150 lines
20 KiB
XML
150 lines
20 KiB
XML
<document id="A5EDC636B263D923AB9CF10635267F05" ID-DOI="10.5281/zenodo.185055" ID-GBIF-Dataset="52c2f57f-276a-4340-a2e3-2c101fb70347" ID-ISSN="1175-5326" ID-Zenodo-Dep="185055" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459950849737" checkinUser="plazi" docAuthor="Duan, Yani & Zhang, Yalin" docDate="2009" docId="921787AA7852FFF795EFFD3CFBF6DF29" docLanguage="en" docName="zt01972p043.pdf" docOrigin="Zootaxa 1972" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leofa (Prasutagus) Distant" docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="6E2EFFD27852FFF69578FA0BFFFBDB0B" masterDocTitle="Review of the grassland leafhopper subgenus Leofa (Prasutagus) Distant (Hemiptera: Cicadellidae: Deltocephalinae: Chiasmini)" masterLastPageNumber="43" masterPageNumber="35" pageNumber="35" updateTime="1698230806230" updateUser="plazi">
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<mods:mods id="2E18DCC823BA95CD749B92F1225B001A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9C7A6938628F4326D1291C63CF974AA5">
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<mods:title id="CBD71B1B32ACA1586018D34603CFEC50">Review of the grassland leafhopper subgenus Leofa (Prasutagus) Distant (Hemiptera: Cicadellidae: Deltocephalinae: Chiasmini)</mods:title>
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<mods:name id="D4F3FFC68C3D6340E97BB2E2BC30FBDF" type="personal">
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<mods:roleTerm id="31AF419E468A0092DC0048E34D526C79">Author</mods:roleTerm>
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<mods:namePart id="AB751556DE0A93347DE71C6E39519E18">Duan, Yani</mods:namePart>
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<mods:name id="1B442D6797D631A06D1AB1A4097F7E3C" type="personal">
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<mods:role id="F3F11AE36912E81C16E2C2F3EA652C73">
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<mods:roleTerm id="FC1EB3A11F8DDE9AE87A5EA8E2B15CE4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="29215D4099EEAEF91370C12D77713B9A">Zhang, Yalin</mods:namePart>
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<mods:typeOfResource id="13EB3EDC0DE9AC904C83ACAB26DB532E">text</mods:typeOfResource>
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<mods:titleInfo id="3FDFF230CEAB527D63CD8709DEA5440C">
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<mods:title id="4487C773C741A87E30C0F15027EBA58E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="94C6B6F80BBF994779B70AC08F5E1901">
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<mods:date id="F91BED1BEAE6B4E056B2DB6ABAA96139">2009</mods:date>
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<mods:detail id="E7ED72D3ECDE8792970711DF9519B698" type="volume">
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<mods:number id="60301D7A8B34AEA3AF232ECD19827CA3">1972</mods:number>
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<mods:extent id="A3E0FEEBB91C7A8573AAD0898A074806" unit="page">
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<mods:start id="E0994E33A2FF2C03FBDC09F4C1533AD3">35</mods:start>
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<mods:end id="85FBB97B3F6F2C6A8A81EC767000F2EF">43</mods:end>
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<mods:classification id="A2A0D1E8A6EAFAB549D8F0F3E84904FF">journal article</mods:classification>
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<mods:identifier id="21CDCA9BAB26EA36FC3FB7F9913F444C" type="DOI">10.5281/zenodo.185055</mods:identifier>
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<mods:identifier id="47537553A95D78F32E6279864D5AE77B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EC6C25656C20B8CC9BD1C89D52ECF2C6" type="Zenodo-Dep">185055</mods:identifier>
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<treatment id="921787AA7852FFF795EFFD3CFBF6DF29" ID-DOI="http://doi.org/10.5281/zenodo.6214625" ID-GBIF-Taxon="119376518" ID-Zenodo-Dep="6214625" LSID="urn:lsid:plazi:treatment:921787AA7852FFF795EFFD3CFBF6DF29" httpUri="http://treatment.plazi.org/id/921787AA7852FFF795EFFD3CFBF6DF29" lastPageId="1" lastPageNumber="36" pageId="0" pageNumber="35">
