226 lines
26 KiB
XML
226 lines
26 KiB
XML
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<mods:title id="3AE2A5AF413CF63BC8DAD8CCCDCEDEEB">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<taxonomicName id="4C62A3F2FF8C385C3BD6FB7093FDFB6A" ID-CoL="4K84B" authority="BOHEMAN, 1857" authorityName="Boheman" authorityYear="1857" box="[234,667,1218,1245]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
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<bibRefCitation id="EFF3A580FF8C385C3AE7FB7393FDFB6A" author="Boheman CH" box="[475,667,1218,1245]" pageId="8" pageNumber="329" refId="ref23181" refString="Boheman CH. 1857. Insecta Caffrariae. Annis 1838 - 1845. Part II. Holmiae, 1 - 395." type="book" year="1857">
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<smallCapsWord id="8D3B4EADFF8C385C3AE7FB73932AFB6D" baselines="1238,1239" box="[475,588,1220,1244]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Boheman" pageId="8" pageNumber="329">BOHEMAN</smallCapsWord>
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, 1857
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(
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<figureCitation id="1359C4F4FF8C385C3A6BFB5490BDFB4D" box="[343,475,1249,1275]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">
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<smallCapsWord id="8D3B4EADFF8C385C3A6BFB5490EAFB4F" baselines="1269,1269" box="[343,396,1251,1275]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="8" pageNumber="329">FIGS</smallCapsWord>
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2E
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<emphasis id="B9160463FF8C385C3A84FB5290A0FB4D" box="[440,454,1253,1275]" italics="true" pageId="8" pageNumber="329">–</emphasis>
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H
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</figureCitation>
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,
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<figureCitation id="1359C4F4FF8C385C3AD0FB569368FB4D" box="[492,526,1249,1275]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="8" pageNumber="329">5B</figureCitation>
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, 6)
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</heading>
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<treatmentCitationGroup id="AB72FF5FFF8C385C3BACFAA590FEFAD3" pageId="8" pageNumber="329">
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<treatmentCitation id="0AC3FE60FF8C385C3BACFAA59328FA9E" author="Boheman CH" box="[144,590,1298,1320]" page="125" pageId="8" pageNumber="329" year="1857">
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<taxonomicName id="4C62A3F2FF8C385C3BACFAA59328FA9E" ID-CoL="4K84B" authority="Boheman, 1857: 125" authorityName="Boheman" authorityPageNumber="125" authorityYear="1857" box="[144,590,1298,1320]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
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<emphasis id="B9160463FF8C385C3BACFAA59030FA91" box="[144,342,1298,1319]" italics="true" pageId="8" pageNumber="329">Pleophylla pilosa</emphasis>
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<bibRefCitation id="EFF3A580FF8C385C3A5CFAA59328FA9E" author="Boheman CH" box="[352,590,1298,1320]" pageId="8" pageNumber="329" refId="ref23181" refString="Boheman CH. 1857. Insecta Caffrariae. Annis 1838 - 1845. Part II. Holmiae, 1 - 395." type="book" year="1857">Boheman, 1857: 125</bibRefCitation>
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</taxonomicName>
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[
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<typeStatus id="54D966D3FF8C385C3963FAA493F4FA9E" box="[607,658,1299,1320]" pageId="8" pageNumber="329">type</typeStatus>
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locality:
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<collectingRegion id="49A61693FF8C385C3BACFA869194FAF1" box="[144,242,1329,1351]" country="South Africa" name="Limpopo" pageId="8" pageNumber="329">Limpopo</collectingRegion>
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river]
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<treatmentCitation id="0AC3FE60FF8C385C3A79FA86935AFAF0" author="Schaufuss LW" box="[325,572,1329,1351]" page="232" pageId="8" pageNumber="329" year="1871">
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<bibRefCitation id="EFF3A580FF8C385C3A79FA86935AFAF0" author="Schaufuss LW" box="[325,572,1329,1351]" pageId="8" pageNumber="329" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 232</bibRefCitation>
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</treatmentCitation>
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; Brenske, 1900: 84; Peringuey, 1904: 8.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BDDD871FF8C385C3BADFAD991A3FA76" blockId="8.[144,759,1298,1472]" pageId="8" pageNumber="329">
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<treatmentCitationGroup id="AB72FF5FFF8C385C3BADFAD991A3FA76" pageId="8" pageNumber="329">
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<treatmentCitation id="0AC3FE60FF8C385C3BADFAD9931EFA35" author="Schaufuss LW" box="[145,632,1390,1412]" page="232" pageId="8" pageNumber="329" year="1871">
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<taxonomicName id="4C62A3F2FF8C385C3BADFAD9931EFA35" ID-CoL="6VQMY" authority="Schaufuss, 1871: 232" authorityName="Schaufuss" authorityPageNumber="232" authorityYear="1871" box="[145,632,1390,1412]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
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<emphasis id="B9160463FF8C385C3BADFAD9900BFA35" box="[145,365,1390,1411]" italics="true" pageId="8" pageNumber="329">Pleophylla opalina</emphasis>
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<bibRefCitation id="EFF3A580FF8C385C3A47FAD9931EFA35" author="Schaufuss LW" box="[379,632,1390,1412]" pageId="8" pageNumber="329" pagination="231 - 233" refId="ref24095" refString="Schaufuss LW. 1871. Die Arten der Gattung Pleophylla Er. Nunquam Otiosus I 6: 231-233." type="journal article" year="1871">Schaufuss, 1871: 232</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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[Caffraria interia]; Brenske, 1900: 84, Peringuey, 1904: 8;
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<emphasis id="B9160463FF8C385C39FFFA3A91A3FA76" bold="true" pageId="8" pageNumber="329">syn. nov.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3788BFAFF8C385C3BACFA5E95DBFC41" pageId="8" pageNumber="329" type="materials_examined">
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<paragraph id="8BDDD871FF8C385C3BACFA5E9084F965" blockId="8.[144,759,1512,1748]" pageId="8" pageNumber="329">
|
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<emphasis id="B9160463FF8C385C3BACFA5E90CAFA4B" box="[144,428,1512,1534]" italics="true" pageId="8" pageNumber="329">Type material examined:</emphasis>
|
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<materialsCitation id="3B0AD22CFF8C385C3A87FA5E9395F9EF" collectionCode="NHRS" collectorName="J. Wahlb." location="Stockholm" pageId="8" pageNumber="329" specimenCount="1" specimenCount-male="1" stateProvince="Stockholm" typeStatus="lectotype">
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<typeStatus id="54D966D3FF8C385C3A87FA5E934EFA48" box="[443,552,1513,1534]" pageId="8" pageNumber="329" type="lectotype">Lectotype</typeStatus>
|
||
(here designated):
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||
<specimenCount id="9D6413F8FF8C385C3BACF9BD91C6F9AA" box="[144,160,1546,1564]" count="1" pageId="8" pageNumber="329" type="male">♂</specimenCount>
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“Caffraria./
|
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<collectorName id="2697BDA7FF8C385C3A12F9B090F0F9AB" box="[302,406,1543,1565]" pageId="8" pageNumber="329">J. Wahlb.</collectorName>
|
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/Type./
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<emphasis id="B9160463FF8C385C3ADEF9B093D5F9AA" box="[482,691,1543,1564]" italics="true" pageId="8" pageNumber="329">
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<taxonomicName id="4C62A3F2FF8C385C3ADEF9B093C0F9AA" authorityName="Boheman" authorityYear="1857" box="[482,678,1543,1564]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">Pleophylla pilosa</taxonomicName>
|
||
/
|
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</emphasis>
|
||
<taxonomicName id="4C62A3F2FF8C385C398FF9B090D0F98D" authority="Bhn. Ent. Kaffr." class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B9160463FF8C385C398FF9B09390F9AA" box="[691,758,1543,1564]" italics="true" pageId="8" pageNumber="329">pilosa</emphasis>
|
||
Bhn. Ent. Kaffr.
|
||
</taxonomicName>
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||
/3680E91 + /Naturhistoriska Riksmuseet
|
||
<collectingRegion id="49A61693FF8C385C3A18F9F390FCF9EC" box="[292,410,1604,1626]" country="Sweden" name="Stockholm" pageId="8" pageNumber="329">Stockholm</collectingRegion>
|
||
Loan no. 1746/07” (
|
||
<collectionCode id="ED7340B4FF8C385C39A3F9F3938AF9EF" box="[671,748,1604,1625]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="8" pageNumber="329">NHRS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0AD22CFF8C385C3BACF9D490B8F965" collectionCode="ZMHB" collectorName="Coll. L. W. Schaufuss" pageId="8" pageNumber="329" specimenCount="1" specimenCount-male="1" stateProvince="Stockholm" typeStatus="lectotype">
|
||
<typeStatus id="54D966D3FF8C385C3BACF9D4919BF9CE" box="[144,253,1635,1656]" pageId="8" pageNumber="329" type="lectotype">Lectotype</typeStatus>
|
||
(
|
||
<taxonomicName id="4C62A3F2FF8C385C3A2FF9D49001F9CE" authorityName="Schaufuss" authorityYear="1871" box="[275,359,1635,1656]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
|
||
<emphasis id="B9160463FF8C385C3A2FF9D49001F9CE" box="[275,359,1635,1656]" italics="true" pageId="8" pageNumber="329">opalina</emphasis>
|
||
</taxonomicName>
|
||
, here designated):
|
||
<specimenCount id="9D6413F8FF8C385C396AF9D29300F9C1" box="[598,614,1637,1655]" count="1" pageId="8" pageNumber="329" type="male">♂</specimenCount>
|
||
“
|
||
<collectorName id="2697BDA7FF8C385C3943F9D49067F921" pageId="8" pageNumber="329">Coll. L.W. Schaufuss</collectorName>
|
||
/Nunqn at pag. 232/
|
||
<taxonomicName id="4C62A3F2FF8C385C3ACAF9369391F921" authority="Schauf. Caffr." authorityName="Schauf. Caffr." box="[502,759,1665,1687]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">opalina Schauf. Caffr.</taxonomicName>
|
||
inter./Type./
|
||
<taxonomicName id="4C62A3F2FF8C385C3A24F9179089F903" box="[280,495,1696,1717]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
|
||
<emphasis id="B9160463FF8C385C3A24F9179089F903" box="[280,495,1696,1717]" italics="true" pageId="8" pageNumber="329">Pleophylla opalina</emphasis>
|
||
</taxonomicName>
|
||
m.
|
||
<taxonomicName id="4C62A3F2FF8C385C3919F9179391F903" authority="Burm." authorityName="Burmeister" authorityYear="1855" box="[549,759,1696,1718]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="navicularis">navicularis Burm.</taxonomicName>
|
||
[sic! canceled] Natal” (
|
||
<collectionCode id="ED7340B4FF8C385C3ABAF90990B1F965" box="[390,471,1726,1747]" pageId="8" pageNumber="329">ZMHB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF8C385C3BACF94C93A0F8A7" blockId="8.[144,710,1787,1809]" box="[144,710,1787,1809]" pageId="8" pageNumber="329">
|
||
<emphasis id="B9160463FF8C385C3BACF94C9089F8A6" box="[144,495,1787,1808]" italics="true" pageId="8" pageNumber="329">Additional material examined:</emphasis>
|
||
See Appendix S1.
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF8C385C3BACF88F9514FC89" blockId="8.[144,759,1848,1901]" lastBlockId="8.[807,1422,199,832]" pageId="8" pageNumber="329">
|
||
<emphasis id="B9160463FF8C385C3BACF88F9057F8FB" box="[144,305,1848,1869]" italics="true" pageId="8" pageNumber="329">Redescription:</emphasis>
|
||
Body length: 9.0 mm, elytral length:
|
||
<quantity id="4C9A7594FF8C385C3BACF8E0918FF8DB" box="[144,233,1879,1901]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="8" pageNumber="329" unit="mm" value="6.1">6.1 mm</quantity>
|
||
, maximum width:
|
||
<quantity id="4C9A7594FF8C385C3AE4F8E09355F8DB" box="[472,563,1879,1901]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="329" unit="mm" value="4.7">4.7 mm</quantity>
|
||
. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially straight; margins of labroclypeus moderately reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.68. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum moderately large. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.33. Metatibia, ratio of width/length: 1/3.87. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
|
||
<figureCitation id="1359C4F4FF8C385C381BFC9D92FBFC89" box="[807,925,810,832]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">
|
||
Fig. 2E
|
||
<emphasis id="B9160463FF8C385C3847FC9A92E1FC89" box="[891,903,813,831]" italics="true" pageId="8" pageNumber="329">–</emphasis>
|
||
G
|
||
</figureCitation>
|
||
. Habitus:
|
||
<figureCitation id="1359C4F4FF8C385C3F28FC9D9509FC89" box="[1044,1135,810,832]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="8" pageNumber="329">Fig. 2H</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDDD871FF8C385C381BFCD095DBFC41" blockId="8.[807,1421,871,1015]" pageId="8" pageNumber="329">
|
||
<emphasis id="B9160463FF8C385C381BFCD092F4FCCA" box="[807,914,871,892]" italics="true" pageId="8" pageNumber="329">Remarks:</emphasis>
|
||
The shape of the parameres of
|
||
<taxonomicName id="4C62A3F2FF8C385C3E2BFCDF94EBFCCA" box="[1303,1421,871,893]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="opalina">
|
||
<emphasis id="B9160463FF8C385C3E2BFCDF94EBFCCA" box="[1303,1421,871,893]" italics="true" pageId="8" pageNumber="329">P. opalina</emphasis>
|
||
</taxonomicName>
|
||
are virtually identical in shape to those of the
|
||
<typeStatus id="54D966D3FF8C385C381BFC1392EBFC0C" box="[807,909,932,954]" pageId="8" pageNumber="329" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C62A3F2FF8C385C38FCFC129542FC0F" box="[960,1060,932,954]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="B9160463FF8C385C38FCFC129542FC0F" box="[960,1060,932,954]" italics="true" pageId="8" pageNumber="329">P. pilosa</emphasis>
|
||
</taxonomicName>
|
||
. In previous publications we referred to this species as ‘sp6’ (Ozg € ul-Siemund € &
|
||
<bibRefCitation id="EFF3A580FF8C385C381BFC5692A7FC41" author="Ahrens D" box="[807,961,993,1015]" pageId="8" pageNumber="329" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF3A580FF8C385C38F1FC5695D7FC41" author="Eberle J & Warnock RCM & Ahrens D" box="[973,1201,993,1015]" pageId="8" pageNumber="329" refId="ref23428" refString="Eberle J, Warnock RCM, Ahrens D. 2016 b. Bayesian species delimitation in Pleophylla chafers (Coleoptera) - the importance of prior choice and morphology. BMC Evolutionary Biology 16: 94." type="journal volume" year="2016">
|
||
Eberle
|
||
<emphasis id="B9160463FF8C385C3F1CFC55953CFC40" box="[1056,1114,993,1015]" italics="true" pageId="8" pageNumber="329">et al.</emphasis>
|
||
, 2016b
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |