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<document id="BD7FB604767BA71276A642E43915990C" ID-DOI="10.1111/zoj.12489" ID-ISSN="0024-4082" ID-Zenodo-Dep="14807083" ID-ZooBank="3663E0A-2E17-4346-A388-6BEE53081814" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1738712105264" checkinUser="plazi" docAuthor="Eberle, Jonas, Beckett, Marc, Ozgul-Siemund, Alev, Frings, Jujina, Fabrizi, Silvia &amp; Ahrens, Dirk" docDate="2017" docId="03CB6967FF8A385B3BA9F9C993BAFD73" docLanguage="en" docName="zoj.12489.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 180 (2)" docSource="https://www.mendeley.com/catalogue/f558ad26-a47a-355d-9f45-4a1c420a1594/" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pleophylla settentrionalis AHRENS, BECKETT, EBERLE &amp; FABRIZI 2017, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="336" masterDocId="FFF2111FFF8438543B3CFFB79166FFB6" masterDocTitle="Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision" masterLastPageNumber="353" masterPageNumber="321" pageNumber="335" updateTime="1738712560284" updateUser="ExternalLinkService">
<mods:mods id="07623E6A0552DB5E54B544B2117405A2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6560E179918CD6D812C5AA84C163F4B0">Afromontane forests hide 19 new species of an ancient chafer lineage (Coleoptera: Scarabaeidae: Melolonthinae): Pleophylla Erichson, 1847 - phylogeny and taxonomic revision</mods:title>
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<mods:namePart id="D173E79BA80ED53660D9C42EE634881E">Eberle, Jonas</mods:namePart>
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<mods:namePart id="EE65DB9E363FC6E0F751A4508C09C656">Beckett, Marc</mods:namePart>
</mods:name>
<mods:name id="1B14A5E1E68C3C7D0F6D17A5C3865331" type="personal">
<mods:role id="3005C2360059F537A6C6CD840255A9BF">
<mods:roleTerm id="8635E1D31866F59A56C4FAF6E18F42AD">Author</mods:roleTerm>
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<mods:namePart id="1AE097FC790F28870EE8C279F89E9DF5">Ozgul-Siemund, Alev</mods:namePart>
</mods:name>
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<mods:namePart id="D74B9B1877AB5C84D2CBBBA9FF9641CB">Frings, Jujina</mods:namePart>
</mods:name>
<mods:name id="FC5392B087C41966BA8F1BDF88E218D2" type="personal">
<mods:role id="DF66296DF8B6AB28542E00B36C62F6F9">
<mods:roleTerm id="35AFA866B858CC5CD9A616CD96000C3B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F38256263FD5AC4B6ADF091B098A5F2">Fabrizi, Silvia</mods:namePart>
</mods:name>
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<mods:role id="48DCEC4267B0041E95F22F80E8AF0019">
<mods:roleTerm id="9D209BF24BDD96405EBFF60830E36382">Author</mods:roleTerm>
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<mods:namePart id="1FC8107CC02041C199C56CDA1A68EDDA">Ahrens, Dirk</mods:namePart>
</mods:name>
<mods:typeOfResource id="710A79984678A116A3522AD8ECC82839">text</mods:typeOfResource>
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<mods:title id="7B1DDF53C8DE486D36E39761A3024B62">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5532BF8128585380AC1FA49AFBACD47B">2017</mods:date>
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<mods:title id="C8BD874BA52B912A61712B7D74B1CEE4">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="D47979398B0809A85FE3B0262D7C9EB9">180</mods:number>
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<mods:number id="A2FE725CDA985DB78F82FB40802DA84C">2</mods:number>
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<treatment id="03CB6967FF8A385B3BA9F9C993BAFD73" LSID="urn:lsid:plazi:treatment:03CB6967FF8A385B3BA9F9C993BAFD73" httpUri="http://treatment.plazi.org/id/03CB6967FF8A385B3BA9F9C993BAFD73" lastPageId="15" lastPageNumber="336" pageId="14" pageNumber="335">
<subSubSection id="C3788BFAFF8A385A3BA9F9C9905EF8B5" pageId="14" pageNumber="335" type="nomenclature">
<paragraph id="8BDDD871FF8A385A3BA9F9C9938BF904" blockId="14.[149,753,1659,1715]" pageId="14" pageNumber="335">
<heading id="D0956F1DFF8A385A3BA9F9C9938BF904" centered="true" fontSize="9" level="2" pageId="14" pageNumber="335" reason="2">
<taxonomicName id="4C62A3F2FF8A385A3BA9F9C99015F907" ID-CoL="6W2MY" authority="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityName="AHRENS, BECKETT, EBERLE &amp; FABRIZI" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="settentrionalis" status="sp. nov.">
<emphasis id="B9160463FF8A385A3BA9F9C9936DF924" bold="true" box="[149,523,1662,1683]" italics="true" pageId="14" pageNumber="335">
<smallCapsWord id="8D3B4EADFF8A385A3BA9F9C99049F924" baselines="1678,1678" box="[149,303,1662,1683]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Pleophylla" pageId="14" pageNumber="335">PLEOPHYLLA</smallCapsWord>
<smallCapsWord id="8D3B4EADFF8A385A3A04F936936DF924" baselines="1678" box="[312,523,1665,1682]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="settentrionalis" pageId="14" pageNumber="335">SETTENTRIONALIS</smallCapsWord>
</emphasis>
<smallCapsWord id="8D3B4EADFF8A385A3929F9CB9316F924" baselines="1678,1678" box="[533,624,1660,1684]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ahrens" pageId="14" pageNumber="335">AHRENS</smallCapsWord>
,
<smallCapsWord id="8D3B4EADFF8A385A39BEF9CB938FF924" baselines="1678,1678" box="[642,745,1660,1684]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Beckett" pageId="14" pageNumber="335">BECKETT</smallCapsWord>
,
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&amp;
<smallCapsWord id="8D3B4EADFF8A385A3A27F92D9015F907" baselines="1708,1709" box="[283,371,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Fabrizi" pageId="14" pageNumber="335">FABRIZI</smallCapsWord>
</taxonomicName>
<emphasis id="B9160463FF8A385A3A40F9179084F905" bold="true" box="[380,482,1688,1715]" pageId="14" pageNumber="335">
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<smallCapsWord id="8D3B4EADFF8A385A3A40F91790FEF907" baselines="1709" box="[380,408,1696,1713]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="sp" pageId="14" pageNumber="335">SP</smallCapsWord>
.
<smallCapsWord id="8D3B4EADFF8A385A3A96F91790BCF907" baselines="1709" box="[426,474,1696,1713]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="nov" pageId="14" pageNumber="335">NOV</smallCapsWord>
.
</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1359C4F4FF8A385A3AC8F92D93F7F906" box="[500,657,1688,1714]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">
<smallCapsWord id="8D3B4EADFF8A385A3AC8F92D934FF907" baselines="1708,1709" box="[500,553,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Figs" pageId="14" pageNumber="335">FIGS</smallCapsWord>
2
<smallCapsWord id="8D3B4EADFF8A385A397FF92D9305F907" baselines="1708,1709" box="[579,611,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ac" pageId="14" pageNumber="335">AC</smallCapsWord>
<emphasis id="B9160463FF8A385A3958F92B9314F904" box="[612,626,1692,1714]" italics="true" pageId="14" pageNumber="335">–</emphasis>
<smallCapsWord id="8D3B4EADFF8A385A394EF92D93F7F906" baselines="1708,1708" box="[626,657,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Af" pageId="14" pageNumber="335">AF</smallCapsWord>
</figureCitation>
,
<figureCitation id="1359C4F4FF8A385A399EF92F93A5F904" box="[674,707,1688,1714]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="14" pageNumber="335">5F</figureCitation>
,
<figureCitation id="1359C4F4FF8A385A39E8F92F9382F904" box="[724,740,1688,1714]" captionStart="Figure 7" captionStartId="14.[144,223,925,944]" captionTargetBox="[144,1428,192,893]" captionTargetId="figure-25@14.[144,1428,192,893]" captionTargetPageId="14" captionText="Figure 7. Distribution of Pleophylla species (left side) and absence records (right side), represented by sampling locali- ties of all other sampled Sericini species without records of Pleophylla (white dots)." figureDoi="http://doi.org/10.5281/zenodo.14807097" httpUri="https://zenodo.org/record/14807097/files/figure.png" pageId="14" pageNumber="335">7</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BDDD871FF8A385A3BADF978905EF8B5" blockId="14.[145,761,1743,1795]" pageId="14" pageNumber="335">
<uri id="FFF3D473FF8A385A3BADF978905EF8B5" pageId="14" pageNumber="335">
urn:lsid:zoobank.org:act:
<uuid id="FFC4E2A4FF8A385A3AA5F978905EF8B5" pageId="14" pageNumber="335">FAA40DFF-EB25-413D-8573- 6993258C3090</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3788BFAFF8A385B3BACF8AD93BAFD73" lastPageId="15" lastPageNumber="336" pageId="14" pageNumber="335" type="materials_examined">
<paragraph id="8BDDD871FF8A385A3BACF8AD9324F8DA" blockId="14.[144,759,1818,1901]" pageId="14" pageNumber="335">
<emphasis id="B9160463FF8A385A3BACF8AD90A7F899" box="[144,449,1818,1839]" italics="true" pageId="14" pageNumber="335">Type material examined:</emphasis>
<materialsCitation id="3B0AD22CFF8A385A3AECF8AD9196F8DA" collectingDate="1951-11" collectionCode="RMCA" collectorName="Coll. Mus. &amp; N. Lac &amp; J. V. Leroy" country="Democratic Republic of the Congo" location="Rwankwi" pageId="14" pageNumber="335" specimenCount="1" specimenCount-male="1" stateProvince="Sud-Kivu" typeStatus="holotype">
<typeStatus id="54D966D3FF8A385A3AECF8AD9350F886" box="[464,566,1818,1840]" pageId="14" pageNumber="335" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6413F8FF8A385A3975F8AA933FF899" box="[585,601,1821,1839]" count="1" pageId="14" pageNumber="335" type="male">♂</specimenCount>
“
<collectorName id="2697BDA7FF8A385A394BF8AD9391F899" box="[631,759,1818,1840]" pageId="14" pageNumber="335">Coll. Mus.</collectorName>
<collectingCountry id="F37598E1FF8A385A3BACF88F91B1F8FB" box="[144,215,1848,1869]" name="Democratic Republic of the Congo" pageId="14" pageNumber="335">Congo</collectingCountry>
<collectorName id="2697BDA7FF8A385A3BDAF88E905CF8F8" box="[230,314,1849,1870]" pageId="14" pageNumber="335">N. Lac</collectorName>
<collectingRegion id="49A61693FF8A385A3A76F88F90E7F8F8" box="[330,385,1848,1870]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="14" pageNumber="335">Kivu</collectingRegion>
<location id="8EBD8EAAFF8A385A3AADF88F909AF8F8" LSID="urn:lsid:plazi:treatment:03CB6967FF8A385B3BA9F9C993BAFD73:8EBD8EAAFF8A385A3AADF88F909AF8F8" box="[401,508,1848,1870]" country="Democratic Republic of the Congo" name="Rwankwi" pageId="14" pageNumber="335" stateProvince="Sud-Kivu">Rwankwi</location>
<date id="FFDCFEB1FF8A385A3930F88F9300F8F8" box="[524,614,1848,1870]" pageId="14" pageNumber="335" value="1951-11">
<collectingDate id="EF980759FF8A385A3930F88F9300F8F8" box="[524,614,1848,1870]" pageId="14" pageNumber="335" value="1951-11">XI-1951</collectingDate>
</date>
<collectorName id="2697BDA7FF8A385A3953F88E938AF8FB" box="[623,748,1848,1870]" pageId="14" pageNumber="335">J.V. Leroy</collectorName>
” (
<collectionCode id="ED7340B4FF8A385A3BABF8E0918FF8DA" box="[151,233,1879,1900]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="14" pageNumber="335" type="Museum">RMCA</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54D966D3FF8A385A3BC7F8E09017F8DA" box="[251,369,1879,1900]" pageId="14" pageNumber="335" type="paratype">Paratypes</typeStatus>
: see Appendix S1.
</paragraph>
<paragraph id="8BDDD871FF8A385A381BFBB7928BF900" blockId="14.[807,1422,1023,1718]" pageId="14" pageNumber="335">
<emphasis id="B9160463FF8A385A381BFBB792C9FBA3" box="[807,943,1024,1045]" italics="true" pageId="14" pageNumber="335">Description:</emphasis>
Body length:
<quantity id="4C9A7594FF8A385A3F53FBB795A1FBA3" box="[1135,1223,1024,1045]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="335" unit="mm" value="7.7">7.7 mm</quantity>
, elytral length:
<quantity id="4C9A7594FF8A385A381BFBA99219FB82" box="[807,895,1054,1076]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="14" pageNumber="335" unit="mm" value="5.6">5.6 mm</quantity>
, maximum width: 4.0 mm. Angle between lateral margins of labroclypeus and ocular canthus indistinct; anterior margin of labroclypeus medially weakly sinuate; margins of labroclypeus strongly reflexed; smooth area in front of eyes twice as wide as long; ratio of eye diameter/interocular width: 0.77. Pronotum with dense and thick erect setae; unicoloured; basal margin of pronotum complete; larger punctures on pronotum very coarse. Hypomeron not carinate. Elytra with dark spots; erect dorsal pilosity on elytra dense (more than ten setae per interval); sutural interval in apical declivity of elytra flat and not delimited by a sharp carina; white, fine pilosity not condensed to longitudinal patches; intervals punctate at least laterally; adpressed white pilosity long, 1/3 to 1/2 of width of elytral intervals; external apical elytral angle evenly convex. Mesosternal process short. Ratio of length of metepisternum/metacoxa: 1/1.35. Metatibia, ratio of width/length: 1/4. Metatarsi dorsally smooth; first metatarsomere as long as subsequent one. Aedeagus:
<figureCitation id="1359C4F4FF8A385A3891F9369288F920" box="[941,1006,1665,1687]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">Fig. 2</figureCitation>
Ac
<emphasis id="B9160463FF8A385A3F37F9339571F920" box="[1035,1047,1668,1686]" italics="true" pageId="14" pageNumber="335">–</emphasis>
Ae. Habitus:
<figureCitation id="1359C4F4FF8A385A3FF8F9369463F921" box="[1220,1285,1665,1687]" captionStart="Figure 2" captionStartId="5.[166,245,1618,1637]" captionTargetBox="[178,1436,207,1589]" captionTargetId="figure-26@5.[163,1443,182,1596]" captionTargetPageId="5" captionText="Figure 2. A–D, Pleophylla fasciatipennis Blanchard (RSA, George); E–H, Pleophylla pilosa Boheman (RSA, Karkloof for.); I–L, Pleophylla pseudopilosa sp. nov. (holotype); M–P, Pleophylla nelshoogteensis sp. nov. (holotype); Q– T, Pleophylla mlilwaneensis sp. nov. (holotype); U–X, Pleophylla burundiensis sp. nov. (holotype); Y–Ab, Pleophylla congoensis sp. nov. (holotype); Ac–Af, Pleophylla settentrionalis sp. nov. (holotype). A, E, I, M, Q, U, Y, Ac, aedeagus left side lateral view; C, G, K, O, S, W, Aa, Ae, aedeagus right side lateral view; B, F, J, N, R, V, Z, Ad, parameres, dorsal view; D, H, L, P, T, X, Ab, Af, habitus, dorsal view. Scale bars = 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.14807087" httpUri="https://zenodo.org/record/14807087/files/figure.png" pageId="14" pageNumber="335">Fig. 2</figureCitation>
Af. Female genitalia:
<figureCitation id="1359C4F4FF8A385A38A8F917928FF900" box="[916,1001,1696,1718]" captionStart="Figure 5" captionStartId="11.[166,245,1717,1736]" captionTargetBox="[237,1338,218,1653]" captionTargetId="figure-123@11.[211,1395,195,1687]" captionTargetPageId="11" captionText="Figure 5. Female genitalia of Pleophylla species. A, Pleophylla fasciatipennis; B, Pleophylla pilosa; C, Pleophylla pseudopilosa sp. nov.; D, Pleophylla nelshoogteensis sp. nov.; E, Pleophylla burundiensis sp. nov.; F, Pleophylla settentrionalis sp. nov.; G, Pleophylla ferruginea; H, Pleophylla murzini sp. nov.; I, Pleophylla navicularis; J, Pleophylla silvatica; K, Pleophylla transkeiensis sp. nov.; L, Pleophylla ruthae sp. nov.; M, Pleophylla warnockae sp. nov.; N, Pleophylla harrisoni sp. nov.; O, Pleophylla stalsi sp. nov. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14807095" httpUri="https://zenodo.org/record/14807095/files/figure.png" pageId="14" pageNumber="335">
Fig.
<specimenCount id="9D6413F8FF8A385A38F5F917928FF900" box="[969,1001,1696,1718]" count="5" pageId="14" pageNumber="335" type="generic">5F</specimenCount>
</figureCitation>
.
</paragraph>
<paragraph id="8BDDD871FF8A385A381BF96992F4F8DB" blockId="14.[807,1422,1757,1901]" pageId="14" pageNumber="335">
<emphasis id="B9160463FF8A385A381BF96995B4F944" box="[807,1234,1757,1779]" italics="true" pageId="14" pageNumber="335">
Diagnosis:
<taxonomicName id="4C62A3F2FF8A385A3890F96A95B4F944" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[940,1234,1757,1778]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="settentrionalis" status="sp. nov.">Pleophylla settentrionalis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A225B918FF8A385A3FE1F96A945FF944" box="[1245,1337,1757,1778]" pageId="14" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
differs from the previous two species,
<taxonomicName id="4C62A3F2FF8A385A3FADF94B944CF8A7" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[1169,1322,1788,1809]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="congoensis" status="sp. nov.">
<emphasis id="B9160463FF8A385A3FADF94B944CF8A7" box="[1169,1322,1788,1809]" italics="true" pageId="14" pageNumber="335">P. congoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8A385A3E08F94B94E8F8A7" box="[1332,1422,1788,1809]" pageId="14" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C62A3F2FF8A385A3866F8AC9569F899" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[858,1039,1818,1840]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="species" species="burundiensis" status="sp. nov.">
<emphasis id="B9160463FF8A385A3866F8AC9569F899" box="[858,1039,1818,1840]" italics="true" pageId="14" pageNumber="335">P. burundiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225B918FF8A385A3F24F8AC9512F886" box="[1048,1140,1819,1840]" pageId="14" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
, by having the centre of the elytral intervals impunctate rather than densely punctate.
</paragraph>
<paragraph id="8BDDD871FF8B385B3B9AFF7093CCFE67" blockId="15.[166,781,199,465]" pageId="15" pageNumber="336">
<emphasis id="B9160463FF8B385B3B9AFF70907CFF6A" box="[166,282,199,220]" italics="true" pageId="15" pageNumber="336">Variation:</emphasis>
Body length: 7.6
<emphasis id="B9160463FF8B385B3AD2FF7E909CFF6D" box="[494,506,201,219]" italics="true" pageId="15" pageNumber="336">–</emphasis>
9.0 mm, elytral length: 5.6
<emphasis id="B9160463FF8B385B3BF4FF5F91B2FF4C" box="[200,212,232,250]" italics="true" pageId="15" pageNumber="336">–</emphasis>
7.0 mm, maximum width: 4.0
<emphasis id="B9160463FF8B385B3900FF5F932EFF4C" box="[572,584,232,250]" italics="true" pageId="15" pageNumber="336">–</emphasis>
<quantity id="4C9A7594FF8B385B3974FF5293C4FF4D" box="[584,674,229,251]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="336" unit="mm" value="4.8">4.8 mm</quantity>
. Density of dark spots may vary, sometimes completely without spots. Female: antennal club of female straight, composed of five antennomeres, slightly longer than the remaining antennomeres combined; sixth antennomere short, slightly shorter than a quarter of club length. Eyes smaller than in male: ratio of eye diameter/interocular width: 0.53.
</paragraph>
<paragraph id="8BDDD871FF8B385B3B9AFE4F9198FDDC" blockId="15.[166,781,504,618]" pageId="15" pageNumber="336">
<emphasis id="B9160463FF8B385B3B9AFE4F9042FDBB" box="[166,292,504,525]" italics="true" pageId="15" pageNumber="336">Etymology:</emphasis>
This new species is named
<taxonomicName id="4C62A3F2FF8B385B3B9AFDA0902BFD9A" authorityName="Eberle &amp; Beckett &amp; Ozgul-Siemund &amp; Frings &amp; Fabrizi &amp; Ahrens" authorityYear="2017" box="[166,333,535,556]" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="settentrionalis">
‘
<emphasis id="B9160463FF8B385B3B97FDA0902BFD9A" box="[171,333,535,556]" italics="true" pageId="15" pageNumber="336">settentrionalis</emphasis>
</taxonomicName>
’ in reference to its occurrence in the northernmost part of the known range of
<taxonomicName id="4C62A3F2FF8B385B39A9FD829198FDDC" class="Insecta" family="Scarabaeidae" genus="Pleophylla" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="336" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9160463FF8B385B39A9FD82926BFDFC" box="[661,781,565,586]" italics="true" pageId="15" pageNumber="336">Pleophylla</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="8BDDD871FF8B385B3B9AFD2693BAFD73" blockId="15.[166,780,657,709]" pageId="15" pageNumber="336">
<emphasis id="B9160463FF8B385B3B9AFD269074FD10" box="[166,274,657,678]" italics="true" pageId="15" pageNumber="336">Remarks:</emphasis>
In a previous publication we referred to this species as ‘sp27’ (Ozg € ul-Siemund € &amp;
<bibRefCitation id="EFF3A580FF8B385B390BFD1893B6FD73" author="Ahrens D" box="[567,720,687,709]" pageId="15" pageNumber="336" pagination="167 - 178" refId="ref23910" refString="Ozg € ul-Siemund € A, Ahrens D. 2015. Taxonomic utility of female copulation organs in Sericini chafers (Coleoptera, Scarabaeidae), with special reference to asymmetry. Contributions to Zoology 84 (2): 167-178." type="journal article" year="2015">Ahrens, 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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