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<subSubSection id="52A465377852FFF695EFFD3CFC47DC92" pageId="0" pageNumber="35" type="nomenclature">
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<paragraph id="1A0136BC7852FFF695EFFD3CFE19DC5A" blockId="0.[151,482,1847,1873]" box="[151,482,1847,1873]" pageId="0" pageNumber="35">
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<heading id="414981D07852FFF695EFFD3CFE19DC5A" bold="true" box="[151,482,1847,1873]" fontSize="11" level="1" pageId="0" pageNumber="35" reason="1">
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<taxonomicName id="DDBE4D3F7852FFF695EFFD3CFE19DC5A" ID-CoL="8JMTL" authority="Distant" authorityName="Distant" box="[151,482,1847,1873]" class="Insecta" family="Cicadellidae" genus="Leofa" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="subGenus" subGenus="Prasutagus">
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<emphasis id="28CAEAAE7852FFF695EFFD3CFE19DC5A" bold="true" box="[151,482,1847,1873]" pageId="0" pageNumber="35">
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<emphasis id="28CAEAAE7852FFF695EFFD3CFF26DC5A" bold="true" box="[151,221,1847,1873]" italics="true" pageId="0" pageNumber="35">Leofa</emphasis>
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(
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<emphasis id="28CAEAAE7852FFF69596FD3CFE8DDC5A" bold="true" box="[238,374,1847,1873]" italics="true" pageId="0" pageNumber="35">Prasutagus</emphasis>
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) Distant
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="1A0136BC7852FFF695EFFD89FC47DC92" blockId="0.[151,1437,1922,2009]" box="[151,956,1922,1945]" pageId="0" pageNumber="35">
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<emphasis id="28CAEAAE7852FFF695EFFD89FEEADC92" box="[151,273,1922,1945]" italics="true" pageId="0" pageNumber="35">Prasutagus</emphasis>
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<bibRefCitation id="7E2F4B4D7852FFF6946FFD89FE51DC92" author="Distant" box="[279,426,1922,1945]" pageId="0" pageNumber="43" refString="Distant, W. L. (1918) The fauna of British India, including Ceylon and Burma, Rhynchota, VII. Homoptera: Appendix, Heteroptera: Addenda. Published under the authority of the Secretary of State for India in Council. Taylor & Francis, London. pp i - vii, 1 - 210." type="book chapter" year="1918">Distant, 1918</bibRefCitation>
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: 53.
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<typeStatus id="C505881E7852FFF694A4FD89FDEADC92" box="[476,529,1922,1945]" pageId="0" pageNumber="35">Type</typeStatus>
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species:
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<taxonomicName id="DDBE4D3F7852FFF6970BFD89FC47DC92" authority="Distant." authorityName="Distant." box="[627,956,1922,1945]" class="Insecta" family="Cicadellidae" genus="Prasutagus" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis id="28CAEAAE7852FFF6970BFD89FC9ADC92" box="[627,865,1922,1945]" italics="true" pageId="0" pageNumber="35">Prasutagus pulchellus</emphasis>
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Distant.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="52A465377852FFF795EFFDA9FC52D839" lastPageId="1" lastPageNumber="36" pageId="0" pageNumber="35" type="description">
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<paragraph id="1A0136BC7852FFF695EFFDA9FD62DCD2" blockId="0.[151,1437,1922,2009]" pageId="0" pageNumber="35">
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<emphasis id="28CAEAAE7852FFF695EFFDA9FEF4DCB2" box="[151,271,1954,1977]" italics="true" pageId="0" pageNumber="35">Oneratulus</emphasis>
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<bibRefCitation id="7E2F4B4D7852FFF6946DFDA9FE49DCB2" author="Vilbaste" box="[277,434,1954,1977]" pageId="0" pageNumber="43" refString="Vilbaste, J. (1975) On some species of Homoptera cicadinea described by Motschulsky. Ecoli NSV Teaduste Akadeemia (Biologia), 24 (3), 228 - 236." type="journal article" year="1975">Vilbaste, 1975</bibRefCitation>
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: 233–244.
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<typeStatus id="C505881E7852FFF6975EFDA9FDA0DCB2" box="[550,603,1954,1977]" pageId="0" pageNumber="35">Type</typeStatus>
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species:
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<taxonomicName id="DDBE4D3F7852FFF697C4FDA9FCFADCB2" box="[700,769,1954,1977]" class="Insecta" family="Cicadellidae" genus="Jassus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">
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<emphasis id="28CAEAAE7852FFF697C4FDA9FCFADCB2" box="[700,769,1954,1977]" italics="true" pageId="0" pageNumber="35">Jassus</emphasis>
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</taxonomicName>
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?
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<taxonomicName id="DDBE4D3F7852FFF6966BFDA9FBA1DCB2" authority="Motschulsky. Implicit" authorityName="Motschulsky. Implicit" box="[787,1114,1954,1977]" class="Insecta" family="Cicadellidae" genus="Leofa" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="curtulus">
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<emphasis id="28CAEAAE7852FFF6966BFDA9FC90DCB2" box="[787,875,1954,1977]" italics="true" pageId="0" pageNumber="35">curtulus</emphasis>
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Motschulsky. Implicit
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</taxonomicName>
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synonymy by
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<bibRefCitation id="7E2F4B4D7852FFF6918EFDA9FA62DCB2" author="Zahniser" box="[1270,1433,1954,1977]" pageId="0" pageNumber="43" refString="Zahniser, J. N. (2008) Seven new species and new distributions of Old World Chiasmini (Hemiptera: Cicadellidae: Deltocephalinae), with a redescription, key to genera, and species. Zootaxa, 1808, 1 - 32." type="journal article" year="2008">Zahniser, 2008</bibRefCitation>
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: 31.
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<taxonomicName id="DDBE4D3F7852FFF69597FDC9FD6EDCD2" authority="Zahniser, 2008: 16" authorityName="Zahniser" authorityPageNumber="16" authorityYear="2008" box="[239,661,1986,2009]" class="Insecta" family="Cicadellidae" genus="Leofa" kingdom="Animalia" order="Hemiptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="subGenus" subGenus="Prasutagus">
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<emphasis id="28CAEAAE7852FFF69597FDC9FED7DCD2" box="[239,300,1986,2009]" italics="true" pageId="0" pageNumber="35">Leofa</emphasis>
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(
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<emphasis id="28CAEAAE7852FFF69443FDC9FE4EDCD2" box="[315,437,1986,2009]" italics="true" pageId="0" pageNumber="35">Prasutagus</emphasis>
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),
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<bibRefCitation id="7E2F4B4D7852FFF694B3FDC9FD96DCD2" author="Zahniser" box="[459,621,1986,2009]" pageId="0" pageNumber="43" refString="Zahniser, J. N. (2008) Seven new species and new distributions of Old World Chiasmini (Hemiptera: Cicadellidae: Deltocephalinae), with a redescription, key to genera, and species. Zootaxa, 1808, 1 - 32." type="journal article" year="2008">Zahniser, 2008</bibRefCitation>
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: 16
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="1A0136BC7853FFF795EFFA9CFBB0DA71" blockId="1.[151,1437,151,1058]" pageId="1" pageNumber="36">
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<emphasis id="28CAEAAE7853FFF795EFFA9CFED4DBBA" bold="true" box="[151,303,151,177]" pageId="1" pageNumber="36">Description.</emphasis>
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Ochraceous with face dark brown; forewing veins white to translucent, usually bordered with brown, clavus with distinct white stripe. Head with eyes as wide as pronotum or slightly wider and distinctly longer than pronotum. Face with frontoclypeus somewhat longer than wide; anteclypeus narrowed slightly apically. Ocelli small, less than their own diameter from eye. Forewings submacropterous, truncate; appendix absent; tip of wing reaching to apex of abdominal tergite VI to exceeding tergite VIII; outer anteapical cell very small and constricted or completely absent. Hindwings completely reduced.
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</paragraph>
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<paragraph id="1A0136BC7853FFF795BDFB8CFC12D9E9" blockId="1.[151,1437,151,1058]" pageId="1" pageNumber="36">
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<emphasis id="28CAEAAE7853FFF795BDFB8CFE7ADAAA" bold="true" box="[197,385,391,417]" pageId="1" pageNumber="36">Male genitalia.</emphasis>
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Pygofer without membranous cleft; caudal lobes with dorsal margin abruptly narrowed near base of anal tube and obliquely sloping posteriorly, two lobe-like areas thus formed with internal sclerotised area and few long macrosetae; large inner appendage arising from base of upper lobe and extending to apex of pygofer. Valve triangular. Subgenital plate triangular with slightly obliquely truncate apex and undulated lateral margin; few irregularly arranged macrosetae apically. Style relatively narrow; apical apophysis curved laterally and tapered to acute apex, foot-like, in dorsal view; lateral lobe weakly to well developed. Connective Y-shaped with stem very short. Aedeagus with shaft cylindrical to laterally compressed, with one or more processes and dorsolateral flange basally, gonopore apical; basal apodeme very short. Elongate, recurved unpaired process articulated between aedeagus and connective.
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</paragraph>
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<paragraph id="1A0136BC7853FFF795BDF8E4FC52D839" blockId="1.[151,1437,151,1058]" pageId="1" pageNumber="36">
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<emphasis id="28CAEAAE7853FFF795BDF8E4FE61D802" bold="true" box="[197,410,751,777]" pageId="1" pageNumber="36">Female genitalia.</emphasis>
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Pygofer almost as long as rest of abdomen.
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<collectingCountry id="62A9762C7853FFF796DBF8FBFC25D801" box="[931,990,752,778]" name="India" pageId="1" pageNumber="36">Hind</collectingCountry>
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margin of VII sternite somewhat protruded in middle. Ovipositor projecting well beyond tip of pygofer.
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</paragraph>
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</subSubSection>
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<subSubSection id="52A465377853FFF795BDF934FCF8D851" box="[197,771,831,858]" pageId="1" pageNumber="36" type="distribution">
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<paragraph id="1A0136BC7853FFF795BDF934FCF8D851" blockId="1.[151,1437,151,1058]" box="[197,771,831,858]" pageId="1" pageNumber="36">
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<emphasis id="28CAEAAE7853FFF795BDF934FE9DD852" bold="true" box="[197,358,831,857]" pageId="1" pageNumber="36">Distribution.</emphasis>
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<collectingCountry id="62A9762C7853FFF79415F94BFE19D851" box="[365,482,832,858]" name="Sri Lanka" pageId="1" pageNumber="36">Sri Lanka</collectingCountry>
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,
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<collectingCountry id="62A9762C7853FFF79496F94BFDD6D851" box="[494,557,832,858]" name="India" pageId="1" pageNumber="36">India</collectingCountry>
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,
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<collectingCountry id="62A9762C7853FFF79741F94BFD84D851" box="[569,639,832,858]" name="China" pageId="1" pageNumber="36">China</collectingCountry>
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and
