847B7040932BF4BD422">180</mods:number>
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<treatment id="03CB6967FF8C385E3803FB9990E4FBCF" LSID="urn:lsid:plazi:treatment:03CB6967FF8C385E3803FB9990E4FBCF" httpUri="http://treatment.plazi.org/id/03CB6967FF8C385E3803FB9990E4FBCF" lastPageId="10" lastPageNumber="331" pageId="8" pageNumber="329">
<subSubSection id="C3788BFAFF8C385C3803FB999286FB77" pageId="8" pageNumber="329" type="nomenclature">
<paragraph id="8BDDD871FF8C385C3803FB999414FBD4" blockId="8.[831,1397,1066,1123]" pageId="8" pageNumber="329">
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<taxonomicName id="4C62A3F2FF8C385C3803FB99957DFBD7" ID-CoL="4K84C" authority="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityName="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pseudopilosa" status="sp. nov.">
<emphasis id="B9160463FF8C385C3803FB9995E9FBF4" bold="true" box="[831,1167,1070,1091]" italics="true" pageId="8" pageNumber="329">
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<smallCapsWord id="8D3B4EADFF8C385C38DDFB8695E9FBF4" baselines="1086" box="[993,1167,1073,1090]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="pseudopilosa" pageId="8" pageNumber="329">PSEUDOPILOSA</smallCapsWord>
</emphasis>
<smallCapsWord id="8D3B4EADFF8C385C3FA5FB9C9593FBF4" baselines="1085,1086" box="[1177,1269,1067,1091]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="8" pageNumber="329">AHRENS</smallCapsWord>
,
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,
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&amp;
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</taxonomicName>
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<smallCapsWord id="8D3B4EADFF8C385C3F19FBF89527FBD6" baselines="1116" box="[1061,1089,1103,1120]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="8" pageNumber="329">SP</smallCapsWord>
.
<smallCapsWord id="8D3B4EADFF8C385C3F6EFBF895E4FBD6" baselines="1116" box="[1106,1154,1103,1120]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="8" pageNumber="329">NOV</smallCapsWord>
.
</taxonomicNameLabel>
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(
<figureCitation id="1359C4F4FF8C385C3FA1FBFD9473FBD4" box="[1181,1301,1096,1122]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">
<smallCapsWord id="8D3B4EADFF8C385C3FA1FBFD95B4FBD7" baselines="1116,1117" box="[1181,1234,1098,1122]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="8" pageNumber="329">FIGS</smallCapsWord>
2I
<emphasis id="B9160463FF8C385C3FCAFBFB9462FBD4" box="[1270,1284,1100,1122]" italics="true" pageId="8" pageNumber="329">–</emphasis>
L
</figureCitation>
,
<figureCitation id="1359C4F4FF8C385C3E1AFBFF942EFBD4" box="[1318,1352,1096,1122]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="8" pageNumber="329">5C</figureCitation>
, 6)
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<uri id="FFF3D473FF8C385C3816FB399286FB77" pageId="8" pageNumber="329">
<emphasis id="B9160463FF8C385C3816FB39923CFB15" box="[810,858,1166,1187]" italics="true" pageId="8" pageNumber="329">urn:</emphasis>
lsid:zoobank.org:act:
<uuid id="FFC4E2A4FF8C385C3F02FB3A9286FB77" pageId="8" pageNumber="329">877B006F-29D1-44E7-AB0D- FB3F8189AE48</uuid>
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<paragraph id="8BDDD871FF8C385C381BFB0094EBFB7A" box="[807,1421,1207,1229]" pageId="8" pageNumber="329">
X-DA0394
<collectingCountry id="F37598E1FF8C385C38A1FB009541FB7B" box="[925,1063,1207,1229]" name="South Africa" pageId="8" pageNumber="329">South Africa</collectingCountry>
Cheerio Farm, Haenertsburg, ca
</paragraph>
<paragraph id="8BDDD871FF8C385C381BFB6E940CFADE" blockId="8.[807,1421,1241,1385]" pageId="8" pageNumber="329">
<emphasis id="B9160463FF8C385C381BFB6E9509FB58" box="[807,1135,1241,1262]" italics="true" pageId="8" pageNumber="329">Type material examined:</emphasis>
<materialsCitation id="3B0AD22CFF8C385C3F42FB6E9548FADE" collectingDate="2006-11-20" collectingDateMax="2006-11-22" collectingDateMin="2006-11-20" collectionCode="TMSA" collectorName="D. Ahrens &amp; S. Fabrizi" country="South Africa" elevation="1492" latitude="-53.01186" location="Cheerio Farm" longLatPrecision="1" longitude="57.002693" municipality="Haenertsburg" pageId="8" pageNumber="329" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54D966D3FF8C385C3F42FB6E9580FB59" box="[1150,1254,1241,1263]" pageId="8" pageNumber="329" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D6413F8FF8C385C3E3AFB6C9470FB5B" box="[1286,1302,1243,1261]" count="1" pageId="8" pageNumber="329" type="male">♂</specimenCount>
“834699 X-DA0394
<collectingCountry id="F37598E1FF8C385C38A1FB409541FABB" box="[925,1063,1271,1293]" name="South Africa" pageId="8" pageNumber="329">South Africa</collectingCountry>
<location id="8EBD8EAAFF8C385C3F10FB4095A0FABB" LSID="urn:lsid:plazi:treatment:03CB6967FF8C385E3803FB9990E4FBCF:8EBD8EAAFF8C385C3F10FB4095A0FABB" box="[1068,1222,1271,1293]" country="South Africa" latitude="-53.01186" longLatPrecision="1" longitude="57.002693" municipality="Haenertsburg" name="Cheerio Farm" pageId="8" pageNumber="329">Cheerio Farm</location>
,
<collectingMunicipality id="6BB9420BFF8C385C3FF3FB40940AFABB" box="[1231,1388,1271,1293]" pageId="8" pageNumber="329">Haenertsburg</collectingMunicipality>
, ca
<locationDeviation id="9A6FC153FF8C385C381BFAA19569FA9D" box="[807,1039,1302,1324]" location="Tzaneen" pageId="8" pageNumber="329">
<quantity id="4C9A7594FF8C385C381BFAA19208FA9A" box="[807,878,1302,1324]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="329" unit="km" value="20.0">20 km</quantity>
W of Tzaneen
</locationDeviation>
,
<quantity id="4C9A7594FF8C385C3F26FAA19508FA9D" box="[1050,1134,1302,1323]" metricMagnitude="3" metricUnit="m" metricValue="1.4919999999999998" pageId="8" pageNumber="329" unit="m" value="1492.0">
<elevation id="004F3F42FF8C385C3F26FAA19508FA9D" box="[1050,1134,1302,1323]" metricMagnitude="3" metricUnit="m" metricValue="1.4919999999999998" pageId="8" pageNumber="329" unit="m" value="1492.0">1492 m</elevation>
</quantity>
023°
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53
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42, 7
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S
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029°
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57
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09, 7
<emphasis id="B9160463FF8C385C3808FA839258FAFF" bold="true" box="[820,830,1332,1353]" italics="true" pageId="8" pageNumber="329">″</emphasis>
E
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<date id="FFDCFEB1FF8C385C3858FA83929BFAFC" box="[868,1021,1332,1354]" pageId="8" pageNumber="329" value="2006-11-20" valueMax="2006-11-22" valueMin="2006-11-20">
<collectingDate id="EF980759FF8C385C3858FA83929BFAFC" box="[868,1021,1332,1354]" pageId="8" pageNumber="329" value="2006-11-20" valueMax="2006-11-22" valueMin="2006-11-20">20-22.XI.2006</collectingDate>
</date>
leg.
<collectorName id="2697BDA7FF8C385C3F73FA8295A9FAFC" box="[1103,1231,1332,1354]" pageId="8" pageNumber="329">D. Ahrens</collectorName>
&amp;
<collectorName id="2697BDA7FF8C385C3E30FA8394EFFAFC" box="[1292,1417,1332,1354]" pageId="8" pageNumber="329">S. Fabrizi</collectorName>
,
<taxonomicName id="4C62A3F2FF8C385C381BFAE492A0FADE" box="[807,966,1363,1384]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B9160463FF8C385C381BFAE492FDFADE" box="[807,923,1363,1384]" italics="true" pageId="8" pageNumber="329">Pleophylla</emphasis>
sp1
</taxonomicName>
<emphasis id="B9160463FF8C385C38F9FAE492A9FADE" bold="true" box="[965,975,1363,1384]" italics="true" pageId="8" pageNumber="329">″</emphasis>
(
<collectionCode id="ED7340B4FF8C385C38E7FAE49541FADE" box="[987,1063,1363,1384]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="329">TMSA</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54D966D3FF8C385C3F0AFAE495CFFADE" box="[1078,1193,1363,1384]" pageId="8" pageNumber="329" type="paratype">Paratypes</typeStatus>
: see Appendix S1.
</paragraph>
<paragraph id="8BDDD871FF8C385E381BFA2793E6FE2F" blockId="8.[807,1422,1424,1904]" lastBlockId="10.[144,762,203,409]" lastPageId="10" lastPageNumber="331" pageId="8" pageNumber="329">
<emphasis id="B9160463FF8C385C381BFA2792C9FA13" box="[807,943,1424,1445]" italics="true" pageId="8" pageNumber="329">Description:</emphasis>
Body length:
<quantity id="4C9A7594FF8C385C3F52FA2795A1FA13" box="[1134,1223,1424,1445]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="329" unit="mm" value="8.3">8.3 mm</quantity>
, elytral length: 6.0 mm, maximum width:
<quantity id="4C9A7594FF8C385C3F53FA1995AFFA72" box="[1135,1225,1454,1476]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="329" unit="mm" value="4.6">4.6 mm</quantity>
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.65. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/ metacoxa: 1/1.34. Metatibia, ratio of width/length: 1/4. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
<figureCitation id="1359C4F4FF8E385E39BFFED39394FECC" box="[643,754,356,378]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
Fig. 2I
<emphasis id="B9160463FF8E385E39ECFED093BAFECF" box="[720,732,359,377]" italics="true" pageId="10" pageNumber="331">–</emphasis>
K
</figureCitation>
. Habitus:
<figureCitation id="1359C4F4FF8E385E3BC4FE349029FE2E" box="[248,335,387,409]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="10" pageNumber="331">
Fig.
<specimenCount id="9D6413F8FF8E385E3A11FE349029FE2E" box="[301,335,387,408]" count="2" pageId="10" pageNumber="331" type="generic">2L</specimenCount>
</figureCitation>
. Female genitalia:
<figureCitation id="1359C4F4FF8E385E3918FE34931AFE2F" box="[548,636,387,409]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="10" pageNumber="331">Fig. 5C</figureCitation>
.
