688 lines
87 KiB
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688 lines
87 KiB
XML
<document id="3B37D3FE4A76ECA894195908991CE3A8" ID-DOI="10.11646/zootaxa.2757.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5291111" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1629712410579" checkinUser="felipe" docAuthor="Malhotra, Anita, Thorpe, Roger S. & Stuart, Bryan L." docDate="2011" docId="277C87B0FFD2FF96FF3C4CB0FA83FD03" docLanguage="en" docName="zt02757p023.pdf" docOrigin="Zootaxa 2757 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2757.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cryptelytrops cardamomensis Malhotra & Thorpe & Stuart 2011, sp. nov." docType="treatment" docVersion="1" lastPageNumber="9" masterDocId="DB45FFC8FFD0FF9EFFAB4A7DFF9DFF80" masterDocTitle="Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia" masterLastPageNumber="23" masterPageNumber="1" pageNumber="3" updateTime="1730369689085" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="58FFFCBE7838412A94BA44D969F12F63">Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia</mods:title>
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<mods:name id="678B7B061BD042AEC94547CD0E9BC429" type="personal">
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<mods:roleTerm id="374385F6FB98050476BD2673D919B2CF">Author</mods:roleTerm>
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<mods:namePart id="21ADF9988FF72DCAC36C0FDB0AD20DA5">Malhotra, Anita</mods:namePart>
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<mods:affiliation id="3F2909CB13EA8C83CE18200904F0F4C7">School of Biological Sciences, College of Natural Sciences, Bangor University, Gwynedd, UK.</mods:affiliation>
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<mods:namePart id="008A5BF9891A42F9038DECAB10BDFBB1">Thorpe, Roger S.</mods:namePart>
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<mods:affiliation id="6EC0DA4A0D3C699AF6E1EDBD8B9D485B">School of Biological Sciences, College of Natural Sciences, Bangor University, Gwynedd, UK.</mods:affiliation>
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<mods:namePart id="60B3001124750CEB5BD1707D69B54A9D">Stuart, Bryan L.</mods:namePart>
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<mods:affiliation id="7A279E90A296C12C0855E587E31FA8EA">North Carolina Museum of Natural Sciences, USA. E-mail: bryan. stuart @ ncdenr. gov</mods:affiliation>
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<mods:title id="E5D31926C286C01CD67521AD8D3421C9">Zootaxa</mods:title>
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<mods:date id="0BF8E2C4341C04F5943E8CB74CEB6BB3">2011</mods:date>
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<mods:number id="34933253504DA6F84A18E40E530D9DB5">2757</mods:number>
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<subSubSection id="E7CF652DFFD2FF9CFF3C4CB0FDD0F965" box="[151,589,1741,1766]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="AF6A36A6FFD2FF9CFF3C4CB0FDD0F965" blockId="2.[151,1036,1741,1801]" box="[151,589,1741,1766]" pageId="2" pageNumber="3">
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<emphasis id="9DA1EAB4FFD2FF9CFF3C4CB0FDD0F965" bold="true" box="[151,589,1741,1766]" pageId="2" pageNumber="3">
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<taxonomicName id="68D54D25FFD2FF9CFF3C4CB0FE71F966" ID-CoL="6BLFK" authority="Malhotra & Thorpe & Stuart, 2011" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[151,492,1741,1766]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">Cryptelytrops cardamomensis</taxonomicName>
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<taxonomicNameLabel id="869257CFFFD2FF9CFE584CB0FDD0F965" box="[499,589,1741,1765]" pageId="2" pageNumber="3" rank="species">
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<emphasis id="9DA1EAB4FFD2FF9CFE584CB0FD89F965" box="[499,532,1741,1765]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">sp.</emphasis>
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nov.
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</paragraph>
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<subSubSection id="E7CF652DFFD2FF9CFDF84CB0FB91F965" box="[595,1036,1741,1765]" pageId="2" pageNumber="3" type="vernacular_names">
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<paragraph id="AF6A36A6FFD2FF9CFDF84CB0FB91F965" blockId="2.[151,1036,1741,1801]" box="[595,1036,1741,1765]" pageId="2" pageNumber="3">(Cardamom Mountains Green Pitviper)</paragraph>
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</subSubSection>
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<subSubSection id="E7CF652DFFD2FF9CFF3C4C8CFE13F889" box="[151,398,1777,1801]" pageId="2" pageNumber="3" type="description">
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<paragraph id="AF6A36A6FFD2FF9CFF3C4C8CFE13F889" blockId="2.[151,1036,1741,1801]" box="[151,398,1777,1801]" pageId="2" pageNumber="3">
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<figureCitation id="37EE2A23FFD2FF9CFF3C4C8CFF79F889" box="[151,228,1777,1801]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1966]" captionTargetBox="[287,1311,209,1911]" captionTargetId="figure-66@5.[268,1319,193,1921]" captionTargetPageId="5" captionText="FIGURE 3. Line drawings of the head scalation of the holotype of Cryptelytrops cardamomensis sp. nov. (FMNH 259191) A. lateral, B. dorsal, C. ventral. Small letters identify the following measurements: a (eye width): 3.3mm, b (supraocular width): 1.55mm, c (bottom edge of rostral): 2.5mm, d: (head length): 18.75 mm." figureDoi="http://doi.org/10.5281/zenodo.5291121" httpUri="https://zenodo.org/record/5291121/files/figure.png" pageId="2" pageNumber="3">Figs. 3</figureCitation>
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,
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<figureCitation id="37EE2A23FFD2FF9CFF5A4C8CFE9FF889" box="[241,258,1777,1801]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1938]" captionTargetBox="[250,1338,202,1900]" captionTargetId="figure-82@6.[242,1344,193,1903]" captionTargetPageId="6" captionText="FIGURE 4. Cryptelytrops cardamomensis sp.nov. A. Close-up of head and mid-body region of adult male (left) and female (right) from Khao Kitchakut National Park, Chantaburi Prov., Thailand (photos: A Malhotra). B. Adult male (left) and female (right) from Kâmpóng Saôm Bay, Koh Kong Prov., Cambodia (photos: A Gumprecht); C. and D. Adult specimens from Mount Da Lai in Cardamom Mountains, southwest Cambodia (photos: Jeremy Holden)." figureDoi="http://doi.org/10.5281/zenodo.5291123" httpUri="https://zenodo.org/record/5291123/files/figure.png" pageId="2" pageNumber="3">4</figureCitation>
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;
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<tableCitation id="E257031DFFD2FF9CFEA54C8CFEF6F889" box="[270,363,1777,1801]" captionStart="TABLE 1" captionStartId="2.[151,234,1482,1505]" captionTargetPageId="2" captionText="TABLE 1. Sample sizes for each sex of the three species in the multivariate analyses." pageId="2" pageNumber="3">Tables 1</tableCitation>
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,
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<tableCitation id="E257031DFFD2FF9CFED14C8CFE16F889" box="[378,395,1777,1801]" captionStart="TABLE 2" captionStartId="9.[151,234,358,381]" captionTargetPageId="9" captionText="TABLE 2. Mean values and standard deviations of morphological characters that in combination can be used to distinguish C. cardamomensis sp. nov. from both C. macrops s.s. and C. rubeus sp. nov. Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm = C. macrops s.s., Cc = C. cardamomensis, Cr = C. rubeus. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from C. cardamomensis (e.g. BTWSUPOC can be used to discriminate both C. macrops and C. rubeus from C. cardamomensis while DV19to17 is only useful to discriminate C. rubeus from C. cardamomensis)." pageId="2" pageNumber="3">2</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="E7CF652DFFD2FF9CFF3C4D47FE7EF82B" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="AF6A36A6FFD2FF9CFF3C4D47FAB1F8EE" blockId="2.[151,1436,1849,1963]" pageId="2" pageNumber="3">
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<taxonomicName id="68D54D25FFD2FF9CFF3C4D47FD91F8CF" authority="Kramer 1977" authorityName="Kramer" authorityYear="1977" box="[151,524,1849,1871]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="macrops">
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<emphasis id="9DA1EAB4FFD2FF9CFF3C4D47FEE7F8CF" box="[151,378,1850,1871]" italics="true" pageId="2" pageNumber="3">Trimeresurus macrops</emphasis>
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<bibRefCitation id="CB444B57FFD2FF9CFE294D47FD91F8CF" author="Kramer, E." box="[386,524,1849,1871]" pageId="2" pageNumber="3" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer 1977</bibRefCitation>
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</taxonomicName>
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||
:
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||
<bibRefCitation id="CB444B57FFD2FF9CFDB34D47FD2EF8CF" author="Kramer, E." box="[536,691,1849,1871]" pageId="2" pageNumber="3" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer (1977)</bibRefCitation>
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||
: 757 (part);
|
||
<bibRefCitation id="CB444B57FFD2FF9CFC994D47FBC2F8CF" author="Regenass, U. & Kramer, E." box="[818,1119,1849,1871]" pageId="2" pageNumber="3" pagination="163 - 205" refId="ref12317" refString="Regenass, U. & Kramer, E. (1981) Zur Systematik der grunen Grubenottern der Gattung Trimeresurus Serpentes, Crotalidae. Revue Suisse Zoologie, 88, 163 - 205." type="journal article" year="1981">Regenass and Kramer (1981)</bibRefCitation>
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||
:184 (part);
|
||
<bibRefCitation id="CB444B57FFD2FF9CFB734D44FA04F8CF" author="Gumprecht, A." box="[1240,1433,1849,1871]" pageId="2" pageNumber="3" pagination="25 - 36" refId="ref11920" refString="Gumprecht, A. (1998) Die Bambusottern der Gattung Trimeresurus Lacepede. Teil II: Die Grossaugen-bambusotter Trimeresurus macrops Kramer, 1977. Sauria, 20, 25 - 36." type="journal article" year="1998">Gumprecht (1998)</bibRefCitation>
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||
: 25 (part);
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||
<bibRefCitation id="CB444B57FFD2FF9CFE874D25FE64F8EE" author="Orlov, N. & Ananjeva, N. & Barabanov, A. & Ryabov, S. & Khalikov, R." box="[300,505,1880,1902]" pageId="2" pageNumber="3" pagination="177 - 218" refId="ref12165" refString="Orlov, N., Ananjeva, N., Barabanov, A., Ryabov, S. & Khalikov, R. (2002 a) Diversity of vipers Azemiopinae, Crotalinae in East, Southeast, and South Asia: Annotated checklist and natural history data. Reptilia: Squamata: Serpentes: Viperidae. Faunistische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 23, 177 - 218." type="journal article" year="2002">
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||
Orlov
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||
<emphasis id="9DA1EAB4FFD2FF9CFEC64D24FE04F8EE" box="[365,409,1880,1902]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
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||
. (2002a)
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||
</bibRefCitation>
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||
: 193 (part);
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||
<bibRefCitation id="CB444B57FFD2FF9CFDDF4D25FCDFF8EE" author="Orlov, N. & Ananjeva, N. & Khalikov, R." box="[628,834,1880,1902]" pageId="2" pageNumber="3" pagination="345 - 360" refId="ref12238" refString="Orlov, N., Ananjeva, N. & Khalikov, R. (2002 b) Natural History of pitvipers in Eastern and Southeastern Asia. In: Schuett, G. W., Hoggren, M., Douglas, M. E. & Greene, H. W. (Eds.), Biology of the vipers. Eagle Mountain Publishing, Eagle Mountain, Utah, pp 345 - 360." type="book chapter" year="2002">
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||
Orlov
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||
<emphasis id="9DA1EAB4FFD2FF9CFD1D4D24FD7CF8EE" box="[694,737,1880,1902]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
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||
. (2002b)
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||
</bibRefCitation>
|
||
: 353 (part);
|
||
<bibRefCitation id="CB444B57FFD2FF9CFC154D25FB2BF8EE" author="Gumprecht, A. & Tillack, F. & Orlov, N. L. & Captain, A. & Ryabov, S." box="[958,1206,1880,1902]" pageId="2" pageNumber="3" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. & Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
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||
Gumprecht
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<emphasis id="9DA1EAB4FFD2FF9CFB9D4D24FBFEF8EE" box="[1078,1123,1880,1902]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
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||
. (2004)
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||
</bibRefCitation>
|
||
: 230 (part).
|
||
</paragraph>
|
||
<paragraph id="AF6A36A6FFD2FF9CFF3C4D0AFE7EF82B" blockId="2.[151,1436,1849,1963]" pageId="2" pageNumber="3">
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<taxonomicName id="68D54D25FFD2FF9CFF3C4D0AFDBFF80C" authority="(Kramer 1977)" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[151,546,1910,1933]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="macrops">
|
||
<emphasis id="9DA1EAB4FFD2FF9CFF3C4D0AFEE3F80C" box="[151,382,1911,1933]" italics="true" pageId="2" pageNumber="3">Cryptelytrops macrops</emphasis>
|
||
(
|
||
<bibRefCitation id="CB444B57FFD2FF9CFE264D0BFD87F80C" author="Kramer, E." box="[397,538,1910,1932]" pageId="2" pageNumber="3" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer 1977</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<bibRefCitation id="CB444B57FFD2FF9CFD864D0BFCA7F80C" author="Stuart, B. L. & Emmett, D. A." box="[557,826,1910,1932]" pageId="2" pageNumber="3" pagination="1 - 27" refId="ref12477" refString="Stuart, B. L. & Emmett, D. A. (2006) A collection of amphibians and reptiles from the Cardamom Mountains, southwestern Cambodia. Fieldiana, New Series, 109, 1 - 27." type="journal article" year="2006">Stuart and Emmett (2006)</bibRefCitation>
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||
: 23;
|
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<bibRefCitation id="CB444B57FFD2FF9CFCC64D0BFBD4F80C" author="Grismer, L. & Chav, T. & Neang, T. & Wood Jr, P. L. & Grismer, J. L. & Youmans, T. M. & Daltry, J. C. & Kaiser, H." box="[877,1097,1910,1933]" pageId="2" pageNumber="3" pagination="216 - 241" refId="ref11548" refString="Grismer, L. L, Chav, T., Neang, T., Wood Jr, P. L., Grismer, J. L., Youmans, T. M., Ponce 1, A., Daltry, J. C. & Kaiser, H. (2007) The herpetofauna of the Phnom Aural Wildlife Sanctuary and checklist of the herpetofauna of the Cardamom mountains, Cambodia. Hamadryad, 31, 216 - 241." type="journal article" year="2007">
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||
Grismer
|
||
<emphasis id="9DA1EAB4FFD2FF9CFC6C4D0AFC68F80D" box="[967,1013,1911,1933]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
|
||
. (2007)
|
||
</bibRefCitation>
|
||
: 232;
|
||
<bibRefCitation id="CB444B57FFD2FF9CFB224D0BFAECF80C" author="Grismer, L. & Neang, T. & Chav, T. & Grismer, J. L." box="[1161,1393,1910,1933]" pageId="2" pageNumber="3" pagination="12 - 28" refId="ref11634" refString="Grismer, L. L, Neang, T., Chav, T. & Grismer, J. L. (2008 a) Checklist of the amphibians and reptiles of the Cardamom region of southwestern Cambodia. Cambodian Journal of Natural History, 2008, 12 - 28." type="journal article" year="2008">
|
||
Grismer
|
||
<emphasis id="9DA1EAB4FFD2FF9CFB484D0AFA8CF80D" box="[1251,1297,1911,1933]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
|
||
. (2008a)
|
||
</bibRefCitation>
|
||
: 24;
|
||
<bibRefCitation id="CB444B57FFD2FF9CFF6C4DE8FE31F82B" author="Grismer, L. L. & Neang, T. & Chav, T. & Wood Jr, P. L. & Oaks, J. R. & Holden, J. & Grismer, J. L. & Szutz, T. R. & Youmans, T. M." box="[199,428,1941,1963]" pageId="2" pageNumber="3" pagination="161 - 175" refId="ref11686" refString="Grismer, L. L., Neang, T., Chav, T., Wood Jr, P. L., Oaks, J. R., Holden, J., Grismer, J. L., Szutz, T. R. & Youmans, T. M. (2008 b) Additional amphibians and reptiles from the Phnom Samkos wildlife sanctuary northwestern Cardamom Mountains, Cambodia, with comments on their taxonomy and the discovery of three new species. Raffles Bulletin of Zoology, 56, 161 - 175." type="journal article" year="2008">
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||
Grismer
|
||
<emphasis id="9DA1EAB4FFD2FF9CFE8B4DEBFED1F82B" box="[288,332,1941,1963]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
|
||
. (2008b)
|
||
</bibRefCitation>
|
||
: 166.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="FBAA662EFFD3FF9DFF3C4DEBFC41F869" ID-DOI="http://doi.org/10.5281/zenodo.5291117" ID-Zenodo-Dep="5291117" httpUri="https://zenodo.org/record/5291117/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,250,1942,1965]" targetBox="[267,1326,203,1889]" targetPageId="3">
|
||
<paragraph id="AF6A36A6FFD3FF9DFF3C4DEBFC41F869" blockId="3.[151,1436,1942,2025]" pageId="3" pageNumber="4">
|
||
<emphasis id="9DA1EAB4FFD3FF9DFF3C4DEBFE8FF82D" bold="true" box="[151,274,1942,1965]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
|
||
Canonical variate plot showing the success of multivariate morphometrics in discriminating the new species (symbols are as given in Fig. 1). Colour pattern and internal characters provide additional resolution, but could not be included in this analysis as they could not be recorded in all specimens. A. Males, B. Females.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E7CF652DFFD4FF9AFF3C4AE5FDCCFD6F" pageId="4" pageNumber="5" type="materials_examined">
|
||
<paragraph id="AF6A36A6FFD4FF9AFF3C4AE5FAECFF77" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
|
||
<materialsCitation id="1FBD3CFBFFD4FF9AFF3C4AE5FAECFF77" collectingDate="2000-08-25" collectorName="S. Platt." country="Cambodia" county="Sre Ambel District" elevation="1" latitude="11.44" location="Phnom Chan Mountain" longLatPrecision="777" longitude="103.79" municipality="Cardamom Mountains" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong" typeStatus="holotype">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFF3C4AE5FE96FF2F" bold="true" box="[151,267,152,176]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">
|
||
<typeStatus id="706E8804FFD4FF9AFF3C4AE5FE99FF30" box="[151,260,152,176]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
FMNH 259191, adult female, collected on a root of a strangler fig growing from a tree trunk
|
||
<quantity id="682D9B43FFD4FF9AFA8C4AEAFACFFF2F" box="[1319,1362,151,175]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">
|
||
<elevation id="24F8D195FFD4FF9AFA8C4AEAFACFFF2F" box="[1319,1362,151,175]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">1 m</elevation>
|
||
</quantity>
|
||
above the ground in semi-evergreen forest on
|
||
<location id="AA0A607DFFD4FF9AFDF64AC1FCF5FF54" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFDF64AC1FCF5FF54" box="[605,872,188,212]" country="Cambodia" county="Sre Ambel District" latitude="11.44" longLatPrecision="777" longitude="103.79" municipality="Cardamom Mountains" name="Phnom Chan Mountain" pageId="4" pageNumber="5" stateProvince="Koh Kong">Phnom Chan Mountain</location>
|
||
,
|
||
<collectingMunicipality id="4F0EACDCFFD4FF9AFCDD4AC1FBEBFF54" box="[886,1142,188,212]" pageId="4" pageNumber="5">Cardamom Mountains</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="460B4E2AFFD4FF9AFB2F4AC1FAC3FF54" box="[1156,1374,188,212]" pageId="4" pageNumber="5">Sre Ambel District</collectingCounty>
|
||
,
|
||
<collectingRegion id="6D11F844FFD4FF9AFAC74AC1FEDFFF77" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D7C27636FFD4FF9AFEE64AA2FE5DFF77" box="[333,448,223,247]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="CAE15061FFD4FF9AFE644AA2FDB0FF77" box="[463,557,223,247]" degrees="11.44" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="11.44">11.44 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CAE15061FFD4FF9AFD934AA2FD3DFF77" box="[568,672,223,247]" degrees="103.79" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="103.79">103.79 E</geoCoordinate>
|
||
),
|
||
<quantity id="682D9B43FFD4FF9AFD1F4AA2FCAEFF77" box="[692,819,223,248]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
|
||
<elevation id="24F8D195FFD4FF9AFD1F4AA2FCAEFF77" box="[692,819,223,248]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
|
||
100
|
||
<emphasis id="9DA1EAB4FFD4FF9AFD754A9CFD71FF78" box="[734,748,225,248]" italics="true" pageId="4" pageNumber="5">–</emphasis>
|
||
200 m
|
||
</elevation>
|
||
</quantity>
|
||
elevation, on
|
||
<date id="DB6B1066FFD4FF9AFC794AA2FB18FF77" box="[978,1157,223,247]" pageId="4" pageNumber="5" value="2000-08-25">
|
||
<collectingDate id="CB2FE98EFFD4FF9AFC794AA2FB18FF77" box="[978,1157,223,247]" pageId="4" pageNumber="5" value="2000-08-25">25 August 2000</collectingDate>
|
||
</date>
|
||
by BLS and
|
||
<collectorName id="02205370FFD4FF9AFAB24AA2FAECFF77" box="[1305,1393,223,247]" pageId="4" pageNumber="5">S. Platt.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="AF6A36A6FFD4FF9AFF6E4B79FACBFEE4" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
|
||
<materialsCitation id="1FBD3CFBFFD4FF9AFF6E4B79FACEFEE4" collectingDate="2000-08-27" collectorName="S. Platt" country="Cambodia" county="Sre Ambel District" elevation="10" latitude="11.105" location="Prek Kroch River" longLatPrecision="77" longitude="103.659" municipality="Prek Sre Ambel River" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong" typeStatus="paratype">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFF6E4B79FEA4FE9C" bold="true" box="[197,313,260,284]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">
|
||
<typeStatus id="706E8804FFD4FF9AFF6E4B79FEA9FE9C" box="[197,308,260,284]" pageId="4" pageNumber="5" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
FMNH 259192, an adult male, collected at 2000 hrs swimming mid–stream in the
|
||
<location id="AA0A607DFFD4FF9AFB7C4B79FA01FE9C" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFB7C4B79FA01FE9C" box="[1239,1436,260,284]" country="Cambodia" county="Sre Ambel District" latitude="11.105" longLatPrecision="77" longitude="103.659" municipality="Prek Sre Ambel River" name="Prek Kroch River" pageId="4" pageNumber="5" stateProvince="Koh Kong">Prek Kroch River</location>
|
||
(tributary of the
|
||
<collectingMunicipality id="4F0EACDCFFD4FF9AFEF34B5AFDC6FEBF" box="[344,603,295,319]" pageId="4" pageNumber="5">Prek Sre Ambel River</collectingMunicipality>
|
||
) in melaleuca/mangrove coastal swamp forest,
|
||
<collectingCounty id="460B4E2AFFD4FF9AFB2A4B5AFAC0FEBF" box="[1153,1373,295,319]" pageId="4" pageNumber="5">Sre Ambel District</collectingCounty>
|
||
,
|
||
<collectingRegion id="6D11F844FFD4FF9AFAC04B5AFEDFFEE4" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D7C27636FFD4FF9AFEE64B31FE5DFEE4" box="[333,448,332,356]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="CAE15061FFD4FF9AFE644B31FDA6FEE3" box="[463,571,332,356]" degrees="11.105" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="55" value="11.105">11.105 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CAE15061FFD4FF9AFDED4B31FD21FEE4" box="[582,700,332,356]" degrees="103.659" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="55" value="103.659">103.659 E</geoCoordinate>
|
||
), <
|
||
<quantity id="682D9B43FFD4FF9AFD4A4B31FC84FEE3" box="[737,793,332,356]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">
|
||
<elevation id="24F8D195FFD4FF9AFD4A4B31FC84FEE3" box="[737,793,332,356]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
elevation, on
|
||
<date id="DB6B1066FFD4FF9AFC134B31FBF6FEE4" box="[952,1131,332,356]" pageId="4" pageNumber="5" value="2000-08-27">
|
||
<collectingDate id="CB2FE98EFFD4FF9AFC134B31FBF6FEE4" box="[952,1131,332,356]" pageId="4" pageNumber="5" value="2000-08-27">27 August 2000</collectingDate>
|
||
</date>
|
||
by BLS and
|
||
<collectorName id="02205370FFD4FF9AFB554B31FACEFEE4" box="[1278,1363,332,356]" pageId="4" pageNumber="5">S. Platt</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="AF6A36A6FFD4FF9AFF6E4B12FC99FDBB" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFF6E4B12FE39FE07" bold="true" box="[197,420,367,392]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Referred material.</emphasis>
|
||
<materialsCitation id="1FBD3CFBFFD4FF9AFE074B0DFB83FE4F" collectingDate="2004-03-01" collectingDateMax="2004-08-24" collectingDateMin="2004-03-01" collectorName="D. Emmett. & D. Emmett" country="Cambodia" county="Thmar Baing District" elevation="5" latitude="11.63" location="Central Cardamoms Protected Forest" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong">
|
||
FMNH 263387, female, collected on bamboo
|
||
<quantity id="682D9B43FFD4FF9AFC124B12FC67FE07" box="[953,1018,367,391]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">
|
||
<elevation id="24F8D195FFD4FF9AFC124B12FC67FE07" box="[953,1018,367,391]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">0.5 m</elevation>
|
||
</quantity>
|
||
above the ground,
|
||
<quantity id="682D9B43FFD4FF9AFB7C4B12FA9EFE07" box="[1239,1283,367,391]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">
|
||
<elevation id="24F8D195FFD4FF9AFB7C4B12FA9EFE07" box="[1239,1283,367,391]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">1 m</elevation>
|
||
</quantity>
|
||
from a small stream in hill evergreen forest,
|
||
<location id="AA0A607DFFD4FF9AFE574BE9FC3FFE2B" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFE574BE9FC3FFE2B" box="[508,930,404,428]" country="Cambodia" county="Thmar Baing District" latitude="11.63" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" name="Central Cardamoms Protected Forest" pageId="4" pageNumber="5" stateProvince="Koh Kong">Central Cardamoms Protected Forest</location>
|
||
,
|
||
<location id="AA0A607DFFD4FF9AFC044BE9FB38FE2C" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFC044BE9FB38FE2C" box="[943,1189,404,428]" country="Cambodia" county="Thmar Baing District" latitude="11.63" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" name="Thmar Baing District" pageId="4" pageNumber="5" stateProvince="Koh Kong">Thmar Baing District</location>
|
||
,
|
||
<collectingRegion id="6D11F844FFD4FF9AFB194BE9FA0AFE2C" box="[1202,1431,404,428]" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D7C27636FFD4FF9AFF3C4BCAFE97FE4F" box="[151,266,439,463]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="CAE15061FFD4FF9AFEB34BCAFEF2FE4F" box="[280,367,439,463]" degrees="11.67" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="11.67">11.67 E</geoCoordinate>
|
||
<geoCoordinate id="CAE15061FFD4FF9AFEDD4BCAFE7DFE4F" box="[374,480,439,463]" degrees="103.71" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="103.71">103.71 N</geoCoordinate>
|
||
),
|
||
<quantity id="682D9B43FFD4FF9AFE5F4BCAFDA6FE4F" box="[500,571,439,463]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="m" value="800.0">
|
||
<elevation id="24F8D195FFD4FF9AFE5F4BCAFDA6FE4F" box="[500,571,439,463]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="m" value="800.0">800 m</elevation>
|
||
</quantity>
|
||
elevation, on
|
||
<date id="DB6B1066FFD4FF9AFD7C4BCAFCEEFE4F" box="[727,883,439,463]" pageId="4" pageNumber="5" value="2004-03-01">
|
||
<collectingDate id="CB2FE98EFFD4FF9AFD7C4BCAFCEEFE4F" box="[727,883,439,463]" pageId="4" pageNumber="5" value="2004-03-01">1 March 2004</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="02205370FFD4FF9AFC374BC5FB83FE4F" box="[924,1054,440,463]" pageId="4" pageNumber="5">D. Emmett.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="1FBD3CFBFFD4FF9AFB8F4BC5FC9DFDBB" collectingDate="2004-03-01" collectingDateMax="2004-08-24" collectingDateMin="2004-03-01" collectorName="D. Emmett. & D. Emmett" country="Cambodia" county="Thmar Baing District" elevation="5" latitude="11.63" location="Central Cardamoms Protected Forest" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong">
|
||
FMNH 267732, female, collected at 2000 hrs on a tree trunk
|
||
<quantity id="682D9B43FFD4FF9AFE694BA1FD9FFE73" box="[450,514,476,500]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">
|
||
<elevation id="24F8D195FFD4FF9AFE694BA1FD9FFE73" box="[450,514,476,500]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">0.5 m</elevation>
|
||
</quantity>
|
||
above the ground,
|
||
<quantity id="682D9B43FFD4FF9AFD734BA1FC8DFE73" box="[728,784,476,500]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">
|
||
<elevation id="24F8D195FFD4FF9AFD734BA1FC8DFE73" box="[728,784,476,500]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
from a river, in seasonally flooded lowland evergreen forest,
|
||
<collectingMunicipality id="4F0EACDCFFD4FF9AFF6D4B82FDF1FD97" box="[198,620,511,535]" pageId="4" pageNumber="5">Central Cardamoms Protected Forest</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="460B4E2AFFD4FF9AFDD34B82FCF3FD97" box="[632,878,511,535]" pageId="4" pageNumber="5">Thmar Baing District</collectingCounty>
|
||
,
|
||
<collectingRegion id="6D11F844FFD4FF9AFCD04B82FBFDFD97" box="[891,1120,511,536]" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D7C27636FFD4FF9AFBC54B82FB7CFD97" box="[1134,1249,511,535]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="CAE15061FFD4FF9AFB5A4B82FACDFD97" box="[1265,1360,511,535]" degrees="11.63" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="11.63">11.63 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CAE15061FFD4FF9AFAF64B82FF36FDBC" degrees="103.5" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="5555" value="103.5">103.5 E</geoCoordinate>
|
||
),
|
||
<quantity id="682D9B43FFD4FF9AFF144859FE9BFDBB" box="[191,262,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="5" unit="m" value="180.0">
|
||
<elevation id="24F8D195FFD4FF9AFF144859FE9BFDBB" box="[191,262,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="5" unit="m" value="180.0">180 m</elevation>
|
||
</quantity>
|
||
elevation, on
|
||
<date id="DB6B1066FFD4FF9AFE0F4859FDCAFDBC" box="[420,599,548,572]" pageId="4" pageNumber="5" value="2004-08-24">
|
||
<collectingDate id="CB2FE98EFFD4FF9AFE0F4859FDCAFDBC" box="[420,599,548,572]" pageId="4" pageNumber="5" value="2004-08-24">24 August 2004</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="02205370FFD4FF9AFD2A4859FC9DFDBB" box="[641,768,548,571]" pageId="4" pageNumber="5">D. Emmett</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="AF6A36A6FFD4FF9AFF6E483AFDCCFD6F" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFF6E483AFE2AFDE0" bold="true" box="[197,439,583,608]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Additional material.</emphasis>
|
||
<materialsCitation id="1FBD3CFBFFD4FF9AFE164835FDD0FD6F" collectingDate="1998-11-11" country="Thailand" county="Khao Kitchakut National Park" elevation="75" latitude="12.8" location="Two" longLatPrecision="775" longitude="102.13" municipality="Krathing Waterfall" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Chanthaburi">
|
||
<location id="AA0A607DFFD4FF9AFE164835FE73FDDF" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFE164835FE73FDDF" box="[445,494,584,607]" country="Thailand" county="Khao Kitchakut National Park" latitude="12.8" longLatPrecision="775" longitude="102.13" municipality="Krathing Waterfall" name="Two" pageId="4" pageNumber="5" stateProvince="Chanthaburi">Two</location>
|
||
live specimens, an adult male and female, collected at 2215 hrs in low vegetation by the fourth tier of
|
||
<collectingMunicipality id="4F0EACDCFFD4FF9AFEC24811FDDEFD04" box="[361,579,620,644]" pageId="4" pageNumber="5">Krathing Waterfall</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="460B4E2AFFD4FF9AFDF94811FC2BFD04" box="[594,950,620,644]" pageId="4" pageNumber="5">Khao Kitchakut National Park</collectingCounty>
|
||
,
|
||
<collectingRegion id="6D11F844FFD4FF9AFC6D4811FB28FD04" box="[966,1205,620,644]" country="Thailand" name="Chanthaburi" pageId="4" pageNumber="5">Chantaburi Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D7C27636FFD4FF9AFB6E4811FAB5FD04" box="[1221,1320,620,644]" name="Thailand" pageId="4" pageNumber="5">Thailand</collectingCountry>
|
||
(
|
||
<geoCoordinate id="CAE15061FFD4FF9AFA974811FA01FD03" box="[1340,1436,620,644]" degrees="12.80" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="12.8">12.80 N</geoCoordinate>
|
||
<geoCoordinate id="CAE15061FFD4FF9AFF3C48F2FE9CFD27" box="[151,257,655,679]" degrees="102.13" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="102.13">102.13 E</geoCoordinate>
|
||
),
|
||
<quantity id="682D9B43FFD4FF9AFEBC48F2FECFFD27" box="[279,338,655,679]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="5" unit="m" value="75.0">
|
||
<elevation id="24F8D195FFD4FF9AFEBC48F2FECFFD27" box="[279,338,655,679]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="5" unit="m" value="75.0">75 m</elevation>
|
||
</quantity>
|
||
elevation, on
|
||
<date id="DB6B1066FFD4FF9AFE5C48F2FD49FD27" box="[503,724,655,679]" pageId="4" pageNumber="5" value="1998-11-11">
|
||
<collectingDate id="CB2FE98EFFD4FF9AFE5C48F2FD49FD27" box="[503,724,655,679]" pageId="4" pageNumber="5" value="1998-11-11">11 November 1998</collectingDate>
|
||
</date>
|
||
, by AM and RST. Data from these living specimens has been used in all analyses, but under the conditions of permission, these specimens could not be preserved as vouchers and were released at the site of capture
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7CF652DFFD4FF9AFF6E4886FD30F814" pageId="4" pageNumber="5" type="diagnosis">
|
||
<paragraph id="AF6A36A6FFD4FF9AFF6E4886FD30F814" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFF6E4886FEA2FC93" bold="true" box="[197,319,763,788]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Diagnosis.</emphasis>
|
||
<taxonomicName id="68D54D25FFD4FF9AFEE24881FD07FC94" authority="Malhotra & Thorpe & Stuart, 2011" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[329,666,764,788]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFEE24881FD07FC94" box="[329,666,764,788]" italics="true" pageId="4" pageNumber="5">Cryptelytrops cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9DA1EAB4FFD4FF9AFD0F4881FC9DFC94" bold="true" box="[676,768,764,788]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFD0F4881FC9DFC94" box="[676,768,764,788]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="68D54D25FFD4FF9AFBAE4881FB14FC94" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[1029,1161,764,788]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="venustus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFBAE4881FB14FC94" box="[1029,1161,764,788]" italics="true" pageId="4" pageNumber="5">C. venustus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="68D54D25FFD4FF9AFB314881FADBFC94" box="[1178,1350,764,788]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="honsonensis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFB314881FADBFC94" box="[1178,1350,764,788]" italics="true" pageId="4" pageNumber="5">C. honsonensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="68D54D25FFD4FF9AFA294881FEBBFCB8" baseAuthorityName="Malhotra & Thorpe" baseAuthorityYear="2004" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="kanburiensis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFA294881FEBBFCB8" italics="true" pageId="4" pageNumber="5">C. kanburiensis</emphasis>
|
||
</taxonomicName>
|
||
, by the absence of purple-red marking (banding on the body and blotches on the head). It is distinguished from all other Asian pitviper species which have the typical "green pitviper" colouration (uniform green dorsal colour and a lateral stripe present on the first few dorsal scale rows in one or both sexes), except other species of
|
||
<taxonomicName id="68D54D25FFD4FF9AFF4249F1FE1FFC24" authorityName="Cope" authorityYear="1860" box="[233,386,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFF4249F1FE1FFC24" box="[233,386,908,932]" italics="true" pageId="4" pageNumber="5">Cryptelytrops</emphasis>
|
||
</taxonomicName>
|
||
(including
|
||
<emphasis id="9DA1EAB4FFD4FF9AFDAF49F1FCB7FC24" box="[516,810,908,932]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFDAF49F1FD07FC24" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[516,666,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
|
||
,
|
||
<taxonomicName id="68D54D25FFD4FF9AFD0C49F1FCB7FC24" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[679,810,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="68D54D25FFD4FF9AFCC949F1FBB5FC24" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[866,1064,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFCC949F1FBB5FC24" box="[866,1064,908,932]" italics="true" pageId="4" pageNumber="5">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
), by the presence of a fused first supralabial and nasal scale. It can be distinguished from
|
||
<emphasis id="9DA1EAB4FFD4FF9AFCA649CDFBADFC48" box="[781,1072,944,968]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFCA649CDFC3EFC48" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[781,931,944,968]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
|
||
,
|
||
<taxonomicName id="68D54D25FFD4FF9AFC0549CDFBADFC48" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[942,1072,944,968]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="68D54D25FFD4FF9AFBCD49CDFAB0FC48" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[1126,1325,944,968]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="septentrionalis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFBCD49CDFAB0FC48" box="[1126,1325,944,968]" italics="true" pageId="4" pageNumber="5">C. septentrionalis</emphasis>
|
||
</taxonomicName>
|
||
primarily by the relatively larger size of the eye (most obvious in adults), the relatively wider supraoculars, and the shape of the head. The latter is more elongate
|
||
<emphasis id="9DA1EAB4FFD4FF9AFD904984FDD4FB90" box="[571,585,1017,1040]" italics="true" pageId="4" pageNumber="5">–</emphasis>
|
||
oval in
|
||
<emphasis id="9DA1EAB4FFD4FF9AFD0A4985FC56FB90" box="[673,971,1016,1040]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFD0A4985FCA7FB90" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[673,826,1016,1040]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
|
||
,
|
||
<taxonomicName id="68D54D25FFD4FF9AFCEC4985FC56FB90" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[839,971,1016,1040]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="9DA1EAB4FFD4FF9AFBAD4985FB4BFB90" box="[1030,1238,1016,1040]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFBAD4985FB4CFB90" authorityName="Malhotra & Thorpe" authorityYear="2004" box="[1030,1233,1016,1040]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="septentrionalis">C. septentrionalis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but widens quite abruptly behind the eyes in
|
||
<emphasis id="9DA1EAB4FFD4FF9AFE7B4E61FCC7FBB4" box="[464,858,1052,1076]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFE7B4E61FD3FFBB4" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[464,674,1052,1076]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">C. cardamomensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="68D54D25FFD4FF9AFD054E61FCB2FBB4" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[686,815,1052,1076]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFC9C4E60FCC7FBB4" box="[823,858,1053,1076]" pageId="4" pageNumber="5" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="68D54D25FFD4FF9AFC394E61FC62FBB4" authority="Malhotra & Thorpe & Stuart, 2011" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[914,1023,1052,1076]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFC394E61FC62FBB4" box="[914,1023,1052,1076]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9DA1EAB4FFD4FF9AFBAC4E61FBFCFBB4" bold="true" box="[1031,1121,1052,1076]" pageId="4" pageNumber="5">
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFBAC4E61FBFCFBB4" box="[1031,1121,1052,1076]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(described below) to give a characteristically triangular shaped head.
|
||
<taxonomicName id="68D54D25FFD4FF9AFDCC4E3DFC2BFBD8" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[615,950,1088,1112]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFDCC4E3DFC2BFBD8" box="[615,950,1088,1112]" italics="true" pageId="4" pageNumber="5">Cryptelytrops cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from both
|
||
<emphasis id="9DA1EAB4FFD4FF9AFAB74E3DFF27FBFC" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFAB74E3DFA01FBD8" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[1308,1436,1088,1112]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFF3C4E18FF27FBFC" box="[151,186,1125,1148]" pageId="4" pageNumber="5" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="68D54D25FFD4FF9AFF5B4E19FEC1FBFC" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[240,348,1124,1148]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFF5B4E19FEC1FBFC" box="[240,348,1124,1148]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
using a combination of the following characters (for further details see
|
||
<tableCitation id="E257031DFFD4FF9AFBD24E19FB50FBFC" box="[1145,1229,1124,1148]" captionStart="TABLE 2" captionStartId="9.[151,234,358,381]" captionTargetPageId="9" captionText="TABLE 2. Mean values and standard deviations of morphological characters that in combination can be used to distinguish C. cardamomensis sp. nov. from both C. macrops s.s. and C. rubeus sp. nov. Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm = C. macrops s.s., Cc = C. cardamomensis, Cr = C. rubeus. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from C. cardamomensis (e.g. BTWSUPOC can be used to discriminate both C. macrops and C. rubeus from C. cardamomensis while DV19to17 is only useful to discriminate C. rubeus from C. cardamomensis)." pageId="4" pageNumber="5">Table 2</tableCitation>
|
||
). In males,
|
||
<taxonomicName id="68D54D25FFD4FF9AFAE54E19FEBCFB20" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFAE54E19FEBCFB20" italics="true" pageId="4" pageNumber="5">C. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
tends to have more scales between the rear edges of the supraoculars (BTWSUPOC2), a relatively larger eye (DEYE), and a more prominent postocular white stripe (SCROC) than either
|
||
<emphasis id="9DA1EAB4FFD4FF9AFB094ED1FAD0FB44" box="[1186,1357,1196,1220]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFB094ED1FABBFB44" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[1186,1318,1196,1220]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFA9A4ED0FAD0FB44" box="[1329,1357,1197,1220]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
|
||
</emphasis>
|
||
, or
|
||
<taxonomicName id="68D54D25FFD4FF9AFA294ED1FF7EFB68" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFA294ED1FF7EFB68" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, compared to
|
||
<emphasis id="9DA1EAB4FFD4FF9AFD8F4EADFD51FB68" box="[548,716,1232,1256]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFD8F4EADFD3BFB68" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[548,678,1232,1256]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFD044EACFD51FB68" box="[687,716,1233,1256]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the scale reduction from 19 to 17 scale rows around the body (VS19TO17) tends to occur further away from the head, it tends to have fewer (larger) scales bordering the supralabial scales (BORSUPOC), and less keeled temporal scales (KTEMP). Compared to
|
||
<taxonomicName id="68D54D25FFD4FF9AFBE74F65FB24FAB0" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[1100,1209,1304,1328]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFBE74F65FB24FAB0" box="[1100,1209,1304,1328]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
, the scale reduction from 19 to 17 scale rows around the body tends to involve higher (i.e., more dorsal) scale rows (DV19TO17); it tends to have more keeled body scales at mid-body (BSCK), and more scales between the last supralabial and the chin shields (VENTEDGE). In females, the lateral white stripe is much more prominent (STRIPE), always involving the first two dorsal scale rows. Furthermore, it tends to cover a larger proportion of the first scale row (SCR1) than in females of
|
||
<taxonomicName id="68D54D25FFD4FF9AFECD4FB1FE78FA64" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[358,485,1484,1508]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFECD4FB1FE78FA64" box="[358,485,1484,1508]" italics="true" pageId="4" pageNumber="5">C. macrops</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFE474FB0FD8DFA63" box="[492,528,1484,1508]" pageId="4" pageNumber="5" sensu="stricto">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFE474FB0FD94FA64" box="[492,521,1485,1508]" italics="true" pageId="4" pageNumber="5">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A small scale is present between the nasal scale and the scale bordering the anterior edge of the pit (NASPIT), whereas this is infrequently present in
|
||
<emphasis id="9DA1EAB4FFD4FF9AFC664F8DFBE2F988" box="[973,1151,1520,1544]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFC664F8DFBCCF988" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[973,1105,1520,1544]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFBF74F8CFBE2F988" box="[1116,1151,1521,1544]" pageId="4" pageNumber="5" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and never present in
|
||
<emphasis id="9DA1EAB4FFD4FF9AFA294F8DFDDFF9AC" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFA294F8DFF7AF9AC" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">C. rubeus</taxonomicName>
|
||
.
|
||
<taxonomicName id="68D54D25FFD4FF9AFF594C69FDDFF9AC" box="[242,578,1556,1580]" class="Reptilia" family="Viperidae" genus="Crytpelytrops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">Crytpelytrops cardamomensis</taxonomicName>
|
||
</emphasis>
|
||
tends to have more (smaller) scales bordering the supralabial scales (BORSUPOC) than
|
||
<taxonomicName id="68D54D25FFD4FF9AFE974C45FE37F9D0" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[316,426,1592,1616]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFE974C45FE37F9D0" box="[316,426,1592,1616]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
, but fewer (larger) than in
|
||
<taxonomicName id="68D54D25FFD4FF9AFD4A4C45FCFEF9D0" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[737,867,1592,1616]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFD4A4C45FCFEF9D0" box="[737,867,1592,1616]" italics="true" pageId="4" pageNumber="5">C. macrops</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFCC04C44FC12F9CF" box="[875,911,1592,1616]" pageId="4" pageNumber="5" sensu="stricto">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFCC04C44FC15F9D0" box="[875,904,1593,1616]" italics="true" pageId="4" pageNumber="5">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
The scale reduction from 19 to 17 scale rows around the body (DV19TO17) tends to involve lower scale rows in
|
||
<taxonomicName id="68D54D25FFD4FF9AFC3E4C21FBF8F9F4" box="[917,1125,1628,1652]" class="Reptilia" family="Viperidae" genus="Crytpelytrops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFC3E4C21FBF8F9F4" box="[917,1125,1628,1652]" italics="true" pageId="4" pageNumber="5">C. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
and the fusion between the first labial and nasal scale tends to be more complete (LABNAS) and involve partial sutures on both sides of the nostril (compare
|
||
<figureCitation id="37EE2A23FFD4FF9AFEF44CD9FE2CF93C" box="[351,433,1700,1724]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1966]" captionTargetBox="[287,1311,209,1911]" captionTargetId="figure-66@5.[268,1319,193,1921]" captionTargetPageId="5" captionText="FIGURE 3. Line drawings of the head scalation of the holotype of Cryptelytrops cardamomensis sp. nov. (FMNH 259191) A. lateral, B. dorsal, C. ventral. Small letters identify the following measurements: a (eye width): 3.3mm, b (supraocular width): 1.55mm, c (bottom edge of rostral): 2.5mm, d: (head length): 18.75 mm." figureDoi="http://doi.org/10.5281/zenodo.5291121" httpUri="https://zenodo.org/record/5291121/files/figure.png" pageId="4" pageNumber="5">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="37EE2A23FFD4FF9AFE684CD9FE4CF93C" box="[451,465,1700,1724]" captionStart="FIGURE 5" captionStartId="10.[151,250,1972,1995]" captionTargetBox="[253,1314,199,1920]" captionTargetId="figure-49@10.[246,1329,193,1928]" captionTargetPageId="10" captionText="FIGURE 5. Line drawings of the head scalation of the holotype of Cryptelytrops rubeus sp. nov. (FMNH 262718). Labelling as in Fig. 3 (a: 4.00mm, b: 2.05mm, c: 3.45mm, d: 24.9mm)." figureDoi="http://doi.org/10.5281/zenodo.5291125" httpUri="https://zenodo.org/record/5291125/files/figure.png" pageId="4" pageNumber="5">5</figureCitation>
|
||
and
|
||
<figureCitation id="37EE2A23FFD4FF9AFDBB4CD9FDBDF93C" box="[528,544,1700,1724]" captionStart="FIGURE 7" captionStartId="16.[151,250,1976,1999]" captionTargetBox="[307,1289,201,1941]" captionTargetId="figure-49@16.[283,1298,193,1942]" captionTargetPageId="16" captionText="FIGURE 7. Line drawings of the head scalation of the holotype of Cryptelytrops macrops s. s. (MHNG 1400.85). Labelling as in Fig. 3 (a: 4.15mm, b: 2.30mm, c: 3.00mm, d: 23.2mm)." figureDoi="http://doi.org/10.5281/zenodo.5291131" httpUri="https://zenodo.org/record/5291131/files/figure.png" pageId="4" pageNumber="5">7</figureCitation>
|
||
), than in either
|
||
<emphasis id="9DA1EAB4FFD4FF9AFD494CD9FC12F93C" box="[738,911,1700,1724]" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFD494CD9FCFBF93C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[738,870,1700,1724]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFCD94CD8FC12F93C" box="[882,911,1701,1724]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
|
||
</emphasis>
|
||
, or
|
||
<taxonomicName id="68D54D25FFD4FF9AFC6F4CD9FBABF93C" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[964,1078,1700,1724]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFC6F4CD9FBABF93C" box="[964,1078,1700,1724]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, compared to
|
||
<emphasis id="9DA1EAB4FFD4FF9AFA294CD9FE81F960" italics="true" pageId="4" pageNumber="5">
|
||
<taxonomicName id="68D54D25FFD4FF9AFA294CD9FF6BF960" baseAuthorityName="Kramer" baseAuthorityYear="1977" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD4FF9AFF544CB4FE81F960" box="[255,284,1737,1760]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
|
||
</emphasis>
|
||
,, the scale reduction from 19 to 17 scale rows around the body (VS19TO17) tends to occur further away from the head, it tends to have more scales between the rear edges of the supraoculars (BTWSUPOC2), a higher number of sublabial scales (SUBLAB), and a relatively smaller eye (DEYE). Compared to
|
||
<taxonomicName id="68D54D25FFD4FF9AFB4E4D6DFACFF8A8" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[1253,1362,1808,1832]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFB4E4D6DFACFF8A8" box="[1253,1362,1808,1832]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
, it has more keeled body scales at mid-body (BSCK), the scale reduction from 12 to 10 scale rows around the tail (SC12O10) also tends to occur further away from the vent, and there are a larger number of ventral scales in
|
||
<taxonomicName id="68D54D25FFD4FF9AFAE54D25FEBFF814" class="Reptilia" family="Viperidae" genus="Crytpelytrops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFAE54D25FEBFF814" italics="true" pageId="4" pageNumber="5">C. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
females than in
|
||
<taxonomicName id="68D54D25FFD4FF9AFE714D01FDDAF814" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[474,583,1916,1940]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
|
||
<emphasis id="9DA1EAB4FFD4FF9AFE714D01FDDAF814" box="[474,583,1916,1940]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
|
||
</taxonomicName>
|
||
females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="FBAA662EFFD5FF9BFF3C4DEAFCEBF86B" ID-DOI="http://doi.org/10.5281/zenodo.5291121" ID-Zenodo-Dep="5291121" httpUri="https://zenodo.org/record/5291121/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,250,1943,1966]" targetBox="[287,1311,209,1911]" targetPageId="5">
|
||
<paragraph id="AF6A36A6FFD5FF9BFF3C4DEAFCEBF86B" blockId="5.[151,1436,1943,2027]" pageId="5" pageNumber="6">
|
||
<emphasis id="9DA1EAB4FFD5FF9BFF3C4DEAFE8FF82E" bold="true" box="[151,274,1943,1966]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
|
||
Line drawings of the head scalation of the holotype of
|
||
<taxonomicName id="68D54D25FFD5FF9BFCEF4DE5FBEFF82E" authority="Malhotra & Thorpe & Stuart, 2011" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[836,1138,1944,1966]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
|
||
<emphasis id="9DA1EAB4FFD5FF9BFCEF4DE5FBEFF82E" box="[836,1138,1944,1966]" italics="true" pageId="5" pageNumber="6">Cryptelytrops cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9DA1EAB4FFD5FF9BFBD24DE5FB57F82D" bold="true" box="[1145,1226,1944,1965]" pageId="5" pageNumber="6">
|
||
<taxonomicNameLabel id="869257CFFFD5FF9BFBD24DE5FB57F82D" box="[1145,1226,1944,1965]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(FMNH 259191) A. lateral, B. dorsal, C. ventral. Small letters identify the following measurements: a (eye width): 3.3mm, b (supraocular width): 1.55mm, c (bottom edge of rostral): 2.5mm, d: (head length): 18.75 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FBAA662EFFD6FF98FF3C4D06FC57F86E" ID-DOI="http://doi.org/10.5281/zenodo.5291123" ID-Zenodo-Dep="5291123" httpUri="https://zenodo.org/record/5291123/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1915,1938]" targetBox="[250,1338,202,1900]" targetPageId="6">
|
||
<paragraph id="AF6A36A6FFD6FF98FF3C4D06FC57F86E" blockId="6.[151,1436,1915,2030]" pageId="6" pageNumber="7">
|
||
<emphasis id="9DA1EAB4FFD6FF98FF3C4D06FE89F812" bold="true" box="[151,276,1915,1938]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
|
||
<taxonomicName id="68D54D25FFD6FF98FEB74D01FDD0F812" authority="Malhotra & Thorpe & Stuart, 2011" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[284,589,1916,1938]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
|
||
<emphasis id="9DA1EAB4FFD6FF98FEB74D01FDD0F812" box="[284,589,1916,1938]" italics="true" pageId="6" pageNumber="7">Cryptelytrops cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD6FF98FDFE4D00FD00F812" box="[597,669,1917,1938]" pageId="6" pageNumber="7" rank="species">sp.nov.</taxonomicNameLabel>
|
||
A. Close-up of head and mid-body region of adult male (left) and female (right) from Khao Kitchakut National Park, Chantaburi Prov., Thailand (photos: A Malhotra). B. Adult male (left) and female (right) from Kâmpóng Saôm Bay, Koh Kong Prov., Cambodia (photos: A Gumprecht); C. and D. Adult specimens from Mount Da Lai in Cardamom Mountains, southwest Cambodia (photos: Jeremy Holden).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E7CF652DFFD7FF96FF6E4AEAFC7CFEBF" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
|
||
<paragraph id="AF6A36A6FFD7FF99FF6E4AEAFEFAFD4C" blockId="7.[151,1436,151,716]" pageId="7" pageNumber="8">
|
||
<emphasis id="9DA1EAB4FFD7FF99FF6E4AEAFE79FF2F" bold="true" box="[197,484,151,176]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">
|
||
Description of
|
||
<typeStatus id="706E8804FFD7FF99FEDD4AE5FE40FF30" box="[374,477,152,176]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Body cylindrical, head triangular in dorsal aspect and very distinct from neck. Canthus rostralis distinct. Head scales small (no large dorsal shields), smooth except in temporal region, region between the suprabials and temporal region, and on the rear of head, which show weak tubercular keeling. Dorsal body scales weakly keeled. 171 ventral scales, 63 pairs subcaudal scales, 21 dorsal scales at mid
|
||
<emphasis id="9DA1EAB4FFD7FF99FB5D4B78FA99FE9C" box="[1270,1284,261,284]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
body. Rostral scale roughly triangular, upper edge about half the width of lower edge. Pupil vertically elliptical. Loreal pit present. Nostril completely enclosed in nasal scale. Nasal scale partially fused with first supralabial, bearing a suture present behind the nostril, but not anterior to it. Ten and 11 supralabials on the right and left sides respectively, similarly 13/12 sublabials, three postocular scales. Shield bordering anterior edge of pit fused with second supralabial, but separated from nasal by one small scale on either side. Subocular scale touches third supralabial, but separated by one scale from fourth and fifth supralabials. At least 11 scales between supraoculars, with 14 scales between their posterior edges. Five teeth on palatine, 16 on pterygoid, and 16 on dentary. Snout
|
||
<emphasis id="9DA1EAB4FFD7FF99FBDD487CFB19FD98" box="[1142,1156,513,536]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
vent length
|
||
<quantity id="682D9B43FFD7FF99FAA74B82FAF7FD97" box="[1292,1386,511,535]" metricMagnitude="-1" metricUnit="m" metricValue="2.91" pageId="7" pageNumber="8" unit="cm" value="29.1">29.1 cm</quantity>
|
||
; tail length
|
||
<quantity id="682D9B43FFD7FF99FF4D4859FEA5FDBB" box="[230,312,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="8" unit="cm" value="5.6">5.6 cm</quantity>
|
||
. Head length measured from tip of snout to posterior edge of lower jawbone
|
||
<quantity id="682D9B43FFD7FF99FB314859FA8CFDBB" box="[1178,1297,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.875" pageId="7" pageNumber="8" unit="mm" value="18.75">18.75 mm</quantity>
|
||
; head width measured between rear outer edges of supraoculars
|
||
<quantity id="682D9B43FFD7FF99FD45483AFCD4FDDF" box="[750,841,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="7" pageNumber="8" unit="mm" value="8.3">8.3 mm</quantity>
|
||
, and at widest point of head
|
||
<quantity id="682D9B43FFD7FF99FB35483AFA9AFDDF" box="[1182,1287,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="7" pageNumber="8" unit="mm" value="12.9">12.9 mm</quantity>
|
||
. Supraocular scale (on right hand side)
|
||
<quantity id="682D9B43FFD7FF99FE144811FD88FD03" box="[447,533,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="8" unit="mm" value="3.9">3.9 mm</quantity>
|
||
long,
|
||
<quantity id="682D9B43FFD7FF99FDF44811FD59FD03" box="[607,708,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="7" pageNumber="8" unit="mm" value="1.55">1.55 mm</quantity>
|
||
wide. Distance from eye to nostril
|
||
<quantity id="682D9B43FFD7FF99FBFC4811FB23FD03" box="[1111,1214,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="7" pageNumber="8" unit="mm" value="3.85">3.85 mm</quantity>
|
||
, eye to pit
|
||
<quantity id="682D9B43FFD7FF99FA954811FA0AFD03" box="[1342,1431,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="mm" value="0.9">0.9 mm</quantity>
|
||
, posterior edge of pit to anterior edge of nostril
|
||
<quantity id="682D9B43FFD7FF99FD0D48F2FC9DFD27" box="[678,768,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="8" unit="mm" value="3.6">3.6 mm</quantity>
|
||
. Eye diameter
|
||
<quantity id="682D9B43FFD7FF99FC0C48F2FB9DFD27" box="[935,1024,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="8" unit="mm" value="3.3">3.3 mm</quantity>
|
||
. Scale reduction formula (derived as in
|
||
<bibRefCitation id="CB444B57FFD7FF99FF1848C9FEC7FD4C" author="Thorpe, R. S." box="[179,346,692,716]" pageId="7" pageNumber="8" pagination="27 - 43" refId="ref12721" refString="Thorpe, R. S. (1975) Quantitative handling of characters useful in snake systematics with particular reference to intraspecific variation in the ringed snake Natrix natrix (L.). Biological Journal of the Linnean Society, 7, 27 - 43." type="journal article" year="1975">Thorpe [1975]</bibRefCitation>
|
||
):
|
||
</paragraph>
|
||
<paragraph id="AF6A36A6FFD7FF99FF6E491AFEABFB8F" blockId="7.[151,1437,871,2012]" pageId="7" pageNumber="8">
|
||
<emphasis id="9DA1EAB4FFD7FF99FF6E491AFDFEFCFF" bold="true" box="[197,611,871,896]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">
|
||
Colour of
|
||
<typeStatus id="706E8804FFD7FF99FE954915FE38FC00" box="[318,421,872,896]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
Dark brownish grey dorsally, the ventral surface somewhat paler than the dorsal surface towards anterior of body, but less distinct posteriorly. Pale lateral stripe well developed, covering approximately one
|
||
<emphasis id="9DA1EAB4FFD7FF99FEC149CCFEE5FC48" box="[362,376,945,968]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
third of first dorsal scale row, increasing in width towards head. Upper and lower labials not distinct in colour from the rest of the head, but chin shields, scales between chin shields and sublabial scales are creamy white.
|
||
</paragraph>
|
||
<paragraph id="AF6A36A6FFD7FF99FF6E4E66FAAEF987" blockId="7.[151,1437,871,2012]" pageId="7" pageNumber="8">
|
||
<emphasis id="9DA1EAB4FFD7FF99FF6E4E66FE59FBB4" box="[197,452,1051,1076]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">
|
||
<emphasis id="9DA1EAB4FFD7FF99FF6E4E66FEFFFBB4" bold="true" box="[197,354,1051,1076]" pageId="7" pageNumber="8">Colour in life</emphasis>
|
||
(
|
||
<figureCitation id="37EE2A23FFD7FF99FED94E61FE2AFBB4" box="[370,439,1052,1076]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1938]" captionTargetBox="[250,1338,202,1900]" captionTargetId="figure-82@6.[242,1344,193,1903]" captionTargetPageId="6" captionText="FIGURE 4. Cryptelytrops cardamomensis sp.nov. A. Close-up of head and mid-body region of adult male (left) and female (right) from Khao Kitchakut National Park, Chantaburi Prov., Thailand (photos: A Malhotra). B. Adult male (left) and female (right) from Kâmpóng Saôm Bay, Koh Kong Prov., Cambodia (photos: A Gumprecht); C. and D. Adult specimens from Mount Da Lai in Cardamom Mountains, southwest Cambodia (photos: Jeremy Holden)." figureDoi="http://doi.org/10.5281/zenodo.5291123" httpUri="https://zenodo.org/record/5291123/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
|
||
).
|
||
</emphasis>
|
||
No notes or photographs were taken of the
|
||
<typeStatus id="706E8804FFD7FF99FC064E61FB92FBB4" box="[941,1039,1052,1076]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
in life. This description is based on photographs of the male and female specimens from Khao Kitchakut NP,
|
||
<collectingCountry id="D7C27636FFD7FF99FC434E42FBD6FBD7" box="[1000,1099,1087,1111]" name="Thailand" pageId="7" pageNumber="8">Thailand</collectingCountry>
|
||
as well as photographs on p. 230 Plate II and p. 232 Plate I (adult male from Khao Soi Dao wildlife sanctuary,
|
||
<collectingRegion id="6D11F844FFD7FF99FB824E19FA89FBFC" box="[1065,1300,1124,1148]" country="Thailand" name="Chanthaburi" pageId="7" pageNumber="8">Chantaburi Province</collectingRegion>
|
||
), and p. 233 Plate II, and p. 235 Pl II (adult male and female respectively, from
|
||
<collectingRegion id="6D11F844FFD7FF99FC274EF5FBF2FB20" box="[908,1135,1160,1184]" country="Cambodia" name="Koh Kong" pageId="7" pageNumber="8">Koh Kong Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="D7C27636FFD7FF99FBD04EF5FB6DFB20" box="[1147,1264,1160,1184]" name="Cambodia" pageId="7" pageNumber="8">Cambodia</collectingCountry>
|
||
) of
|
||
<bibRefCitation id="CB444B57FFD7FF99FAB54EF5FEB8FB44" author="Gumprecht, A. & Tillack, F. & Orlov, N. L. & Captain, A. & Ryabov, S." pageId="7" pageNumber="8" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. & Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
|
||
Gumprecht
|
||
<emphasis id="9DA1EAB4FFD7FF99FF3C4ED0FF54FB44" box="[151,201,1196,1220]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
|
||
. (2004)
|
||
</bibRefCitation>
|
||
. Dorsal and ventral surface grass green, very similar in shade. Lateral white stripe more prominent in male, extending onto the second dorsal scale row (confined to the first in female). In female, obvious bright blue dot on the upper rear quadrant of first scale row; present in some males, but less obvious. In male only, lateral white stripe extends onto the head as postocular stripe. Upper lip slightly paler than rest of dorsal surface of head, more obvious in female. Sublabials, and to lesser extent supralabials, also suffused with pale blue, mostly confined to edges of scales. Same pale blue also covers most scales on ventral surface of head to first part of body, more saturated and extensive on underside of head than in
|
||
<taxonomicName id="68D54D25FFD7FF99FD144FF9FCA2FA1C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[703,831,1412,1436]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
|
||
<emphasis id="9DA1EAB4FFD7FF99FD144FF9FCA2FA1C" box="[703,831,1412,1436]" italics="true" pageId="7" pageNumber="8">C. macrops</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD7FF99FCED4FF9FCF7FA1B" box="[838,874,1412,1435]" pageId="7" pageNumber="8" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
Occasional small patches of white and green pigment on scales of underside of head. Interstitial skin bright blue, black banding sometimes on anterior body, scales here have narrow edging of black pigment at base. Tail solid brick red, to vent on dorsal surface, approximately one-third (male) to half (female) distance to the vent on ventral surface. Eye golden yellow in both sexes.
|
||
</paragraph>
|
||
<paragraph id="AF6A36A6FFD7FF96FF6E4C6EFC7CFEBF" blockId="7.[151,1437,871,2012]" lastBlockId="8.[151,1436,151,644]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
<emphasis id="9DA1EAB4FFD7FF99FF6E4C6EFEA1F9AB" bold="true" box="[197,316,1555,1580]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Variation.</emphasis>
|
||
Sexual dimorphism is not very obvious in coloration. Females never have postocular stripes, but they may be only indistinct white streaks in males. The lateral white stripe may be much more prominent in males. Females have relatively shorter tails and fewer subcaudals (
|
||
<quantity id="682D9B43FFD7FF99FC844C21FC0EF9F4" box="[815,915,1628,1652]" metricMagnitude="0" metricUnit="m" metricValue="1.5367000000000002" metricValueMax="1.6002000000000003" metricValueMin="1.4731999999999998" pageId="7" pageNumber="8" unit="in" value="60.5" valueMax="63.0" valueMin="58.0">
|
||
58
|
||
<emphasis id="9DA1EAB4FFD7FF99FCE74C20FCC7F9F4" box="[844,858,1629,1652]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
63 in
|
||
</quantity>
|
||
females, versus
|
||
<quantity id="682D9B43FFD7FF99FBE64C21FB18F9F4" box="[1101,1157,1628,1652]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" pageId="7" pageNumber="8" unit="in" value="69.0">69 in</quantity>
|
||
the single male in which this count could be made). Females also are likely to reach an overall larger size (maximum recorded
|
||
<quantity id="682D9B43FFD7FF99FAAC4CFDFAFDF917" box="[1287,1376,1664,1688]" metricMagnitude="-1" metricUnit="m" metricValue="5.36" pageId="7" pageNumber="8" unit="cm" value="53.6">53.6 cm</quantity>
|
||
SVL versus
|
||
<quantity id="682D9B43FFD7FF99FF4D4CD9FEDFF93B" box="[230,322,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="4.99" pageId="7" pageNumber="8" unit="cm" value="49.9">49.9 cm</quantity>
|
||
for males), but the sample is small. The number of ventral scales is similar in both sexes, varying between 169 and 173. Keeling of body scales at mid
|
||
<emphasis id="9DA1EAB4FFD7FF99FD504CB4FC94F960" box="[763,777,1737,1760]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
body is usually strong. Temporal scales and scales in the region between the temporal scales and supralabials show weak tubercular keeling, and keeling is strong on the scales covering the rear of the head. The ratio between the lengths of the upper and lower edges of the rostral scale varies between 0.35 and 0.56. Supralabials vary between 10 and 11 and sublabials between 11 and 14. The suture between the nasal scale and first labial scale is always more extensive to the rear of the nostril, usually absent in front of the nostril, but there may be a slight kink in the anterior margin of the fused scale at the likely position. The minimum number of scales between supraoculars varies between 8
|
||
<emphasis id="9DA1EAB4FFD7FF99FC294DDCFC0DF838" box="[898,912,1953,1976]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
12 and there are 13
|
||
<emphasis id="9DA1EAB4FFD7FF99FBCE4DDCFBEEF838" box="[1125,1139,1953,1976]" italics="true" pageId="7" pageNumber="8">–</emphasis>
|
||
16 scales between the posterior edges of the supraoculars. The number of scales between the nasals and shield bordering the anterior of the pit varies between 0
|
||
<emphasis id="9DA1EAB4FFD8FF96FED14AE4FE15FF30" box="[378,392,153,176]" italics="true" pageId="8" pageNumber="9">–</emphasis>
|
||
2, and there are 0
|
||
<emphasis id="9DA1EAB4FFD8FF96FDE64AE4FDC6FF30" box="[589,603,153,176]" italics="true" pageId="8" pageNumber="9">–</emphasis>
|
||
1 scales separating the internasals. There are 0
|
||
<emphasis id="9DA1EAB4FFD8FF96FBC24AE4FBEAFF30" box="[1129,1143,153,176]" italics="true" pageId="8" pageNumber="9">–</emphasis>
|
||
1 scales between the third supralabial and the subocular, one scale between the fourth supralabial, and 1
|
||
<emphasis id="9DA1EAB4FFD8FF96FC594AC0FB9DFF54" box="[1010,1024,189,212]" italics="true" pageId="8" pageNumber="9">–</emphasis>
|
||
2 scales between the fifth supralabial and the subocular scale. Hemipenes were examined in FMNH 259192, in which they were both partly everted, and do not differ notably from those of
|
||
<taxonomicName id="68D54D25FFD8FF96FD8F4B79FD3EFE9C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[548,675,260,284]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
|
||
<emphasis id="9DA1EAB4FFD8FF96FD8F4B79FD3EFE9C" box="[548,675,260,284]" italics="true" pageId="8" pageNumber="9">C. macrops</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="869257CFFFD8FF96FD004B78FD52FE9B" box="[683,719,260,284]" pageId="8" pageNumber="9" sensu="stricto">
|
||
<emphasis id="9DA1EAB4FFD8FF96FD004B78FD5AFE9C" box="[683,711,261,284]" italics="true" pageId="8" pageNumber="9">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Teeth were also counted in this specimen, and were identical to the
|
||
<typeStatus id="706E8804FFD8FF96FF6B4B5AFEBFFEBF" box="[192,290,295,319]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
except for the number of pterygoid teeth, which numbered 14.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7CF652DFFD8FF96FF6E4B36FDBDFE4F" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="AF6A36A6FFD8FF96FF6E4B36FDBDFE4F" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="9">
|
||
<emphasis id="9DA1EAB4FFD8FF96FF6E4B36FEC3FEE4" bold="true" box="[197,350,331,356]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Distribution.</emphasis>
|
||
<taxonomicName id="68D54D25FFD8FF96FEC14B31FD20FEE4" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[362,701,332,356]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="9DA1EAB4FFD8FF96FEC14B31FD20FEE4" box="[362,701,332,356]" italics="true" pageId="8" pageNumber="9">Cryptelytrops cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
is currently known from
|
||
<collectingRegion id="6D11F844FFD8FF96FC5A4B31FB47FEE4" box="[1009,1242,332,356]" country="Cambodia" name="Koh Kong" pageId="8" pageNumber="9">Koh Kong Province</collectingRegion>
|
||
in southwestern
|
||
<collectingCountry id="D7C27636FFD8FF96FF3C4B12FE97FE07" box="[151,266,367,391]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
|
||
and
|
||
<collectingRegion id="6D11F844FFD8FF96FEE94B12FDB4FE07" box="[322,553,367,391]" country="Thailand" name="Chanthaburi" pageId="8" pageNumber="9">Chantaburi Province</collectingRegion>
|
||
in southeastern
|
||
<collectingCountry id="D7C27636FFD8FF96FD494B12FCD4FE07" box="[738,841,367,391]" name="Thailand" pageId="8" pageNumber="9">Thailand</collectingCountry>
|
||
. (
|
||
<figureCitation id="37EE2A23FFD8FF96FCF74B12FC03FE07" box="[860,926,367,391]" captionStart="FIGURE 1" captionStartId="1.[151,250,1878,1901]" captionTargetBox="[176,1411,773,1840]" captionTargetId="figure-285@1.[173,1417,770,1842]" captionTargetPageId="1" captionText="FIGURE 1. Map of Southeast Asia, showing the distribution of the two new species and Cryptelytrops macrops s.s. Symbols indicate sampling localities used in the genetic analysis to define species. Grey circles: C. macrops s.s; open diamonds: C. cardamomensis sp. nov.; black triangles: C. rubeus sp. nov. The empty circle and triangle denotes localities of specimens that have either been referred to the corresponding species on the basis of photographs, or for which the locality is uncertain and requires confirmation." figureDoi="http://doi.org/10.5281/zenodo.5291115" httpUri="https://zenodo.org/record/5291115/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
|
||
). The species probably occurs in appropriate habitat throughout the Cardamom Mountains in southwestern
|
||
<collectingCountry id="D7C27636FFD8FF96FCF14BE9FC50FE2C" box="[858,973,404,428]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
|
||
and adjacent southeastern
|
||
<collectingCountry id="D7C27636FFD8FF96FAAB4BE9FAFBFE2C" box="[1280,1382,404,428]" name="Thailand" pageId="8" pageNumber="9">Thailand</collectingCountry>
|
||
, and offshore islands (e.g., Koh Chang).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7CF652DFFD8FF96FF6E4BA1FF44FDE0" pageId="8" pageNumber="9" type="biology_ecology">
|
||
<paragraph id="AF6A36A6FFD8FF96FF6E4BA1FF44FDE0" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="9">
|
||
<emphasis id="9DA1EAB4FFD8FF96FF6E4BA1FEB7FE73" bold="true" box="[197,298,476,500]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Ecology.</emphasis>
|
||
<taxonomicName id="68D54D25FFD8FF96FE9A4BA1FD1DFE74" authorityName="Malhotra & Thorpe & Stuart" authorityYear="2011" box="[305,640,476,500]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="9DA1EAB4FFD8FF96FE9A4BA1FD1DFE74" box="[305,640,476,500]" italics="true" pageId="8" pageNumber="9">Cryptelytrops cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
is currently known from low-lying coastal areas up to
|
||
<quantity id="682D9B43FFD8FF96FB4C4BA1FAB3FE73" box="[1255,1326,476,500]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="m" value="800.0">800 m</quantity>
|
||
elevation in semi-evergreen, evergreen, and mangrove/melaleuca swamp forest. All specimens were found within a couple of metres above the ground, and most within
|
||
<quantity id="682D9B43FFD8FF96FDD64859FD25FDBB" box="[637,696,548,572]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="10.0">10 m</quantity>
|
||
of water. No food items were found in the digestive tracts of the
|
||
<typeStatus id="706E8804FFD8FF96FF3C4835FF4BFDE0" box="[151,214,584,608]" pageId="8" pageNumber="9">types</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7CF652DFFD8FF96FF6E4811FA83FD03" box="[197,1310,620,644]" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="AF6A36A6FFD8FF96FF6E4811FA83FD03" blockId="8.[151,1436,151,644]" box="[197,1310,620,644]" pageId="8" pageNumber="9">
|
||
<emphasis id="9DA1EAB4FFD8FF96FF6E4811FED1FD04" bold="true" box="[197,332,620,644]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Etymology.</emphasis>
|
||
The name refers to the mountain range wherein the
|
||
<typeStatus id="706E8804FFD8FF96FC3E4811FC58FD04" box="[917,965,620,644]" pageId="8" pageNumber="9">type</typeStatus>
|
||
locality of this species occurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |