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<mods:title id="E9E3A862666CD6D3E115939BFB61E2EE">First records of some coral reef-associated brachyuran crabs from the Nicobar archipelago, India</mods:title>
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<mods:namePart id="917BD597A29CB8133DAB33D00A2E0229">Padate, Vinay P.</mods:namePart>
<mods:affiliation id="B1DB4D17D23CB86F435E8FC6C25585DE">Centre for Marine Living Resources &amp; Ecology, Ministry of Earth Sciences, Government of India, Atal Bhavan, LNG Terminus Road, Puthuvype, Kochi, 682508, India.</mods:affiliation>
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<mods:namePart id="7E83DC4ACD96FD022E2A5C40FE7D422E">Cubelio, Sherine Sonia</mods:namePart>
<mods:affiliation id="2E35242C687D12B1747E6E96623BF045">Centre for Marine Living Resources &amp; Ecology, Ministry of Earth Sciences, Government of India, Atal Bhavan, LNG Terminus Road, Puthuvype, Kochi, 682508, India.</mods:affiliation>
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<mods:namePart id="3192CB0BBF95FE349DCEEF97BF76182F">Takeda, Masatsune</mods:namePart>
<mods:affiliation id="77F7BC032BC586132F36AF01D00CDCE2">Department of Zoology, National Museum of Nature and Science, Tokyo. 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<paragraph id="8BFC36877F20FF83FF79FABEB7D9D25D" blockId="11.[151,656,1374,1437]" box="[151,656,1374,1401]" pageId="11" pageNumber="487">
<heading id="D0B481EB7F20FF83FF79FABEB7D9D25D" bold="true" box="[151,656,1374,1401]" fontSize="11" level="1" pageId="11" pageNumber="487" reason="1">
<taxonomicName id="4C434D047F20FF83FF79FABEB7D9D25D" ID-CoL="33LG7" authority="(Rathbun, 1906)" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[151,656,1374,1401]" class="Malacostraca" family="Xanthidae" genus="Cyrtocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="487" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B937EA957F20FF83FF79FABEB7D9D25D" bold="true" box="[151,656,1374,1401]" pageId="11" pageNumber="487">
<emphasis id="B937EA957F20FF83FF79FABEB4F7D25C" bold="true" box="[151,446,1375,1401]" italics="true" pageId="11" pageNumber="487">Cyrtocarcinus truncatus</emphasis>
(
<bibRefCitation id="EFD24B767F20FF83FE20FABEB7C1D25D" author="Rathbun, M. J." box="[462,648,1374,1401]" pageId="11" pageNumber="487" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</emphasis>
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</paragraph>
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<subSubSection id="C359650C7F20FF83FF79FA62B4F1D2B8" box="[151,440,1411,1437]" pageId="11" pageNumber="487" type="description">
<paragraph id="8BFC36877F20FF83FF79FA62B4F1D2B8" blockId="11.[151,656,1374,1437]" box="[151,440,1411,1437]" pageId="11" pageNumber="487">
(
<figureCitation id="13782A027F20FF83FF71FA62B5B7D2B8" box="[159,254,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="11" pageNumber="487">Figs. 2E</figureCitation>
,
<figureCitation id="13782A027F20FF83FEE2FA62B405D2B8" box="[268,332,1411,1437]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="11" pageNumber="487">7AF</figureCitation>
,
<figureCitation id="13782A027F20FF83FEB9FA62B4E7D2B8" box="[343,430,1411,1437]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="11" pageNumber="487">10MO</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F20FF83FF79FA29B0D5D17F" pageId="11" pageNumber="487" type="reference_group">
<paragraph id="8BFC36877F20FF83FF79FA29B0D5D2FA" blockId="11.[151,1436,1479,1657]" box="[151,1436,1479,1503]" pageId="11" pageNumber="487">
<treatmentCitationGroup id="AB5311A97F20FF83FF79FA29B0D5D2FA" box="[151,1436,1479,1503]" pageId="11" pageNumber="487">
<treatmentCitation id="0AE210967F20FF83FF79FA29B768D2FA" author="Rathbun, M. J." box="[151,545,1479,1503]" page="832" pageId="11" pageNumber="487" year="1906">
<taxonomicName id="4C434D047F20FF83FF79FA29B768D2FA" authority="Rathbun, 1906: 832" authorityName="Rathbun" authorityPageNumber="832" authorityYear="1906" box="[151,545,1479,1503]" class="Malacostraca" family="Pilumnidae" genus="Harrovia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="487" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="B937EA957F20FF83FF79FA29B418D2FA" box="[151,337,1480,1503]" italics="true" pageId="11" pageNumber="487">Harrovia truncata</emphasis>
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<bibRefCitation id="EFD24B767F20FF83FEB6FA26B768D2FA" author="Rathbun, M. J." box="[344,545,1479,1503]" pageId="11" pageNumber="487" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun, 1906: 832</bibRefCitation>
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(list), 886, fig. 40, pl. 14 fig. 8 (
<typeStatus id="54F888257F20FF83FC85FA26B6D1D2FA" box="[875,920,1479,1503]" pageId="11" pageNumber="487">type</typeStatus>
locality:
<collectingMunicipality id="6B98ACFD7F20FF83FC17FA26B171D2FA" box="[1017,1080,1479,1503]" pageId="11" pageNumber="487">Kauai</collectingMunicipality>
,
<collectingRegion id="4987F8657F20FF83FBACFA26B1C7D2FA" box="[1090,1166,1479,1503]" country="United States of America" name="Hawaii" pageId="11" pageNumber="487">Hawaii</collectingRegion>
,
<emphasis id="B937EA957F20FF83FB76FA26B1B1D2FA" box="[1176,1272,1479,1503]" italics="true" pageId="11" pageNumber="487">
<location id="8E9C605C7F20FF83FB76FA26B1B1D2FA" LSID="urn:lsid:plazi:treatment:03EA87917F20FF85FF79FABEB481D4EC:8E9C605C7F20FF83FB76FA26B1B1D2FA" box="[1176,1272,1479,1503]" municipality="Kauai" name="Albatross" pageId="11" pageNumber="487" stateProvince="Hawaii">Albatross</location>
</emphasis>
stn. 3982, 233
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</paragraph>
<paragraph id="8BFC36877F20FF83FF29FA07B0D5D17F" blockId="11.[151,1436,1479,1657]" pageId="11" pageNumber="487">
240 fathoms (=
<quantity id="4CBB9B627F20FF83FE87FA07B494D2DB" box="[361,477,1510,1534]" metricMagnitude="2" metricUnit="m" metricValue="4.33" metricValueMax="4.39" metricValueMin="4.27" pageId="11" pageNumber="487" unit="m" value="433.0" valueMax="439.0" valueMin="427.0">427439 m</quantity>
) depth).—
<bibRefCitation id="EFD24B767F20FF83FDA4FA07B67BD2DB" author="Edmondson, C. H." box="[586,818,1510,1534]" pageId="11" pageNumber="487" pagination="183 - 243" refId="ref11542" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951: 217</bibRefCitation>
, figs. 21, 22.
<taxonomicName id="4C434D047F20FF83FF79F9E4B418D139" authorityName="Rathbun" authorityYear="1906" box="[151,337,1541,1564]" class="Malacostraca" family="Pilumnidae" genus="Harrovia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="487" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="B937EA957F20FF83FF79F9E4B418D139" box="[151,337,1541,1564]" italics="true" pageId="11" pageNumber="487">Harrovia truncata</emphasis>
</taxonomicName>
<bibRefCitation id="EFD24B767F20FF83FE85F9E5B743D138" author="Flipse, H. J." box="[363,522,1540,1565]" pageId="11" pageNumber="487" pagination="1 - 96" refId="ref11616" refString="Flipse, H. J. (1930) Die Decapoda Brachyura der Siboga-Expedition. VI. Oxyrhyncha: Parthenopidae. Siboga Expeditie, Monograph, 39 (c 2), 1 - 96." type="journal article" year="1930">Flipse 1930: 80</bibRefCitation>
, 90.—
<bibRefCitation id="EFD24B767F20FF83FDA5F9E4B673D139" author="Serene, R. &amp; Tran, V. D. &amp; Nguyen, V. L." box="[587,826,1540,1565]" pageId="11" pageNumber="487" pagination="135 - 242" refId="ref14841" refString="Serene, R., Tran, V. D. &amp; Nguyen, V. L. (1958) Eumedoninae du Viet-Nam (Crustacea) (avec un bibliographie de la sous-famille). Treubia, 24 (2), 135 - 242." type="journal article" year="1958">
Serène
<emphasis id="B937EA957F20FF83FD77F9E4B78CD139" box="[665,709,1540,1564]" italics="true" pageId="11" pageNumber="487">et al</emphasis>
. 1958: 196
</bibRefCitation>
, 199 (key), fig. 7A.—
<bibRefCitation id="EFD24B767F20FF83FBF6F9E4B1F6D139" author="Serene, R." box="[1048,1215,1540,1565]" pageId="11" pageNumber="487" pagination="1 - 122" refId="ref14708" refString="Serene, R. (1968) Prodromus for a check list of the non-planctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication, 1, 1 - 122." type="journal article" year="1968">Serène 1968: 63</bibRefCitation>
(list).
<taxonomicName id="4C434D047F20FF83FF79F9C2B4D0D11E" box="[151,409,1571,1595]" class="Malacostraca" family="Xanthidae" genus="Glyptocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="487" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B937EA957F20FF83FF79F9C2B4D0D11E" box="[151,409,1571,1595]" italics="true" pageId="11" pageNumber="487">Glyptocarcinus truncatus</emphasis>
</taxonomicName>
<bibRefCitation id="EFD24B767F20FF83FE5CF9C2B713D118" author="Takeda, M." box="[434,602,1571,1597]" pageId="11" pageNumber="487" pagination="68 - 72" refId="ref15307" refString="Takeda, M. (1979) Generic and specific validity of Glyptocarcinus lophopus Takeda (Crustacea, Brachyura). Proceedings of the Japanese Society of Systematic Zoology, 17, 68 - 72." type="journal article" year="1979">Takeda 1979: 68</bibRefCitation>
.—
<bibRefCitation id="EFD24B767F20FF83FD96F9C5B63FD11E" author="Stevcic, Z. &amp; Castro, P. &amp; Gore, R. H." box="[632,886,1571,1597]" pageId="11" pageNumber="487" pagination="1301 - 1324" refId="ref15251" refString="Stevcic, Z., Castro, P. &amp; Gore, R. H. (1988) Re-establishment of the family Eumedonidae Dana, 1853 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 22, 1301 - 1324. https: // doi. org / 10.1080 / 00222938800770801" type="journal article" year="1988">
Števčic
<emphasis id="B937EA957F20FF83FD25F9C5B7BED11E" box="[715,759,1571,1595]" italics="true" pageId="11" pageNumber="487">et al</emphasis>
. 1988: 1311
</bibRefCitation>
(list).
<taxonomicName id="4C434D047F20FF83FF79F9A3B4C4D17F" ID-CoL="33LG7" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[151,397,1602,1626]" class="Malacostraca" family="Xanthidae" genus="Cyrtocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="487" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B937EA957F20FF83FF79F9A3B4C4D17F" box="[151,397,1602,1626]" italics="true" pageId="11" pageNumber="487">Cyrtocarcinus truncatus</emphasis>
</taxonomicName>
<bibRefCitation id="EFD24B767F20FF83FE48F9A3B7C9D17F" author="Ng, P. K. L. &amp; Chia, D. G. B." box="[422,640,1602,1626]" pageId="11" pageNumber="487" pagination="701 - 731" refId="ref13262" refString="Ng, P. K. L. &amp; Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 731." type="journal article" year="1994">Ng &amp; Chia 1994: 725</bibRefCitation>
, figs. 1113.—
<bibRefCitation id="EFD24B767F20FF83FCF4F9A3B178D17F" author="Eldredge, L. G. &amp; Miller, S. E." box="[794,1073,1602,1626]" pageId="11" pageNumber="487" pagination="3 - 22" refId="ref11574" refString="Eldredge, L. G. &amp; Miller, S. E. (1997) Numbers of Hawaiian species: supplement 2, including a review of freshwater invertebrates. Bishop Museum Occasional Papers, 48, 3 - 22." type="journal article" year="1997">Eldredge &amp; Miller 1997: 13</bibRefCitation>
(list).—
<bibRefCitation id="EFD24B767F20FF83FB6BF9A3B00ED17F" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1157,1351,1602,1626]" pageId="11" pageNumber="487" pagination="1 - 286" refId="ref13438" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B937EA957F20FF83FB45F9A2B194D17F" box="[1195,1245,1602,1626]" italics="true" pageId="11" pageNumber="487">et al.</emphasis>
2008: 196
</bibRefCitation>
(list).—
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F20FF83FF29F980B172D15D" box="[199,1083,1632,1657]" pageId="11" pageNumber="487" type="description">
<paragraph id="8BFC36877F20FF83FF29F980B172D15D" blockId="11.[151,1436,1479,1657]" box="[199,1083,1632,1657]" pageId="11" pageNumber="487">
<bibRefCitation id="EFD24B767F20FF83FF29F980B421D15C" author="Castro, P." box="[199,360,1633,1657]" pageId="11" pageNumber="487" pagination="1 - 154" refId="ref10943" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947 (1), 1 - 154. https: // doi. org / 10.11646 / zootaxa. 2947.1.1" type="journal article" year="2011">Castro 2011: 88</bibRefCitation>
(list).—
<bibRefCitation id="EFD24B767F20FF83FE50F981B7CFD15C" author="Lai, J. C. &amp; Mendoza, J. C. E. &amp; Guinot, D. &amp; Clark, P. F. &amp; Ng, P. K. L." box="[446,646,1632,1657]" pageId="11" pageNumber="487" pagination="407 - 448" refId="ref12286" refString="Lai, J. C., Mendoza, J. C. E., Guinot, D., Clark, P. F. &amp; Ng, P. K. L. (2011) Xanthidae MacLeay, 1838 (Decapoda: Brachyura: Xanthoidea) systematics: a multi-gene approach with support from adult and zoeal morphology. Zoologischer Anzeiger, 250 (4), 407 - 448. https: // doi. org / 10.1016 / j. jcz. 2011.07.002" type="journal article" year="2011">
Lai
<emphasis id="B937EA957F20FF83FE08F980B75AD15D" box="[486,531,1632,1656]" italics="true" pageId="11" pageNumber="487">et al</emphasis>
. 2011: 440
</bibRefCitation>
, Table 1.—
<bibRefCitation id="EFD24B767F20FF83FD1BF980B686D15C" author="Naruse, T. &amp; Richer de Forges, M. &amp; Richer de Forges, B. &amp; Uyeno, D." box="[757,975,1632,1657]" pageId="11" pageNumber="487" pagination="1 - 7" refId="ref13129" refString="Naruse, T., Richer de Forges, M., Richer de Forges, B. &amp; Uyeno, D. (2020) Records of Cyrtocarcinus truncatus (Rathbun, 1906) (Decapoda: Xanthidae) from the Loyalty Islands, New Caledonia, and the Ryukyu Islands, Japan. Crustacean Research, 49, 1 - 7. https: // doi. org / 10.18353 / crustacea. 49.0 _ 33" type="journal article" year="2020">
Naruse
<emphasis id="B937EA957F20FF83FCA7F980B635D15D" box="[841,892,1632,1656]" italics="true" pageId="11" pageNumber="487">et al.</emphasis>
2020: 1
</bibRefCitation>
, figs. 15.
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F20FF83FF79F947B1A7D1C0" pageId="11" pageNumber="487" type="materials_examined">
<paragraph id="8BFC36877F20FF83FF79F947B1A7D1C0" blockId="11.[151,1436,1702,2017]" pageId="11" pageNumber="487">
<emphasis id="B937EA957F20FF83FF79F947B4C9D1E5" bold="true" box="[151,384,1702,1728]" pageId="11" pageNumber="487">Material examined.</emphasis>
<materialsCitation id="3B2B3CDA7F20FF83FE6BF946B1A3D1C0" collectingDate="2019-08-13" collectionCode="FORVSS" collectorName="V. P. Padate" country="India" elevation="69" latitude="8.44" location="Tillangchong Island" longLatPrecision="781" longitude="93.59" municipality="Nicobars" pageId="11" pageNumber="487" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D45FD0E7F20FF83FE6BF946B486D1E4" box="[389,463,1703,1729]" count="1" pageId="11" pageNumber="487" type="male">1 male</specimenCount>
(8.6 ×
<quantity id="4CBB9B627F20FF83FDF0F946B73DD1E4" box="[542,628,1703,1729]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="11" pageNumber="487" unit="mm" value="6.1">6.1 mm</quantity>
) (IO/SS/BRC/00193),
<collectionCode id="ED52AE427F20FF83FC9DF946B692D1E4" box="[883,987,1703,1729]" pageId="11" pageNumber="487">FORVSS</collectionCode>
stn. 38811, west of
<location id="8E9C605C7F20FF83FB59F946B0D1D1E4" LSID="urn:lsid:plazi:treatment:03EA87917F20FF85FF79FABEB481D4EC:8E9C605C7F20FF83FB59F946B0D1D1E4" box="[1207,1432,1703,1729]" country="India" latitude="8.44" longLatPrecision="781" longitude="93.59" municipality="Nicobars" name="Tillangchong Island" pageId="11" pageNumber="487">Tillangchong Island</location>
,
<collectingMunicipality id="6B98ACFD7F20FF83FF79F92AB5B6D1C0" box="[151,255,1739,1765]" pageId="11" pageNumber="487">Nicobars</collectingMunicipality>
,
<collectingCountry id="F35476177F20FF83FEE4F92AB40FD1C0" box="[266,326,1739,1765]" name="India" pageId="11" pageNumber="487">India</collectingCountry>
,
<geoCoordinate id="EE7750407F20FF83FEBCF92AB4ECD1C0" box="[338,421,1739,1765]" degrees="8.44" direction="north" orientation="latitude" pageId="11" pageNumber="487" precision="555" value="8.44">8.44°N</geoCoordinate>
,
<geoCoordinate id="EE7750407F20FF83FE5EF92BB742D1C0" box="[432,523,1738,1765]" degrees="93.59" direction="east" orientation="longitude" pageId="11" pageNumber="487" precision="555" value="93.59">93.59°E</geoCoordinate>
,
<elevation id="006ED1B47F20FF83FDF4F92AB7D3D1C0" box="[538,666,1739,1765]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="11" pageNumber="487" unit="m" value="69.0">
<quantity id="4CBB9B627F20FF83FDF4F92AB71AD1C0" box="[538,595,1739,1765]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="11" pageNumber="487" unit="m" value="69.0">69 m</quantity>
depth
</elevation>
, chain bag dredge, coll.
<collectorName id="26B653517F20FF83FC44F92AB163D1C0" box="[938,1066,1739,1765]" pageId="11" pageNumber="487">V.P. Padate</collectorName>
,
<date id="FFFD10477F20FF83FBDBF92AB1A3D1C0" box="[1077,1258,1739,1765]" pageId="11" pageNumber="487" value="2019-08-13">
<collectingDate id="EFB9E9AF7F20FF83FBDBF92AB1A3D1C0" box="[1077,1258,1739,1765]" pageId="11" pageNumber="487" value="2019-08-13">13 August 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F20FF85FF29F90FB60ED665" lastPageId="13" lastPageNumber="489" pageId="11" pageNumber="487" type="description">
<paragraph id="8BFC36877F20FF83FF29F90FB0C9D0BC" blockId="11.[151,1436,1702,2017]" pageId="11" pageNumber="487">
<emphasis id="B937EA957F20FF83FF29F90FB410D02D" bold="true" box="[199,345,1774,1800]" pageId="11" pageNumber="487">Description.</emphasis>
Carapace dorsal surface uneven, granular, regions 2M, 3M, 5L moderately elevated, 1P faintly bilobed, overhanging onto intestinal region, intestinal region low, flat (
<figureCitation id="13782A027F20FF83FC4BF8F2B14AD008" box="[933,1027,1811,1837]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="11" pageNumber="487">Figs. 2E</figureCitation>
,
<figureCitation id="13782A027F20FF83FBFEF8F2B17CD008" box="[1040,1077,1811,1837]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="11" pageNumber="487">7A</figureCitation>
). Frontal margin gently sinuous, slightly deflexed ventrally, bilobed by weak median fissure; supraorbital margin with 2 sutures, external orbital angle small; anterolateral margin with 3 triangular teeth, separated by sutures, second tooth largest; posterolateral margin nearly straight, with a V-shaped notch at level of cardiac protrusion; posterior margin concave (
<figureCitation id="13782A027F20FF83FAF7F89EB03AD0BC" box="[1305,1395,1919,1945]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="11" pageNumber="487">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F20FF84FF29F842B66DD7F0" blockId="11.[151,1436,1702,2017]" lastBlockId="12.[151,1437,151,537]" lastPageId="12" lastPageNumber="488" pageId="11" pageNumber="487">
Antennules folding obliquely; basal antennal article rectangular, flagellum lodged in orbital hiatus (
<figureCitation id="13782A027F20FF83FADAF842B0C7D098" box="[1332,1422,1955,1981]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="11" pageNumber="487">Fig. 7B</figureCitation>
); epistome well-developed, posterior margin carinate, with median projection and sublateral concavities (
<figureCitation id="13782A027F20FF83FADAF826B0C7D0C4" box="[1332,1422,1991,2017]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="11" pageNumber="487">Fig. 7B</figureCitation>
); buccal cavity subquadrate; mxp3 bearing papilliform setae, merus subquadrate, 0.6 times as long as ischium, with an oblique elevation separating two depressions (
<figureCitation id="13782A027F27FF84FD50FF5AB65ED7F0" box="[702,791,187,213]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="12" pageNumber="488">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F27FF84FF29FF3EB6DCD688" blockId="12.[151,1437,151,537]" pageId="12" pageNumber="488">
Chelipeds subequal, 2.5 times as long as CL; merus with strong tooth distally on lower surface, anterior margin carinate with subdistal rounded lobe, posterior margin with large subdistal tooth; palm with 4 longitudinal ridges on outer surface; dactylus slightly shorter than palm (
<figureCitation id="13782A027F27FF84FD1AFEC6B606D664" box="[756,847,295,321]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="12" pageNumber="488">Fig. 7D</figureCitation>
). Pereopods 25 slightly shorter than cheliped, meri with granular carinate margins; carpi with 2 triangular lobes on outer margin, subdistal lobe larger than subproximal lobe, propodi with triangular lobe submedially on outer margin, dactyli as long as propodi, terminating in corneous spine, inner margins of propodi and dactyli densely setose (
<figureCitation id="13782A027F27FF84FCDFFE72B6CDD688" box="[817,900,403,429]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="12" pageNumber="488">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F27FF84FF29FE56B699D53C" blockId="12.[151,1437,151,537]" pageId="12" pageNumber="488">
Male pleon subtriangular, with 6 free somites and telson, somite 3 widest, telson subtriangular, half as long as wide (
<figureCitation id="13782A027F27FF84FF31FE3AB47ED6D0" box="[223,311,475,501]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="12" pageNumber="488">Fig. 7F</figureCitation>
). G1 slender, gently curved, distal portion tapering, margins armed with short, strong spines (
<figureCitation id="13782A027F27FF84FA81FE3AB5BAD53C" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="12" pageNumber="488">Fig. 10MN</figureCitation>
); G2 0.4 times G
<quantity id="4CBB9B627F27FF84FE5BFE1EB494D53C" box="[437,477,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="488" unit="in" value="1.0">1 in</quantity>
length, distal segment triangular (
<figureCitation id="13782A027F27FF84FCB5FE1EB689D53C" box="[859,960,511,537]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="12" pageNumber="488">Fig. 10O</figureCitation>
).
</paragraph>
<caption id="DF3C660F7F27FF84FF76F864B116D0C0" ID-DOI="http://doi.org/10.5281/zenodo.13210290" ID-Zenodo-Dep="13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="12" pageNumber="488" startId="12.[152,255,1925,1949]" targetBox="[151,1436,561,1902]" targetPageId="12" targetType="figure">
<paragraph id="8BFC36877F27FF84FF76F864B116D0C0" blockId="12.[151,1436,1925,2021]" pageId="12" pageNumber="488">
<emphasis id="B937EA957F27FF84FF76F864B45CD0B8" bold="true" box="[152,277,1925,1949]" pageId="12" pageNumber="488">FIGURE 7</emphasis>
.
<taxonomicName id="4C434D047F27FF84FECAF867B798D0B8" authority="(Rathbun, 1906)" baseAuthorityName="Rathbun" baseAuthorityYear="1906" box="[292,721,1925,1949]" class="Malacostraca" family="Xanthidae" genus="Cyrtocarcinus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="488" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B937EA957F27FF84FECAF867B754D0B8" box="[292,541,1926,1949]" italics="true" pageId="12" pageNumber="488">Cyrtocarcinus truncatus</emphasis>
(
<bibRefCitation id="EFD24B767F27FF84FDC1F864B783D0B8" author="Rathbun, M. J." box="[559,714,1925,1949]" pageId="12" pageNumber="488" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</taxonomicName>
, male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm.
</paragraph>
</caption>
<paragraph id="8BFC36877F26FF85FF29FF77B60ED665" blockId="13.[151,1437,150,969]" pageId="13" pageNumber="489">
<emphasis id="B937EA957F26FF85FF29FF77B416D795" bold="true" box="[199,351,150,176]" pageId="13" pageNumber="489">Colouration.</emphasis>
Fresh specimen: Carapace dorsal surface with pinkish margins enclosing bright red colouration, cardiac regions with small median and a pair of submedian elongate whitish patches (
<figureCitation id="13782A027F26FF85FBB8FF5BB1E3D7F0" box="[1110,1194,186,213]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="13" pageNumber="489">Fig. 2E</figureCitation>
). Chelipeds dark red, distal half of fingers brown (
<figureCitation id="13782A027F26FF85FE0AFF3FB773D7DC" box="[484,570,222,249]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="13" pageNumber="489">Fig. 2E</figureCitation>
). Pereopods 23 with bright red colouration from meri to proximal half of propodi, P4 bright red on proximal half of merus, entire carpus and proximal half of propodus, and P5 bright red only on proximal half of merus; other parts whitish (
<figureCitation id="13782A027F26FF85FD0DFEC7B67FD664" box="[739,822,294,321]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="13" pageNumber="489">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F26FF85FF29FEABB1A6D6AD" pageId="13" pageNumber="489" type="biology_ecology">
<paragraph id="8BFC36877F26FF85FF29FEABB1A6D6AD" blockId="13.[151,1437,150,969]" pageId="13" pageNumber="489">
<emphasis id="B937EA957F26FF85FF29FEABB46ED641" bold="true" box="[199,295,330,356]" pageId="13" pageNumber="489">Biology.</emphasis>
The present specimens were collected along with gorgonians and sponges from
<quantity id="4CBB9B627F26FF85FB4AFEABB195D640" box="[1188,1244,330,357]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="13" pageNumber="489" unit="m" value="69.0">69 m</quantity>
depth. Known to occur at depths from
<quantity id="4CBB9B627F26FF85FE6BFE8FB4A1D6AC" box="[389,488,366,393]" metricMagnitude="2" metricUnit="m" metricValue="2.21" metricValueMax="4.39" metricValueMin="0.03" pageId="13" pageNumber="489" unit="m" value="221.0" valueMax="439.0" valueMin="3.0">3439 m</quantity>
(
<bibRefCitation id="EFD24B767F26FF85FE16FE8FB7D0D6AD" author="Rathbun, M. J." box="[504,665,366,392]" pageId="13" pageNumber="489" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
;
<bibRefCitation id="EFD24B767F26FF85FD4BFE8FB664D6AD" author="Castro, P." box="[677,813,366,392]" pageId="13" pageNumber="489" pagination="1 - 154" refId="ref10943" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947 (1), 1 - 154. https: // doi. org / 10.11646 / zootaxa. 2947.1.1" type="journal article" year="2011">Castro 2011</bibRefCitation>
) on rocky substrates (
<bibRefCitation id="EFD24B767F26FF85FBCDFE8FB1AAD6AD" author="Ng, P. K. L. &amp; Chia, D. G. B." box="[1059,1251,366,392]" pageId="13" pageNumber="489" pagination="701 - 731" refId="ref13262" refString="Ng, P. K. L. &amp; Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 731." type="journal article" year="1994">Ng &amp; Chia 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F26FF85FF29FE73B481D4EC" pageId="13" pageNumber="489" type="discussion">
<paragraph id="8BFC36877F26FF85FF29FE73B43DD41D" blockId="13.[151,1437,150,969]" pageId="13" pageNumber="489">
<emphasis id="B937EA957F26FF85FF29FE73B472D689" bold="true" box="[199,315,402,428]" pageId="13" pageNumber="489">Remarks.</emphasis>
<taxonomicName id="4C434D047F26FF85FEADFE72B75BD689" authorityName="Rathbun" authorityYear="1906" box="[323,530,403,428]" class="Malacostraca" family="Pilumnidae" genus="Harrovia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="489" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="B937EA957F26FF85FEADFE72B75BD689" box="[323,530,403,428]" italics="true" pageId="13" pageNumber="489">Harrovia truncata</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFD24B767F26FF85FD0BFE73B6DED689" author="Rathbun, M. J." box="[741,919,402,428]" pageId="13" pageNumber="489" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun (1906)</bibRefCitation>
from an immature male collected from deepwaters off Kauai, Hawaiian Islands. The Indian specimen resembles the description of the
<typeStatus id="54F888257F26FF85FB6AFE57B1AFD6F5" box="[1156,1254,438,464]" pageId="13" pageNumber="489" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFD24B767F26FF85FB18FE57B0DDD6F5" author="Rathbun, M. J." box="[1270,1428,438,464]" pageId="13" pageNumber="489" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
) in the morphology of the carapace, chelipeds and pereopods.
<bibRefCitation id="EFD24B767F26FF85FCBAFE3AB16CD6D1" author="Ng, P. K. L. &amp; Chia, D. G. B." box="[852,1061,474,501]" pageId="13" pageNumber="489" pagination="701 - 731" refId="ref13262" refString="Ng, P. K. L. &amp; Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 731." type="journal article" year="1994">Ng &amp; Chia (1994)</bibRefCitation>
transferred this species to a new genus,
<taxonomicName id="4C434D047F26FF85FF08FE1EB4CCD53C" authorityName="Ng &amp; Chia" authorityYear="1994" box="[230,389,511,537]" class="Malacostraca" family="Xanthidae" genus="Cyrtocarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA957F26FF85FF08FE1EB4CCD53C" box="[230,389,511,537]" italics="true" pageId="13" pageNumber="489">Cyrtocarcinus</emphasis>
</taxonomicName>
, which differed from
<taxonomicName id="4C434D047F26FF85FD96FE1EB692D53D" authority="Ng &amp; Chia, 1994" authorityName="Ng &amp; Chia" authorityYear="1994" box="[632,987,510,537]" class="Malacostraca" family="Xanthidae" genus="Antrocarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA957F26FF85FD96FE1EB65FD53C" box="[632,790,511,537]" italics="true" pageId="13" pageNumber="489">Antrocarcinus</emphasis>
<bibRefCitation id="EFD24B767F26FF85FCF2FE1EB692D53D" author="Ng, P. K. L. &amp; Chia, D. G. B." box="[796,987,510,537]" pageId="13" pageNumber="489" pagination="701 - 731" refId="ref13262" refString="Ng, P. K. L. &amp; Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 731." type="journal article" year="1994">Ng &amp; Chia, 1994</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C434D047F26FF85FBFEFE1FB011D53D" authority="Takeda, 1973" authorityName="Takeda" authorityYear="1973" box="[1040,1368,510,536]" class="Malacostraca" family="Xanthidae" genus="Glyptocarcinus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="489" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA957F26FF85FBFEFE1FB1F5D53D" box="[1040,1212,510,536]" italics="true" pageId="13" pageNumber="489">Glyptocarcinus</emphasis>
Takeda,
<quantity id="4CBB9B627F26FF85FACEFE1FB011D53D" box="[1312,1368,510,536]" metricMagnitude="1" metricUnit="m" metricValue="5.011419999999999" pageId="13" pageNumber="489" unit="in" value="1973.0">1973</quantity>
</taxonomicName>
in the presence of a distinct ridge and a depression on the fourth thoracic sternite formed by an abrupt downward sloping of its posterior part. The Indian specimen possesses a larger third tooth on the lateral carapace margin (
<figureCitation id="13782A027F26FF85FADAFDA7B0C6D545" box="[1332,1423,582,608]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="13" pageNumber="489">Fig. 7A</figureCitation>
), as compared to the
<typeStatus id="54F888257F26FF85FE9DFD8BB49CD5A1" box="[371,469,618,644]" pageId="13" pageNumber="489" type="holotype">holotype</typeStatus>
male (6.5 ×
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;
<bibRefCitation id="EFD24B767F26FF85FD24FD8AB6C0D5A1" author="Ng, P. K. L. &amp; Chia, D. G. B." box="[714,905,618,645]" pageId="13" pageNumber="489" pagination="701 - 731" refId="ref13262" refString="Ng, P. K. L. &amp; Chia, D. G. B. (1994) The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42 (3), 701 - 731." type="journal article" year="1994">Ng &amp; Chia 1994</bibRefCitation>
: figs. 11A, 12A), which according to
<bibRefCitation id="EFD24B767F26FF85FADFFD8AB443D58D" author="Naruse, T. &amp; Richer de Forges, M. &amp; Richer de Forges, B. &amp; Uyeno, D." pageId="13" pageNumber="489" pagination="1 - 7" refId="ref13129" refString="Naruse, T., Richer de Forges, M., Richer de Forges, B. &amp; Uyeno, D. (2020) Records of Cyrtocarcinus truncatus (Rathbun, 1906) (Decapoda: Xanthidae) from the Loyalty Islands, New Caledonia, and the Ryukyu Islands, Japan. Crustacean Research, 49, 1 - 7. https: // doi. org / 10.18353 / crustacea. 49.0 _ 33" type="journal article" year="2020">
Naruse
<emphasis id="B937EA957F26FF85FA66FD8AB5FDD58D" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2020)
</bibRefCitation>
, could be attributed to the difference in their carapace size. The Indian specimen closely resembles the morphology and colouration of the Japanese female specimen reported by
<bibRefCitation id="EFD24B767F26FF85FC1AFD52B19ED5E9" author="Naruse, T. &amp; Richer de Forges, M. &amp; Richer de Forges, B. &amp; Uyeno, D." box="[1012,1239,690,717]" pageId="13" pageNumber="489" pagination="1 - 7" refId="ref13129" refString="Naruse, T., Richer de Forges, M., Richer de Forges, B. &amp; Uyeno, D. (2020) Records of Cyrtocarcinus truncatus (Rathbun, 1906) (Decapoda: Xanthidae) from the Loyalty Islands, New Caledonia, and the Ryukyu Islands, Japan. Crustacean Research, 49, 1 - 7. https: // doi. org / 10.18353 / crustacea. 49.0 _ 33" type="journal article" year="2020">
Naruse
<emphasis id="B937EA957F26FF85FBA3FD52B1C1D5E9" box="[1101,1160,690,716]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2020
</bibRefCitation>
: fig. 1C). On the other hand, the Indian specimen lacks a subproximal tooth on the outer surface of cheliped merus (
<figureCitation id="13782A027F26FF85FB0BFD37B074D5D5" box="[1253,1341,726,752]" captionStart="FIGURE 7" captionStartId="12.[152,255,1925,1949]" captionTargetBox="[151,1436,561,1902]" captionTargetId="figure-191@12.[151,1436,561,1902]" captionTargetPageId="12" captionText="FIGURE 7. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, frontal view; (C) right mxp3, ventral view; (D) left cheliped, lateral view; (E) left P2, dorsal view; (F) male pleon and telson, ventral view. Scale bars: AB, F, 2 mm, C, E, 1 mm, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210290" httpUri="https://zenodo.org/record/13210290/files/figure.png" pageId="13" pageNumber="489">Fig. 7D</figureCitation>
) (versus cheliped merus with a subproximal tooth on outer surface in the New Caledonian specimen reported by
<bibRefCitation id="EFD24B767F26FF85FADFFD1AB448D41D" author="Naruse, T. &amp; Richer de Forges, M. &amp; Richer de Forges, B. &amp; Uyeno, D." pageId="13" pageNumber="489" pagination="1 - 7" refId="ref13129" refString="Naruse, T., Richer de Forges, M., Richer de Forges, B. &amp; Uyeno, D. (2020) Records of Cyrtocarcinus truncatus (Rathbun, 1906) (Decapoda: Xanthidae) from the Loyalty Islands, New Caledonia, and the Ryukyu Islands, Japan. Crustacean Research, 49, 1 - 7. https: // doi. org / 10.18353 / crustacea. 49.0 _ 33" type="journal article" year="2020">
Naruse
<emphasis id="B937EA957F26FF85FA66FD1AB5FDD41D" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2020
</bibRefCitation>
: fig. 2A)).
</paragraph>
<paragraph id="8BFC36877F26FF85FF29FCA3B481D4EC" blockId="13.[151,1437,150,969]" pageId="13" pageNumber="489">
<emphasis id="B937EA957F26FF85FF29FCA3B74FD479" bold="true" box="[199,518,834,860]" pageId="13" pageNumber="489">Geographical distribution.</emphasis>
Hawaiian Islands (
<bibRefCitation id="EFD24B767F26FF85FD0FFCA3B6CAD479" author="Rathbun, M. J." box="[737,899,834,860]" pageId="13" pageNumber="489" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
;
<bibRefCitation id="EFD24B767F26FF85FC61FCA3B111D478" author="Edmondson, C. H." box="[911,1112,834,861]" pageId="13" pageNumber="489" pagination="183 - 243" refId="ref11542" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951</bibRefCitation>
;
<bibRefCitation id="EFD24B767F26FF85FB8AFCA3B1BED479" author="Takeda, M." box="[1124,1271,834,860]" pageId="13" pageNumber="489" pagination="68 - 72" refId="ref15307" refString="Takeda, M. (1979) Generic and specific validity of Glyptocarcinus lophopus Takeda (Crustacea, Brachyura). Proceedings of the Japanese Society of Systematic Zoology, 17, 68 - 72." type="journal article" year="1979">Takeda 1979</bibRefCitation>
;
<bibRefCitation id="EFD24B767F26FF85FAEAFCA3B0D9D479" author="Castro, P." box="[1284,1424,834,860]" pageId="13" pageNumber="489" pagination="1 - 154" refId="ref10943" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947 (1), 1 - 154. https: // doi. org / 10.11646 / zootaxa. 2947.1.1" type="journal article" year="2011">Castro 2011</bibRefCitation>
),
<collectingRegion id="4987F8657F26FF85FF79FC87B402D4A5" box="[151,331,870,896]" country="New Caledonia" name="Province des Iles" pageId="13" pageNumber="489">Loyalty Islands</collectingRegion>
,
<collectingCountry id="F35476177F26FF85FEB4FC86B745D4A5" box="[346,524,870,897]" name="New Caledonia" pageId="13" pageNumber="489">New Caledonia</collectingCountry>
(
<bibRefCitation id="EFD24B767F26FF85FDF1FC87B787D4A5" author="Lai, J. C. &amp; Mendoza, J. C. E. &amp; Guinot, D. &amp; Clark, P. F. &amp; Ng, P. K. L." box="[543,718,870,896]" pageId="13" pageNumber="489" pagination="407 - 448" refId="ref12286" refString="Lai, J. C., Mendoza, J. C. E., Guinot, D., Clark, P. F. &amp; Ng, P. K. L. (2011) Xanthidae MacLeay, 1838 (Decapoda: Brachyura: Xanthoidea) systematics: a multi-gene approach with support from adult and zoeal morphology. Zoologischer Anzeiger, 250 (4), 407 - 448. https: // doi. org / 10.1016 / j. jcz. 2011.07.002" type="journal article" year="2011">
Lai
<emphasis id="B937EA957F26FF85FDA1FC86B7C2D4A5" box="[591,651,870,896]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
2011
</bibRefCitation>
) and Ishigaki Island,
<collectingCountry id="F35476177F26FF85FC21FC86B146D4A4" box="[975,1039,871,897]" name="Japan" pageId="13" pageNumber="489">Japan</collectingCountry>
(
<bibRefCitation id="EFD24B767F26FF85FBCEFC87B049D4A5" author="Naruse, T. &amp; Richer de Forges, M. &amp; Richer de Forges, B. &amp; Uyeno, D." box="[1056,1280,870,896]" pageId="13" pageNumber="489" pagination="1 - 7" refId="ref13129" refString="Naruse, T., Richer de Forges, M., Richer de Forges, B. &amp; Uyeno, D. (2020) Records of Cyrtocarcinus truncatus (Rathbun, 1906) (Decapoda: Xanthidae) from the Loyalty Islands, New Caledonia, and the Ryukyu Islands, Japan. Crustacean Research, 49, 1 - 7. https: // doi. org / 10.18353 / crustacea. 49.0 _ 33" type="journal article" year="2020">
Naruse
<emphasis id="B937EA957F26FF85FB92FC86B1F1D4A5" box="[1148,1208,870,896]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
2020
</bibRefCitation>
). The present observation is the first record of the genus and species from Indian Ocean indicating westward extension of the known geographical range.
</paragraph>
</subSubSection>
</treatment>
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