</paragraph>
<caption id="DF1D88F9FF8D385D3B9AF93B9569F884" ID-DOI="http://doi.org/10.5281/zenodo.14807093" ID-Zenodo-Dep="14807093" httpUri="https://zenodo.org/record/14807093/files/figure.png" pageId="9" pageNumber="330" startId="9.[166,245,1676,1695]" targetBox="[189,1421,218,1624]" targetPageId="9" targetType="figure">
<paragraph id="8BDDD871FF8D385D3B9AF93B9569F884" blockId="9.[166,1444,1676,1842]" pageId="9" pageNumber="330">
<emphasis id="B9160463FF8D385D3B9AF93B9076F929" bold="true" box="[166,272,1676,1695]" pageId="9" pageNumber="330">Figure 4.</emphasis>
A
<emphasis id="B9160463FF8D385D3A11F938905EF929" box="[301,312,1679,1695]" italics="true" pageId="9" pageNumber="330">–</emphasis>
D,
<emphasis id="B9160463FF8D385D3A60F93B938FF916" bold="true" box="[348,745,1676,1696]" pageId="9" pageNumber="330">
<taxonomicName id="4C62A3F2FF8D385D3A60F93B93E6F929" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[348,640,1676,1695]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="mpumalanga" status="sp. nov.">
<emphasis id="B9160463FF8D385D3A60F93B93E6F929" bold="true" box="[348,640,1676,1695]" italics="true" pageId="9" pageNumber="330">Pleophylla mpumalanga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8D385D39B7F93A938FF916" box="[651,745,1677,1696]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); E
<emphasis id="B9160463FF8D385D3843F93892ECF929" box="[895,906,1679,1695]" italics="true" pageId="9" pageNumber="330">–</emphasis>
H,
<emphasis id="B9160463FF8D385D3893F93B9592F916" bold="true" box="[943,1268,1676,1696]" pageId="9" pageNumber="330">
<taxonomicName id="4C62A3F2FF8D385D3893F93B95EDF929" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[943,1163,1676,1695]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="charlyi" status="sp. nov.">
<emphasis id="B9160463FF8D385D3893F93B95EDF929" bold="true" box="[943,1163,1676,1695]" italics="true" pageId="9" pageNumber="330">Pleophylla charlyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8D385D3FABF93A9592F916" box="[1175,1268,1677,1696]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); I
<emphasis id="B9160463FF8D385D3EBFF93894E8F929" box="[1411,1422,1679,1695]" italics="true" pageId="9" pageNumber="330">–</emphasis>
L,
<emphasis id="B9160463FF8D385D3B9AF91E90BEF90B" bold="true" box="[166,472,1705,1725]" pageId="9" pageNumber="330">
<taxonomicName id="4C62A3F2FF8D385D3B9AF91E9014F90A" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[166,370,1705,1724]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="lizleri" status="sp. nov.">
<emphasis id="B9160463FF8D385D3B9AF91E9014F90A" bold="true" box="[166,370,1705,1724]" italics="true" pageId="9" pageNumber="330">Pleophylla lizleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8D385D3A40F91D90BEF90B" box="[380,472,1706,1725]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); M
<emphasis id="B9160463FF8D385D3953F91B931CF90A" box="[623,634,1708,1724]" italics="true" pageId="9" pageNumber="330">–</emphasis>
P,
<emphasis id="B9160463FF8D385D39A5F91E9291F90B" bold="true" box="[665,1015,1705,1725]" pageId="9" pageNumber="330">
<taxonomicName id="4C62A3F2FF8D385D39A5F91E92F4F90B" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[665,914,1705,1725]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="taitaensis" status="sp. nov.">
<emphasis id="B9160463FF8D385D39A5F91E92F4F90B" bold="true" box="[665,914,1705,1725]" italics="true" pageId="9" pageNumber="330">Pleophylla taitaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8D385D38A7F91D9291F90B" box="[923,1015,1706,1725]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); Q
<emphasis id="B9160463FF8D385D3FB6F91B95F3F90A" box="[1162,1173,1708,1724]" italics="true" pageId="9" pageNumber="330">–</emphasis>
T,
<emphasis id="B9160463FF8D385D3F88F91E91BEF96C" bold="true" pageId="9" pageNumber="330">
<taxonomicName id="4C62A3F2FF8D385D3F88F91E941FF90A" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[1204,1401,1705,1724]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="stalsi" status="sp. nov.">
<emphasis id="B9160463FF8D385D3F88F91E941FF90A" bold="true" box="[1204,1401,1705,1724]" italics="true" pageId="9" pageNumber="330">Pleophylla stalsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8D385D3EBFF91D91BEF96C" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); U
<emphasis id="B9160463FF8D385D3A52F97E901FF96F" box="[366,377,1737,1753]" italics="true" pageId="9" pageNumber="330">–</emphasis>
X,
<emphasis id="B9160463FF8D385D3AA6F970920EF96C" bold="true" box="[410,872,1735,1754]" pageId="9" pageNumber="330">
<taxonomicName id="4C62A3F2FF8D385D3AA6F9709266F96C" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[410,768,1735,1754]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="wakkerstroomensis" status="sp. nov.">
<emphasis id="B9160463FF8D385D3AA6F9709266F96C" bold="true" box="[410,768,1735,1754]" italics="true" pageId="9" pageNumber="330">Pleophylla wakkerstroomensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8D385D3837F970920EF96C" box="[779,872,1735,1754]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype); Y
<emphasis id="B9160463FF8D385D38C7F97E9560F96F" box="[1019,1030,1737,1753]" italics="true" pageId="9" pageNumber="330">–</emphasis>
Ab,
<taxonomicName id="4C62A3F2FF8D385D3F08F97094C2F96C" authority="Peringuey" authorityName="Peringuey" authorityYear="1904" box="[1076,1444,1735,1754]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="tongaatsana">
<emphasis id="B9160463FF8D385D3F08F9709448F96C" box="[1076,1326,1735,1754]" italics="true" pageId="9" pageNumber="330">Pleophylla tongaatsana</emphasis>
Peringuey
</taxonomicName>
(RSA, Sjonajona); Ac
<emphasis id="B9160463FF8D385D3AB9F95090F6F941" box="[389,400,1767,1783]" italics="true" pageId="9" pageNumber="330">–</emphasis>
Af,
<emphasis id="B9160463FF8D385D3A84F9539261F941" bold="true" box="[440,775,1764,1783]" pageId="9" pageNumber="330">
<taxonomicName id="4C62A3F2FF8D385D3A84F95393C4F941" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[440,674,1764,1783]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="kruegeri" status="sp. nov.">
<emphasis id="B9160463FF8D385D3A84F95393C4F941" bold="true" box="[440,674,1764,1783]" italics="true" pageId="9" pageNumber="330">Pleophylla kruegeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8D385D3997F9539261F941" box="[683,775,1764,1783]" pageId="9" pageNumber="330" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale.
</paragraph>
</caption>
<paragraph id="8BDDD871FF8E385E3BADFE1993F8FD8B" blockId="10.[145,760,430,574]" pageId="10" pageNumber="331">
<emphasis id="B9160463FF8E385E3BADFE199356FE75" box="[145,560,430,451]" italics="true" pageId="10" pageNumber="331">
Diagnosis:
<taxonomicName id="4C62A3F2FF8E385E3A2BFE199356FE75" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[279,560,430,451]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pseudopilosa" status="sp. nov.">Pleophylla pseudopilosa</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A225B918FF8E385E397CFE1993C6FE75" box="[576,672,430,451]" pageId="10" pageNumber="331" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C62A3F2FF8E385E3BE6FE7A9059FE57" box="[218,319,460,482]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B9160463FF8E385E3BE6FE7A9059FE57" box="[218,319,460,482]" italics="true" pageId="10" pageNumber="331">P. pilosa</emphasis>
</taxonomicName>
in the parameres by having the ventral lobe in lateral view wider and shorter; the base of the dorsal lobe of left paramere has a single narrow medial tooth (not two as in
<taxonomicName id="4C62A3F2FF8E385E3916FD9F93E8FD8B" box="[554,654,552,573]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B9160463FF8E385E3916FD9F93E8FD8B" box="[554,654,552,573]" italics="true" pageId="10" pageNumber="331">P. pilosa</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BDDD871FF8E385E3BADFDEF938BFCD4" blockId="10.[145,760,599,866]" pageId="10" pageNumber="331">
<emphasis id="B9160463FF8E385E3BADFDEF9063FDDB" box="[145,261,600,621]" italics="true" pageId="10" pageNumber="331">Variation:</emphasis>
Body length:
<quantity id="4C9A7594FF8E385E3A8BFDE09359FDDB" box="[439,575,599,621]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.5" metricValueMin="6.8" pageId="10" pageNumber="331" unit="mm" value="7.65" valueMax="8.5" valueMin="6.8">
6.8
<emphasis id="B9160463FF8E385E3AE5FDED9083FDDA" box="[473,485,602,620]" italics="true" pageId="10" pageNumber="331">–</emphasis>
8.5 mm
</quantity>
, elytral length:
<quantity id="4C9A7594FF8E385E3BADFDC1907FFD3A" box="[145,281,630,652]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="6.1" metricValueMin="4.2" pageId="10" pageNumber="331" unit="mm" value="5.15" valueMax="6.1" valueMin="4.2">
4.2
<emphasis id="B9160463FF8E385E3B8FFDCE91D9FD3D" box="[179,191,633,651]" italics="true" pageId="10" pageNumber="331">–</emphasis>
6.1 mm
</quantity>
, maximum width:
<quantity id="4C9A7594FF8E385E3932FDC193F0FD3A" box="[526,662,630,652]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.8" metricValueMin="3.8" pageId="10" pageNumber="331" unit="mm" value="4.3" valueMax="4.8" valueMin="3.8">
3.8
<emphasis id="B9160463FF8E385E390CFDCE935AFD3D" box="[560,572,633,651]" italics="true" pageId="10" pageNumber="331">–</emphasis>
4.8 mm
</quantity>
. Colour varies from reddish brown with dark spot to entirely yellowish brown without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere subequal a quarter of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.5.
</paragraph>
<paragraph id="8BDDD871FF8E385E3BAEFCCD9086FBBC" blockId="10.[146,761,890,1034]" pageId="10" pageNumber="331">
<emphasis id="B9160463FF8E385E3BAEFCCD9076FC39" box="[146,272,890,911]" italics="true" pageId="10" pageNumber="331">Etymology:</emphasis>
The name of the new species is composed of the Latinized Greek prefix pseudo- (false) and pilosa (species name of
<taxonomicName id="4C62A3F2FF8E385E3910FC009394FC7A" box="[556,754,951,972]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B9160463FF8E385E3910FC009394FC7A" box="[556,754,951,972]" italics="true" pageId="10" pageNumber="331">Pleophylla pilosa</emphasis>
</taxonomicName>
, with reference to its general similarity with this species) (noun in apposition).
</paragraph>
<paragraph id="8BDDD871FF8E385E3BAEFB9190E4FBCF" blockId="10.[146,761,1062,1145]" pageId="10" pageNumber="331">
<emphasis id="B9160463FF8E385E3BAEFB919198FB8D" box="[146,254,1062,1083]" italics="true" pageId="10" pageNumber="331">Remarks:</emphasis>
In previous publications we referred to this species as ‘sp1
<emphasis id="B9160463FF8E385E3A51FBF79014FBF9" bold="true" box="[365,370,1088,1103]" italics="true" pageId="10" pageNumber="331">
<superScript id="7C177539FF8E385E3A51FBF79014FBF9" attach="left" box="[365,370,1088,1103]" fontSize="6" pageId="10" pageNumber="331">0</superScript>
</emphasis>
(Ozg € ul-Siemund € &amp;
<bibRefCitation id="EFF3A580FF8E385E3966FBF39393FBEC" author="Ahrens D" box="[602,757,1092,1114]" pageId="10" pageNumber="331" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
;
<bibRefCitation id="EFF3A580FF8E385E3BAEFBD49010FBCF" author="Eberle J &amp; Warnock RCM &amp; Ahrens D" box="[146,374,1123,1145]" pageId="10" pageNumber="331" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
Eberle
<emphasis id="B9160463FF8E385E3BD9FBD39079FBCE" box="[229,287,1123,1145]" italics="true" pageId="10" pageNumber="331">et al.</emphasis>
, 2016b
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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