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<collectingCountry id="62A9762C7853FFF797C0F94BFD04D851" box="[696,767,832,858]" name="Spain" pageId="1" pageNumber="36">Spain</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="52A465377853FFF795BEF96CFBF6DF29" pageId="1" pageNumber="36" type="discussion">
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<paragraph id="1A0136BC7853FFF795BEF96CFBF6DF29" blockId="1.[151,1437,151,1058]" pageId="1" pageNumber="36">
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<emphasis id="28CAEAAE7853FFF795BEF96CFEBBD88A" bold="true" box="[198,320,871,897]" pageId="1" pageNumber="36">Remarks.</emphasis>
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Members of the subgenus
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<taxonomicName id="DDBE4D3F7853FFF79706F963FC99D889" authority="Prasutagus" authorityName="Prasutagus" box="[638,866,872,898]" class="Insecta" family="Cicadellidae" genus="Leofa" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="28CAEAAE7853FFF79706F963FD3AD88A" box="[638,705,872,897]" italics="true" pageId="1" pageNumber="36">Leofa</emphasis>
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(
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<emphasis id="28CAEAAE7853FFF797AAF963FCACD88A" box="[722,855,872,897]" italics="true" pageId="1" pageNumber="36">Prasutagus</emphasis>
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)
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</taxonomicName>
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are distinguished by the slightly shortened forewings (submacropterous) with a white stripe on the clavus (
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<figureCitation id="82852A397853FFF79637F99BFC57D8A1" box="[847,940,912,938]" captionStart="FIGURE 1. A – J" captionStartId="2.[151,255,1807,1831]" captionTargetBox="[284,1273,209,1720]" captionTargetId="figure@2.[226,1360,197,1761]" captionTargetPageId="2" captionText="FIGURE 1. A – J, Leofa (Prasutagus) species. A – H, Leofa (Prasutagus) pulchellus Distant (A – G, India H, China), A: head and thorax; B: face; C: male pygofer, lateral view; D: subgenital plate, style and connective, dorsal view; E: aedeagus and paraphysis dorsal view; F: aedeagus and paraphysis, left lateral view; G: aedeagus ventral view; H: apex of left pygofer process; I – J: Leofa (Prasutagus) curtulus (Motschulsky), I, aedeagus, left lateral view (type male, Jassus? curtulus Motschulsky, from Vilbaste, 1975); J, dorsal habitus (syntype male Deltocephalus oneratus Melichar, from Distant, 1908)." httpUri="https://zenodo.org/record/185056/files/figure.png" pageId="1" pageNumber="36">Figs. 1J</figureCitation>
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,
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<figureCitation id="82852A397853FFF796C1F99BFBF4D8A1" box="[953,1039,912,938]" captionStart="FIGURE 4. A, B" captionStartId="6.[151,255,1954,1978]" captionTargetBox="[250,1323,684,1891]" captionTargetId="figure@6.[226,1360,677,1900]" captionTargetPageId="6" captionText="FIGURE 4. A, B. Leofa (Prasutagus) yangae Duan and Zhang sp. n. A: dorsal habitus (♂); B: dorsal habitus (Ψ)." httpUri="https://zenodo.org/record/185059/files/figure.png" pageId="1" pageNumber="36">4A, 4B</figureCitation>
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) and the pygofer side with a large inner appendage arising from the base of the upper lobe that extends to the apex of the pygofer (
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<figureCitation id="82852A397853FFF79185F9B3FA98D8D9" box="[1277,1379,952,978]" captionStart="FIGURE 1. A – J" captionStartId="2.[151,255,1807,1831]" captionTargetBox="[284,1273,209,1720]" captionTargetId="figure@2.[226,1360,197,1761]" captionTargetPageId="2" captionText="FIGURE 1. A – J, Leofa (Prasutagus) species. A – H, Leofa (Prasutagus) pulchellus Distant (A – G, India H, China), A: head and thorax; B: face; C: male pygofer, lateral view; D: subgenital plate, style and connective, dorsal view; E: aedeagus and paraphysis dorsal view; F: aedeagus and paraphysis, left lateral view; G: aedeagus ventral view; H: apex of left pygofer process; I – J: Leofa (Prasutagus) curtulus (Motschulsky), I, aedeagus, left lateral view (type male, Jassus? curtulus Motschulsky, from Vilbaste, 1975); J, dorsal habitus (syntype male Deltocephalus oneratus Melichar, from Distant, 1908)." httpUri="https://zenodo.org/record/185056/files/figure.png" pageId="1" pageNumber="36">Figs. 1C</figureCitation>
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,
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<figureCitation id="82852A397853FFF79008F9B3FA6CD8D9" box="[1392,1431,952,978]" captionStart="FIGURE 3. A – H" captionStartId="5.[151,255,1388,1412]" captionTargetBox="[247,1339,203,1338]" captionTargetId="figure@5.[226,1360,197,1362]" captionTargetPageId="5" captionText="FIGURE 3. A – H, Leofa (Prasutagus) forcipatus Duan and Zhang sp. n. A: pygofer, lateral view; B: apex of pygofer appendage, lateral view; C: valve and subgenital plate, ventral view; D: style, dorsal view; E: aedeagus, paraphysis and connective, lateral view; F: aedeagus, posterior view; G: connective, dorsal view; H: female genitalia." httpUri="https://zenodo.org/record/185058/files/figure.png" pageId="1" pageNumber="36">3A</figureCitation>
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,
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<figureCitation id="82852A397853FFF795EFF9EBFF47D8F1" box="[151,188,992,1018]" captionStart-0="FIGURE 4. A, B" captionStart-1="FIGURE 4. C – K" captionStartId-0="6.[151,255,1954,1978]" captionStartId-1="7.[151,255,1532,1556]" captionTargetBox-0="[250,1323,684,1891]" captionTargetBox-1="[237,1339,209,1474]" captionTargetId-0="figure@6.[226,1360,677,1900]" captionTargetId-1="figure@7.[226,1360,194,1486]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. A, B. Leofa (Prasutagus) yangae Duan and Zhang sp. n. A: dorsal habitus (♂); B: dorsal habitus (Ψ)." captionText-1="FIGURE 4. C – K, Leofa (Prasutagus) yangae Duan and Zhang sp. n. C: female genitalia; D: pygofer, dorsal view; E: pygofer, lateral view; F: apex of pygofer appendage, lateral view; G: valve and subgenital plate, ventral view; H: style, dorsal view; I: aedeagus, paraphysis and connective, lateral view; J: aedeagus, posterior view; K: connective, dorsal view." httpUri-0="https://zenodo.org/record/185059/files/figure.png" httpUri-1="https://zenodo.org/record/185060/files/figure.png" pageId="1" pageNumber="36">4E</figureCitation>
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).
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<bibRefCitation id="7E2F4B4D7853FFF795ABF9EBFD83D8F1" author="Viraktamath" box="[211,632,992,1018]" pageId="1" pageNumber="43" refString="Viraktamath, C. A. & Viraktamath, S. (1992) Revision of the deltocephaline leafhoppers of the grass feeding genus Leofa Distant (Insecta, Homoptera, Auchenorrhyncha: Cicadellidae). Entomologische Abhandlungen, 55, 1 - 12." type="journal article" year="1992">Viraktamath & Viraktamath (1992)</bibRefCitation>
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treated the subgenus as a distinct genus based on its pygofer processes etc, but here we follow
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<bibRefCitation id="7E2F4B4D7853FFF79484FE03FD4CDF29" author="Zahniser" box="[508,695,1032,1058]" pageId="1" pageNumber="43" refString="Zahniser, J. N. (2008) Seven new species and new distributions of Old World Chiasmini (Hemiptera: Cicadellidae: Deltocephalinae), with a redescription, key to genera, and species. Zootaxa, 1808, 1 - 32." type="journal article" year="2008">Zahniser (2008)</bibRefCitation>
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who placed it as a subgenus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |