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<mods:title id="58FFFCBE7838412A94BA44D969F12F63">Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia</mods:title>
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<mods:namePart id="21ADF9988FF72DCAC36C0FDB0AD20DA5">Malhotra, Anita</mods:namePart>
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<mods:namePart id="008A5BF9891A42F9038DECAB10BDFBB1">Thorpe, Roger S.</mods:namePart>
<mods:affiliation id="6EC0DA4A0D3C699AF6E1EDBD8B9D485B">School of Biological Sciences, College of Natural Sciences, Bangor University, Gwynedd, UK.</mods:affiliation>
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<mods:namePart id="60B3001124750CEB5BD1707D69B54A9D">Stuart, Bryan L.</mods:namePart>
<mods:affiliation id="7A279E90A296C12C0855E587E31FA8EA">North Carolina Museum of Natural Sciences, USA. E-mail: bryan. stuart @ ncdenr. gov</mods:affiliation>
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<taxonomicName id="68D54D25FFD2FF9CFF3C4CB0FE71F966" ID-CoL="6BLFK" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[151,492,1741,1766]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">Cryptelytrops cardamomensis</taxonomicName>
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<emphasis id="9DA1EAB4FFD2FF9CFE584CB0FD89F965" box="[499,532,1741,1765]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">sp.</emphasis>
nov.
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<paragraph id="AF6A36A6FFD2FF9CFDF84CB0FB91F965" blockId="2.[151,1036,1741,1801]" box="[595,1036,1741,1765]" pageId="2" pageNumber="3">(Cardamom Mountains Green Pitviper)</paragraph>
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<paragraph id="AF6A36A6FFD2FF9CFF3C4C8CFE13F889" blockId="2.[151,1036,1741,1801]" box="[151,398,1777,1801]" pageId="2" pageNumber="3">
<figureCitation id="37EE2A23FFD2FF9CFF3C4C8CFF79F889" box="[151,228,1777,1801]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1966]" captionTargetBox="[287,1311,209,1911]" captionTargetId="figure-66@5.[268,1319,193,1921]" captionTargetPageId="5" captionText="FIGURE 3. Line drawings of the head scalation of the holotype of Cryptelytrops cardamomensis sp. nov. (FMNH 259191) A. lateral, B. dorsal, C. ventral. Small letters identify the following measurements: a (eye width): 3.3mm, b (supraocular width): 1.55mm, c (bottom edge of rostral): 2.5mm, d: (head length): 18.75 mm." figureDoi="http://doi.org/10.5281/zenodo.5291121" httpUri="https://zenodo.org/record/5291121/files/figure.png" pageId="2" pageNumber="3">Figs. 3</figureCitation>
,
<figureCitation id="37EE2A23FFD2FF9CFF5A4C8CFE9FF889" box="[241,258,1777,1801]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1938]" captionTargetBox="[250,1338,202,1900]" captionTargetId="figure-82@6.[242,1344,193,1903]" captionTargetPageId="6" captionText="FIGURE 4. Cryptelytrops cardamomensis sp.nov. A. Close-up of head and mid-body region of adult male (left) and female (right) from Khao Kitchakut National Park, Chantaburi Prov., Thailand (photos: A Malhotra). B. Adult male (left) and female (right) from Kâmpóng Saôm Bay, Koh Kong Prov., Cambodia (photos: A Gumprecht); C. and D. Adult specimens from Mount Da Lai in Cardamom Mountains, southwest Cambodia (photos: Jeremy Holden)." figureDoi="http://doi.org/10.5281/zenodo.5291123" httpUri="https://zenodo.org/record/5291123/files/figure.png" pageId="2" pageNumber="3">4</figureCitation>
;
<tableCitation id="E257031DFFD2FF9CFEA54C8CFEF6F889" box="[270,363,1777,1801]" captionStart="TABLE 1" captionStartId="2.[151,234,1482,1505]" captionTargetPageId="2" captionText="TABLE 1. Sample sizes for each sex of the three species in the multivariate analyses." pageId="2" pageNumber="3">Tables 1</tableCitation>
,
<tableCitation id="E257031DFFD2FF9CFED14C8CFE16F889" box="[378,395,1777,1801]" captionStart="TABLE 2" captionStartId="9.[151,234,358,381]" captionTargetPageId="9" captionText="TABLE 2. Mean values and standard deviations of morphological characters that in combination can be used to distinguish C. cardamomensis sp. nov. from both C. macrops s.s. and C. rubeus sp. nov. Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm = C. macrops s.s., Cc = C. cardamomensis, Cr = C. rubeus. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from C. cardamomensis (e.g. BTWSUPOC can be used to discriminate both C. macrops and C. rubeus from C. cardamomensis while DV19to17 is only useful to discriminate C. rubeus from C. cardamomensis)." pageId="2" pageNumber="3">2</tableCitation>
.
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<taxonomicName id="68D54D25FFD2FF9CFF3C4D47FD91F8CF" authority="Kramer 1977" authorityName="Kramer" authorityYear="1977" box="[151,524,1849,1871]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="macrops">
<emphasis id="9DA1EAB4FFD2FF9CFF3C4D47FEE7F8CF" box="[151,378,1850,1871]" italics="true" pageId="2" pageNumber="3">Trimeresurus macrops</emphasis>
<bibRefCitation id="CB444B57FFD2FF9CFE294D47FD91F8CF" author="Kramer, E." box="[386,524,1849,1871]" pageId="2" pageNumber="3" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer 1977</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="CB444B57FFD2FF9CFDB34D47FD2EF8CF" author="Kramer, E." box="[536,691,1849,1871]" pageId="2" pageNumber="3" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer (1977)</bibRefCitation>
: 757 (part);
<bibRefCitation id="CB444B57FFD2FF9CFC994D47FBC2F8CF" author="Regenass, U. &amp; Kramer, E." box="[818,1119,1849,1871]" pageId="2" pageNumber="3" pagination="163 - 205" refId="ref12317" refString="Regenass, U. &amp; Kramer, E. (1981) Zur Systematik der grunen Grubenottern der Gattung Trimeresurus Serpentes, Crotalidae. Revue Suisse Zoologie, 88, 163 - 205." type="journal article" year="1981">Regenass and Kramer (1981)</bibRefCitation>
:184 (part);
<bibRefCitation id="CB444B57FFD2FF9CFB734D44FA04F8CF" author="Gumprecht, A." box="[1240,1433,1849,1871]" pageId="2" pageNumber="3" pagination="25 - 36" refId="ref11920" refString="Gumprecht, A. (1998) Die Bambusottern der Gattung Trimeresurus Lacepede. Teil II: Die Grossaugen-bambusotter Trimeresurus macrops Kramer, 1977. Sauria, 20, 25 - 36." type="journal article" year="1998">Gumprecht (1998)</bibRefCitation>
: 25 (part);
<bibRefCitation id="CB444B57FFD2FF9CFE874D25FE64F8EE" author="Orlov, N. &amp; Ananjeva, N. &amp; Barabanov, A. &amp; Ryabov, S. &amp; Khalikov, R." box="[300,505,1880,1902]" pageId="2" pageNumber="3" pagination="177 - 218" refId="ref12165" refString="Orlov, N., Ananjeva, N., Barabanov, A., Ryabov, S. &amp; Khalikov, R. (2002 a) Diversity of vipers Azemiopinae, Crotalinae in East, Southeast, and South Asia: Annotated checklist and natural history data. Reptilia: Squamata: Serpentes: Viperidae. Faunistische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 23, 177 - 218." type="journal article" year="2002">
Orlov
<emphasis id="9DA1EAB4FFD2FF9CFEC64D24FE04F8EE" box="[365,409,1880,1902]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. (2002a)
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: 193 (part);
<bibRefCitation id="CB444B57FFD2FF9CFDDF4D25FCDFF8EE" author="Orlov, N. &amp; Ananjeva, N. &amp; Khalikov, R." box="[628,834,1880,1902]" pageId="2" pageNumber="3" pagination="345 - 360" refId="ref12238" refString="Orlov, N., Ananjeva, N. &amp; Khalikov, R. (2002 b) Natural History of pitvipers in Eastern and Southeastern Asia. In: Schuett, G. W., Hoggren, M., Douglas, M. E. &amp; Greene, H. W. (Eds.), Biology of the vipers. Eagle Mountain Publishing, Eagle Mountain, Utah, pp 345 - 360." type="book chapter" year="2002">
Orlov
<emphasis id="9DA1EAB4FFD2FF9CFD1D4D24FD7CF8EE" box="[694,737,1880,1902]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. (2002b)
</bibRefCitation>
: 353 (part);
<bibRefCitation id="CB444B57FFD2FF9CFC154D25FB2BF8EE" author="Gumprecht, A. &amp; Tillack, F. &amp; Orlov, N. L. &amp; Captain, A. &amp; Ryabov, S." box="[958,1206,1880,1902]" pageId="2" pageNumber="3" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. &amp; Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
Gumprecht
<emphasis id="9DA1EAB4FFD2FF9CFB9D4D24FBFEF8EE" box="[1078,1123,1880,1902]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. (2004)
</bibRefCitation>
: 230 (part).
</paragraph>
<paragraph id="AF6A36A6FFD2FF9CFF3C4D0AFE7EF82B" blockId="2.[151,1436,1849,1963]" pageId="2" pageNumber="3">
<taxonomicName id="68D54D25FFD2FF9CFF3C4D0AFDBFF80C" authority="(Kramer 1977)" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[151,546,1910,1933]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="macrops">
<emphasis id="9DA1EAB4FFD2FF9CFF3C4D0AFEE3F80C" box="[151,382,1911,1933]" italics="true" pageId="2" pageNumber="3">Cryptelytrops macrops</emphasis>
(
<bibRefCitation id="CB444B57FFD2FF9CFE264D0BFD87F80C" author="Kramer, E." box="[397,538,1910,1932]" pageId="2" pageNumber="3" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer 1977</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="CB444B57FFD2FF9CFD864D0BFCA7F80C" author="Stuart, B. L. &amp; Emmett, D. A." box="[557,826,1910,1932]" pageId="2" pageNumber="3" pagination="1 - 27" refId="ref12477" refString="Stuart, B. L. &amp; Emmett, D. A. (2006) A collection of amphibians and reptiles from the Cardamom Mountains, southwestern Cambodia. Fieldiana, New Series, 109, 1 - 27." type="journal article" year="2006">Stuart and Emmett (2006)</bibRefCitation>
: 23;
<bibRefCitation id="CB444B57FFD2FF9CFCC64D0BFBD4F80C" author="Grismer, L. &amp; Chav, T. &amp; Neang, T. &amp; Wood Jr, P. L. &amp; Grismer, J. L. &amp; Youmans, T. M. &amp; Daltry, J. C. &amp; Kaiser, H." box="[877,1097,1910,1933]" pageId="2" pageNumber="3" pagination="216 - 241" refId="ref11548" refString="Grismer, L. L, Chav, T., Neang, T., Wood Jr, P. L., Grismer, J. L., Youmans, T. M., Ponce 1, A., Daltry, J. C. &amp; Kaiser, H. (2007) The herpetofauna of the Phnom Aural Wildlife Sanctuary and checklist of the herpetofauna of the Cardamom mountains, Cambodia. Hamadryad, 31, 216 - 241." type="journal article" year="2007">
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<emphasis id="9DA1EAB4FFD2FF9CFC6C4D0AFC68F80D" box="[967,1013,1911,1933]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. (2007)
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: 232;
<bibRefCitation id="CB444B57FFD2FF9CFB224D0BFAECF80C" author="Grismer, L. &amp; Neang, T. &amp; Chav, T. &amp; Grismer, J. L." box="[1161,1393,1910,1933]" pageId="2" pageNumber="3" pagination="12 - 28" refId="ref11634" refString="Grismer, L. L, Neang, T., Chav, T. &amp; Grismer, J. L. (2008 a) Checklist of the amphibians and reptiles of the Cardamom region of southwestern Cambodia. Cambodian Journal of Natural History, 2008, 12 - 28." type="journal article" year="2008">
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: 24;
<bibRefCitation id="CB444B57FFD2FF9CFF6C4DE8FE31F82B" author="Grismer, L. L. &amp; Neang, T. &amp; Chav, T. &amp; Wood Jr, P. L. &amp; Oaks, J. R. &amp; Holden, J. &amp; Grismer, J. L. &amp; Szutz, T. R. &amp; Youmans, T. M." box="[199,428,1941,1963]" pageId="2" pageNumber="3" pagination="161 - 175" refId="ref11686" refString="Grismer, L. L., Neang, T., Chav, T., Wood Jr, P. L., Oaks, J. R., Holden, J., Grismer, J. L., Szutz, T. R. &amp; Youmans, T. M. (2008 b) Additional amphibians and reptiles from the Phnom Samkos wildlife sanctuary northwestern Cardamom Mountains, Cambodia, with comments on their taxonomy and the discovery of three new species. Raffles Bulletin of Zoology, 56, 161 - 175." type="journal article" year="2008">
Grismer
<emphasis id="9DA1EAB4FFD2FF9CFE8B4DEBFED1F82B" box="[288,332,1941,1963]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. (2008b)
</bibRefCitation>
: 166.
</paragraph>
</subSubSection>
<caption id="FBAA662EFFD3FF9DFF3C4DEBFC41F869" ID-DOI="http://doi.org/10.5281/zenodo.5291117" ID-Zenodo-Dep="5291117" httpUri="https://zenodo.org/record/5291117/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,250,1942,1965]" targetBox="[267,1326,203,1889]" targetPageId="3">
<paragraph id="AF6A36A6FFD3FF9DFF3C4DEBFC41F869" blockId="3.[151,1436,1942,2025]" pageId="3" pageNumber="4">
<emphasis id="9DA1EAB4FFD3FF9DFF3C4DEBFE8FF82D" bold="true" box="[151,274,1942,1965]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
Canonical variate plot showing the success of multivariate morphometrics in discriminating the new species (symbols are as given in Fig. 1). Colour pattern and internal characters provide additional resolution, but could not be included in this analysis as they could not be recorded in all specimens. A. Males, B. Females.
</paragraph>
</caption>
<subSubSection id="E7CF652DFFD4FF9AFF3C4AE5FDCCFD6F" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="AF6A36A6FFD4FF9AFF3C4AE5FAECFF77" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<materialsCitation id="1FBD3CFBFFD4FF9AFF3C4AE5FAECFF77" collectingDate="2000-08-25" collectorName="S. Platt." country="Cambodia" county="Sre Ambel District" elevation="1" latitude="11.44" location="Phnom Chan Mountain" longLatPrecision="777" longitude="103.79" municipality="Cardamom Mountains" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong" typeStatus="holotype">
<emphasis id="9DA1EAB4FFD4FF9AFF3C4AE5FE96FF2F" bold="true" box="[151,267,152,176]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">
<typeStatus id="706E8804FFD4FF9AFF3C4AE5FE99FF30" box="[151,260,152,176]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
.
</emphasis>
FMNH 259191, adult female, collected on a root of a strangler fig growing from a tree trunk
<quantity id="682D9B43FFD4FF9AFA8C4AEAFACFFF2F" box="[1319,1362,151,175]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">
<elevation id="24F8D195FFD4FF9AFA8C4AEAFACFFF2F" box="[1319,1362,151,175]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">1 m</elevation>
</quantity>
above the ground in semi-evergreen forest on
<location id="AA0A607DFFD4FF9AFDF64AC1FCF5FF54" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFDF64AC1FCF5FF54" box="[605,872,188,212]" country="Cambodia" county="Sre Ambel District" latitude="11.44" longLatPrecision="777" longitude="103.79" municipality="Cardamom Mountains" name="Phnom Chan Mountain" pageId="4" pageNumber="5" stateProvince="Koh Kong">Phnom Chan Mountain</location>
,
<collectingMunicipality id="4F0EACDCFFD4FF9AFCDD4AC1FBEBFF54" box="[886,1142,188,212]" pageId="4" pageNumber="5">Cardamom Mountains</collectingMunicipality>
,
<collectingCounty id="460B4E2AFFD4FF9AFB2F4AC1FAC3FF54" box="[1156,1374,188,212]" pageId="4" pageNumber="5">Sre Ambel District</collectingCounty>
,
<collectingRegion id="6D11F844FFD4FF9AFAC74AC1FEDFFF77" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD4FF9AFEE64AA2FE5DFF77" box="[333,448,223,247]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
(
<geoCoordinate id="CAE15061FFD4FF9AFE644AA2FDB0FF77" box="[463,557,223,247]" degrees="11.44" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="11.44">11.44 N</geoCoordinate>
,
<geoCoordinate id="CAE15061FFD4FF9AFD934AA2FD3DFF77" box="[568,672,223,247]" degrees="103.79" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="103.79">103.79 E</geoCoordinate>
),
<quantity id="682D9B43FFD4FF9AFD1F4AA2FCAEFF77" box="[692,819,223,248]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<elevation id="24F8D195FFD4FF9AFD1F4AA2FCAEFF77" box="[692,819,223,248]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
100
<emphasis id="9DA1EAB4FFD4FF9AFD754A9CFD71FF78" box="[734,748,225,248]" italics="true" pageId="4" pageNumber="5"></emphasis>
200 m
</elevation>
</quantity>
elevation, on
<date id="DB6B1066FFD4FF9AFC794AA2FB18FF77" box="[978,1157,223,247]" pageId="4" pageNumber="5" value="2000-08-25">
<collectingDate id="CB2FE98EFFD4FF9AFC794AA2FB18FF77" box="[978,1157,223,247]" pageId="4" pageNumber="5" value="2000-08-25">25 August 2000</collectingDate>
</date>
by BLS and
<collectorName id="02205370FFD4FF9AFAB24AA2FAECFF77" box="[1305,1393,223,247]" pageId="4" pageNumber="5">S. Platt.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="AF6A36A6FFD4FF9AFF6E4B79FACBFEE4" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<materialsCitation id="1FBD3CFBFFD4FF9AFF6E4B79FACEFEE4" collectingDate="2000-08-27" collectorName="S. Platt" country="Cambodia" county="Sre Ambel District" elevation="10" latitude="11.105" location="Prek Kroch River" longLatPrecision="77" longitude="103.659" municipality="Prek Sre Ambel River" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong" typeStatus="paratype">
<emphasis id="9DA1EAB4FFD4FF9AFF6E4B79FEA4FE9C" bold="true" box="[197,313,260,284]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">
<typeStatus id="706E8804FFD4FF9AFF6E4B79FEA9FE9C" box="[197,308,260,284]" pageId="4" pageNumber="5" type="paratype">Paratype</typeStatus>
.
</emphasis>
FMNH 259192, an adult male, collected at 2000 hrs swimming midstream in the
<location id="AA0A607DFFD4FF9AFB7C4B79FA01FE9C" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFB7C4B79FA01FE9C" box="[1239,1436,260,284]" country="Cambodia" county="Sre Ambel District" latitude="11.105" longLatPrecision="77" longitude="103.659" municipality="Prek Sre Ambel River" name="Prek Kroch River" pageId="4" pageNumber="5" stateProvince="Koh Kong">Prek Kroch River</location>
(tributary of the
<collectingMunicipality id="4F0EACDCFFD4FF9AFEF34B5AFDC6FEBF" box="[344,603,295,319]" pageId="4" pageNumber="5">Prek Sre Ambel River</collectingMunicipality>
) in melaleuca/mangrove coastal swamp forest,
<collectingCounty id="460B4E2AFFD4FF9AFB2A4B5AFAC0FEBF" box="[1153,1373,295,319]" pageId="4" pageNumber="5">Sre Ambel District</collectingCounty>
,
<collectingRegion id="6D11F844FFD4FF9AFAC04B5AFEDFFEE4" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD4FF9AFEE64B31FE5DFEE4" box="[333,448,332,356]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
(
<geoCoordinate id="CAE15061FFD4FF9AFE644B31FDA6FEE3" box="[463,571,332,356]" degrees="11.105" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="55" value="11.105">11.105 N</geoCoordinate>
,
<geoCoordinate id="CAE15061FFD4FF9AFDED4B31FD21FEE4" box="[582,700,332,356]" degrees="103.659" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="55" value="103.659">103.659 E</geoCoordinate>
), &lt;
<quantity id="682D9B43FFD4FF9AFD4A4B31FC84FEE3" box="[737,793,332,356]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">
<elevation id="24F8D195FFD4FF9AFD4A4B31FC84FEE3" box="[737,793,332,356]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">10 m</elevation>
</quantity>
elevation, on
<date id="DB6B1066FFD4FF9AFC134B31FBF6FEE4" box="[952,1131,332,356]" pageId="4" pageNumber="5" value="2000-08-27">
<collectingDate id="CB2FE98EFFD4FF9AFC134B31FBF6FEE4" box="[952,1131,332,356]" pageId="4" pageNumber="5" value="2000-08-27">27 August 2000</collectingDate>
</date>
by BLS and
<collectorName id="02205370FFD4FF9AFB554B31FACEFEE4" box="[1278,1363,332,356]" pageId="4" pageNumber="5">S. Platt</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="AF6A36A6FFD4FF9AFF6E4B12FC99FDBB" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<emphasis id="9DA1EAB4FFD4FF9AFF6E4B12FE39FE07" bold="true" box="[197,420,367,392]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Referred material.</emphasis>
<materialsCitation id="1FBD3CFBFFD4FF9AFE074B0DFB83FE4F" collectingDate="2004-03-01" collectingDateMax="2004-08-24" collectingDateMin="2004-03-01" collectorName="D. Emmett. &amp; D. Emmett" country="Cambodia" county="Thmar Baing District" elevation="5" latitude="11.63" location="Central Cardamoms Protected Forest" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong">
FMNH 263387, female, collected on bamboo
<quantity id="682D9B43FFD4FF9AFC124B12FC67FE07" box="[953,1018,367,391]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">
<elevation id="24F8D195FFD4FF9AFC124B12FC67FE07" box="[953,1018,367,391]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">0.5 m</elevation>
</quantity>
above the ground,
<quantity id="682D9B43FFD4FF9AFB7C4B12FA9EFE07" box="[1239,1283,367,391]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">
<elevation id="24F8D195FFD4FF9AFB7C4B12FA9EFE07" box="[1239,1283,367,391]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1.0">1 m</elevation>
</quantity>
from a small stream in hill evergreen forest,
<location id="AA0A607DFFD4FF9AFE574BE9FC3FFE2B" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFE574BE9FC3FFE2B" box="[508,930,404,428]" country="Cambodia" county="Thmar Baing District" latitude="11.63" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" name="Central Cardamoms Protected Forest" pageId="4" pageNumber="5" stateProvince="Koh Kong">Central Cardamoms Protected Forest</location>
,
<location id="AA0A607DFFD4FF9AFC044BE9FB38FE2C" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFC044BE9FB38FE2C" box="[943,1189,404,428]" country="Cambodia" county="Thmar Baing District" latitude="11.63" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" name="Thmar Baing District" pageId="4" pageNumber="5" stateProvince="Koh Kong">Thmar Baing District</location>
,
<collectingRegion id="6D11F844FFD4FF9AFB194BE9FA0AFE2C" box="[1202,1431,404,428]" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD4FF9AFF3C4BCAFE97FE4F" box="[151,266,439,463]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
(
<geoCoordinate id="CAE15061FFD4FF9AFEB34BCAFEF2FE4F" box="[280,367,439,463]" degrees="11.67" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="11.67">11.67 E</geoCoordinate>
<geoCoordinate id="CAE15061FFD4FF9AFEDD4BCAFE7DFE4F" box="[374,480,439,463]" degrees="103.71" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="103.71">103.71 N</geoCoordinate>
),
<quantity id="682D9B43FFD4FF9AFE5F4BCAFDA6FE4F" box="[500,571,439,463]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="m" value="800.0">
<elevation id="24F8D195FFD4FF9AFE5F4BCAFDA6FE4F" box="[500,571,439,463]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="m" value="800.0">800 m</elevation>
</quantity>
elevation, on
<date id="DB6B1066FFD4FF9AFD7C4BCAFCEEFE4F" box="[727,883,439,463]" pageId="4" pageNumber="5" value="2004-03-01">
<collectingDate id="CB2FE98EFFD4FF9AFD7C4BCAFCEEFE4F" box="[727,883,439,463]" pageId="4" pageNumber="5" value="2004-03-01">1 March 2004</collectingDate>
</date>
by
<collectorName id="02205370FFD4FF9AFC374BC5FB83FE4F" box="[924,1054,440,463]" pageId="4" pageNumber="5">D. Emmett.</collectorName>
</materialsCitation>
<materialsCitation id="1FBD3CFBFFD4FF9AFB8F4BC5FC9DFDBB" collectingDate="2004-03-01" collectingDateMax="2004-08-24" collectingDateMin="2004-03-01" collectorName="D. Emmett. &amp; D. Emmett" country="Cambodia" county="Thmar Baing District" elevation="5" latitude="11.63" location="Central Cardamoms Protected Forest" longLatPrecision="5469" longitude="103.5" municipality="Central Cardamoms Protected Forest" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Koh Kong">
FMNH 267732, female, collected at 2000 hrs on a tree trunk
<quantity id="682D9B43FFD4FF9AFE694BA1FD9FFE73" box="[450,514,476,500]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">
<elevation id="24F8D195FFD4FF9AFE694BA1FD9FFE73" box="[450,514,476,500]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="0.5">0.5 m</elevation>
</quantity>
above the ground,
<quantity id="682D9B43FFD4FF9AFD734BA1FC8DFE73" box="[728,784,476,500]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">
<elevation id="24F8D195FFD4FF9AFD734BA1FC8DFE73" box="[728,784,476,500]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="10.0">10 m</elevation>
</quantity>
from a river, in seasonally flooded lowland evergreen forest,
<collectingMunicipality id="4F0EACDCFFD4FF9AFF6D4B82FDF1FD97" box="[198,620,511,535]" pageId="4" pageNumber="5">Central Cardamoms Protected Forest</collectingMunicipality>
,
<collectingCounty id="460B4E2AFFD4FF9AFDD34B82FCF3FD97" box="[632,878,511,535]" pageId="4" pageNumber="5">Thmar Baing District</collectingCounty>
,
<collectingRegion id="6D11F844FFD4FF9AFCD04B82FBFDFD97" box="[891,1120,511,536]" country="Cambodia" name="Koh Kong" pageId="4" pageNumber="5">Koh Kong Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD4FF9AFBC54B82FB7CFD97" box="[1134,1249,511,535]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
(
<geoCoordinate id="CAE15061FFD4FF9AFB5A4B82FACDFD97" box="[1265,1360,511,535]" degrees="11.63" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="11.63">11.63 N</geoCoordinate>
,
<geoCoordinate id="CAE15061FFD4FF9AFAF64B82FF36FDBC" degrees="103.5" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="5555" value="103.5">103.5 E</geoCoordinate>
),
<quantity id="682D9B43FFD4FF9AFF144859FE9BFDBB" box="[191,262,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="5" unit="m" value="180.0">
<elevation id="24F8D195FFD4FF9AFF144859FE9BFDBB" box="[191,262,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="5" unit="m" value="180.0">180 m</elevation>
</quantity>
elevation, on
<date id="DB6B1066FFD4FF9AFE0F4859FDCAFDBC" box="[420,599,548,572]" pageId="4" pageNumber="5" value="2004-08-24">
<collectingDate id="CB2FE98EFFD4FF9AFE0F4859FDCAFDBC" box="[420,599,548,572]" pageId="4" pageNumber="5" value="2004-08-24">24 August 2004</collectingDate>
</date>
by
<collectorName id="02205370FFD4FF9AFD2A4859FC9DFDBB" box="[641,768,548,571]" pageId="4" pageNumber="5">D. Emmett</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="AF6A36A6FFD4FF9AFF6E483AFDCCFD6F" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<emphasis id="9DA1EAB4FFD4FF9AFF6E483AFE2AFDE0" bold="true" box="[197,439,583,608]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Additional material.</emphasis>
<materialsCitation id="1FBD3CFBFFD4FF9AFE164835FDD0FD6F" collectingDate="1998-11-11" country="Thailand" county="Khao Kitchakut National Park" elevation="75" latitude="12.8" location="Two" longLatPrecision="775" longitude="102.13" municipality="Krathing Waterfall" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Chanthaburi">
<location id="AA0A607DFFD4FF9AFE164835FE73FDDF" LSID="urn:lsid:plazi:treatment:277C87B0FFD2FF96FF3C4CB0FA83FD03:AA0A607DFFD4FF9AFE164835FE73FDDF" box="[445,494,584,607]" country="Thailand" county="Khao Kitchakut National Park" latitude="12.8" longLatPrecision="775" longitude="102.13" municipality="Krathing Waterfall" name="Two" pageId="4" pageNumber="5" stateProvince="Chanthaburi">Two</location>
live specimens, an adult male and female, collected at 2215 hrs in low vegetation by the fourth tier of
<collectingMunicipality id="4F0EACDCFFD4FF9AFEC24811FDDEFD04" box="[361,579,620,644]" pageId="4" pageNumber="5">Krathing Waterfall</collectingMunicipality>
,
<collectingCounty id="460B4E2AFFD4FF9AFDF94811FC2BFD04" box="[594,950,620,644]" pageId="4" pageNumber="5">Khao Kitchakut National Park</collectingCounty>
,
<collectingRegion id="6D11F844FFD4FF9AFC6D4811FB28FD04" box="[966,1205,620,644]" country="Thailand" name="Chanthaburi" pageId="4" pageNumber="5">Chantaburi Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD4FF9AFB6E4811FAB5FD04" box="[1221,1320,620,644]" name="Thailand" pageId="4" pageNumber="5">Thailand</collectingCountry>
(
<geoCoordinate id="CAE15061FFD4FF9AFA974811FA01FD03" box="[1340,1436,620,644]" degrees="12.80" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="555" value="12.8">12.80 N</geoCoordinate>
<geoCoordinate id="CAE15061FFD4FF9AFF3C48F2FE9CFD27" box="[151,257,655,679]" degrees="102.13" direction="east" orientation="longitude" pageId="4" pageNumber="5" precision="555" value="102.13">102.13 E</geoCoordinate>
),
<quantity id="682D9B43FFD4FF9AFEBC48F2FECFFD27" box="[279,338,655,679]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="5" unit="m" value="75.0">
<elevation id="24F8D195FFD4FF9AFEBC48F2FECFFD27" box="[279,338,655,679]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="5" unit="m" value="75.0">75 m</elevation>
</quantity>
elevation, on
<date id="DB6B1066FFD4FF9AFE5C48F2FD49FD27" box="[503,724,655,679]" pageId="4" pageNumber="5" value="1998-11-11">
<collectingDate id="CB2FE98EFFD4FF9AFE5C48F2FD49FD27" box="[503,724,655,679]" pageId="4" pageNumber="5" value="1998-11-11">11 November 1998</collectingDate>
</date>
, by AM and RST. Data from these living specimens has been used in all analyses, but under the conditions of permission, these specimens could not be preserved as vouchers and were released at the site of capture
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFD4FF9AFF6E4886FD30F814" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="AF6A36A6FFD4FF9AFF6E4886FD30F814" blockId="4.[151,1437,151,1940]" pageId="4" pageNumber="5">
<emphasis id="9DA1EAB4FFD4FF9AFF6E4886FEA2FC93" bold="true" box="[197,319,763,788]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Diagnosis.</emphasis>
<taxonomicName id="68D54D25FFD4FF9AFEE24881FD07FC94" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[329,666,764,788]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
<emphasis id="9DA1EAB4FFD4FF9AFEE24881FD07FC94" box="[329,666,764,788]" italics="true" pageId="4" pageNumber="5">Cryptelytrops cardamomensis</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFD4FF9AFD0F4881FC9DFC94" bold="true" box="[676,768,764,788]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="869257CFFFD4FF9AFD0F4881FC9DFC94" box="[676,768,764,788]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="68D54D25FFD4FF9AFBAE4881FB14FC94" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[1029,1161,764,788]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="venustus">
<emphasis id="9DA1EAB4FFD4FF9AFBAE4881FB14FC94" box="[1029,1161,764,788]" italics="true" pageId="4" pageNumber="5">C. venustus</emphasis>
</taxonomicName>
,
<taxonomicName id="68D54D25FFD4FF9AFB314881FADBFC94" box="[1178,1350,764,788]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="honsonensis">
<emphasis id="9DA1EAB4FFD4FF9AFB314881FADBFC94" box="[1178,1350,764,788]" italics="true" pageId="4" pageNumber="5">C. honsonensis</emphasis>
</taxonomicName>
and
<taxonomicName id="68D54D25FFD4FF9AFA294881FEBBFCB8" baseAuthorityName="Malhotra &amp; Thorpe" baseAuthorityYear="2004" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="kanburiensis">
<emphasis id="9DA1EAB4FFD4FF9AFA294881FEBBFCB8" italics="true" pageId="4" pageNumber="5">C. kanburiensis</emphasis>
</taxonomicName>
, by the absence of purple-red marking (banding on the body and blotches on the head). It is distinguished from all other Asian pitviper species which have the typical &quot;green pitviper&quot; colouration (uniform green dorsal colour and a lateral stripe present on the first few dorsal scale rows in one or both sexes), except other species of
<taxonomicName id="68D54D25FFD4FF9AFF4249F1FE1FFC24" authorityName="Cope" authorityYear="1860" box="[233,386,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="9DA1EAB4FFD4FF9AFF4249F1FE1FFC24" box="[233,386,908,932]" italics="true" pageId="4" pageNumber="5">Cryptelytrops</emphasis>
</taxonomicName>
(including
<emphasis id="9DA1EAB4FFD4FF9AFDAF49F1FCB7FC24" box="[516,810,908,932]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFDAF49F1FD07FC24" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[516,666,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
,
<taxonomicName id="68D54D25FFD4FF9AFD0C49F1FCB7FC24" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[679,810,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
</emphasis>
and
<taxonomicName id="68D54D25FFD4FF9AFCC949F1FBB5FC24" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[866,1064,908,932]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="9DA1EAB4FFD4FF9AFCC949F1FBB5FC24" box="[866,1064,908,932]" italics="true" pageId="4" pageNumber="5">C. septentrionalis</emphasis>
</taxonomicName>
), by the presence of a fused first supralabial and nasal scale. It can be distinguished from
<emphasis id="9DA1EAB4FFD4FF9AFCA649CDFBADFC48" box="[781,1072,944,968]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFCA649CDFC3EFC48" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[781,931,944,968]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
,
<taxonomicName id="68D54D25FFD4FF9AFC0549CDFBADFC48" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[942,1072,944,968]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
</emphasis>
and
<taxonomicName id="68D54D25FFD4FF9AFBCD49CDFAB0FC48" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[1126,1325,944,968]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="9DA1EAB4FFD4FF9AFBCD49CDFAB0FC48" box="[1126,1325,944,968]" italics="true" pageId="4" pageNumber="5">C. septentrionalis</emphasis>
</taxonomicName>
primarily by the relatively larger size of the eye (most obvious in adults), the relatively wider supraoculars, and the shape of the head. The latter is more elongate
<emphasis id="9DA1EAB4FFD4FF9AFD904984FDD4FB90" box="[571,585,1017,1040]" italics="true" pageId="4" pageNumber="5"></emphasis>
oval in
<emphasis id="9DA1EAB4FFD4FF9AFD0A4985FC56FB90" box="[673,971,1016,1040]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFD0A4985FCA7FB90" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[673,826,1016,1040]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
,
<taxonomicName id="68D54D25FFD4FF9AFCEC4985FC56FB90" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[839,971,1016,1040]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
</emphasis>
and
<emphasis id="9DA1EAB4FFD4FF9AFBAD4985FB4BFB90" box="[1030,1238,1016,1040]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFBAD4985FB4CFB90" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[1030,1233,1016,1040]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="septentrionalis">C. septentrionalis</taxonomicName>
,
</emphasis>
but widens quite abruptly behind the eyes in
<emphasis id="9DA1EAB4FFD4FF9AFE7B4E61FCC7FBB4" box="[464,858,1052,1076]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFE7B4E61FD3FFBB4" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[464,674,1052,1076]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">C. cardamomensis</taxonomicName>
,
<taxonomicName id="68D54D25FFD4FF9AFD054E61FCB2FBB4" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[686,815,1052,1076]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFC9C4E60FCC7FBB4" box="[823,858,1053,1076]" pageId="4" pageNumber="5" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="68D54D25FFD4FF9AFC394E61FC62FBB4" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[914,1023,1052,1076]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
<emphasis id="9DA1EAB4FFD4FF9AFC394E61FC62FBB4" box="[914,1023,1052,1076]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFD4FF9AFBAC4E61FBFCFBB4" bold="true" box="[1031,1121,1052,1076]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="869257CFFFD4FF9AFBAC4E61FBFCFBB4" box="[1031,1121,1052,1076]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(described below) to give a characteristically triangular shaped head.
<taxonomicName id="68D54D25FFD4FF9AFDCC4E3DFC2BFBD8" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[615,950,1088,1112]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD4FF9AFDCC4E3DFC2BFBD8" box="[615,950,1088,1112]" italics="true" pageId="4" pageNumber="5">Cryptelytrops cardamomensis</emphasis>
</taxonomicName>
can be distinguished from both
<emphasis id="9DA1EAB4FFD4FF9AFAB74E3DFF27FBFC" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFAB74E3DFA01FBD8" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[1308,1436,1088,1112]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFF3C4E18FF27FBFC" box="[151,186,1125,1148]" pageId="4" pageNumber="5" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="68D54D25FFD4FF9AFF5B4E19FEC1FBFC" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[240,348,1124,1148]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD4FF9AFF5B4E19FEC1FBFC" box="[240,348,1124,1148]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
using a combination of the following characters (for further details see
<tableCitation id="E257031DFFD4FF9AFBD24E19FB50FBFC" box="[1145,1229,1124,1148]" captionStart="TABLE 2" captionStartId="9.[151,234,358,381]" captionTargetPageId="9" captionText="TABLE 2. Mean values and standard deviations of morphological characters that in combination can be used to distinguish C. cardamomensis sp. nov. from both C. macrops s.s. and C. rubeus sp. nov. Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm = C. macrops s.s., Cc = C. cardamomensis, Cr = C. rubeus. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from C. cardamomensis (e.g. BTWSUPOC can be used to discriminate both C. macrops and C. rubeus from C. cardamomensis while DV19to17 is only useful to discriminate C. rubeus from C. cardamomensis)." pageId="4" pageNumber="5">Table 2</tableCitation>
). In males,
<taxonomicName id="68D54D25FFD4FF9AFAE54E19FEBCFB20" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD4FF9AFAE54E19FEBCFB20" italics="true" pageId="4" pageNumber="5">C. cardamomensis</emphasis>
</taxonomicName>
tends to have more scales between the rear edges of the supraoculars (BTWSUPOC2), a relatively larger eye (DEYE), and a more prominent postocular white stripe (SCROC) than either
<emphasis id="9DA1EAB4FFD4FF9AFB094ED1FAD0FB44" box="[1186,1357,1196,1220]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFB094ED1FABBFB44" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[1186,1318,1196,1220]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFA9A4ED0FAD0FB44" box="[1329,1357,1197,1220]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
</emphasis>
, or
<taxonomicName id="68D54D25FFD4FF9AFA294ED1FF7EFB68" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD4FF9AFA294ED1FF7EFB68" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
. Furthermore, compared to
<emphasis id="9DA1EAB4FFD4FF9AFD8F4EADFD51FB68" box="[548,716,1232,1256]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFD8F4EADFD3BFB68" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[548,678,1232,1256]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFD044EACFD51FB68" box="[687,716,1233,1256]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
</emphasis>
, the scale reduction from 19 to 17 scale rows around the body (VS19TO17) tends to occur further away from the head, it tends to have fewer (larger) scales bordering the supralabial scales (BORSUPOC), and less keeled temporal scales (KTEMP). Compared to
<taxonomicName id="68D54D25FFD4FF9AFBE74F65FB24FAB0" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[1100,1209,1304,1328]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD4FF9AFBE74F65FB24FAB0" box="[1100,1209,1304,1328]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
, the scale reduction from 19 to 17 scale rows around the body tends to involve higher (i.e., more dorsal) scale rows (DV19TO17); it tends to have more keeled body scales at mid-body (BSCK), and more scales between the last supralabial and the chin shields (VENTEDGE). In females, the lateral white stripe is much more prominent (STRIPE), always involving the first two dorsal scale rows. Furthermore, it tends to cover a larger proportion of the first scale row (SCR1) than in females of
<taxonomicName id="68D54D25FFD4FF9AFECD4FB1FE78FA64" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[358,485,1484,1508]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFD4FF9AFECD4FB1FE78FA64" box="[358,485,1484,1508]" italics="true" pageId="4" pageNumber="5">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFE474FB0FD8DFA63" box="[492,528,1484,1508]" pageId="4" pageNumber="5" sensu="stricto">
<emphasis id="9DA1EAB4FFD4FF9AFE474FB0FD94FA64" box="[492,521,1485,1508]" italics="true" pageId="4" pageNumber="5">s.s</emphasis>
.
</taxonomicNameLabel>
A small scale is present between the nasal scale and the scale bordering the anterior edge of the pit (NASPIT), whereas this is infrequently present in
<emphasis id="9DA1EAB4FFD4FF9AFC664F8DFBE2F988" box="[973,1151,1520,1544]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFC664F8DFBCCF988" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[973,1105,1520,1544]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFBF74F8CFBE2F988" box="[1116,1151,1521,1544]" pageId="4" pageNumber="5" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and never present in
<emphasis id="9DA1EAB4FFD4FF9AFA294F8DFDDFF9AC" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFA294F8DFF7AF9AC" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">C. rubeus</taxonomicName>
.
<taxonomicName id="68D54D25FFD4FF9AFF594C69FDDFF9AC" box="[242,578,1556,1580]" class="Reptilia" family="Viperidae" genus="Crytpelytrops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">Crytpelytrops cardamomensis</taxonomicName>
</emphasis>
tends to have more (smaller) scales bordering the supralabial scales (BORSUPOC) than
<taxonomicName id="68D54D25FFD4FF9AFE974C45FE37F9D0" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[316,426,1592,1616]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD4FF9AFE974C45FE37F9D0" box="[316,426,1592,1616]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
, but fewer (larger) than in
<taxonomicName id="68D54D25FFD4FF9AFD4A4C45FCFEF9D0" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[737,867,1592,1616]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFD4FF9AFD4A4C45FCFEF9D0" box="[737,867,1592,1616]" italics="true" pageId="4" pageNumber="5">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFCC04C44FC12F9CF" box="[875,911,1592,1616]" pageId="4" pageNumber="5" sensu="stricto">
<emphasis id="9DA1EAB4FFD4FF9AFCC04C44FC15F9D0" box="[875,904,1593,1616]" italics="true" pageId="4" pageNumber="5">s.s</emphasis>
.
</taxonomicNameLabel>
The scale reduction from 19 to 17 scale rows around the body (DV19TO17) tends to involve lower scale rows in
<taxonomicName id="68D54D25FFD4FF9AFC3E4C21FBF8F9F4" box="[917,1125,1628,1652]" class="Reptilia" family="Viperidae" genus="Crytpelytrops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD4FF9AFC3E4C21FBF8F9F4" box="[917,1125,1628,1652]" italics="true" pageId="4" pageNumber="5">C. cardamomensis</emphasis>
</taxonomicName>
and the fusion between the first labial and nasal scale tends to be more complete (LABNAS) and involve partial sutures on both sides of the nostril (compare
<figureCitation id="37EE2A23FFD4FF9AFEF44CD9FE2CF93C" box="[351,433,1700,1724]" captionStart="FIGURE 3" captionStartId="5.[151,250,1943,1966]" captionTargetBox="[287,1311,209,1911]" captionTargetId="figure-66@5.[268,1319,193,1921]" captionTargetPageId="5" captionText="FIGURE 3. Line drawings of the head scalation of the holotype of Cryptelytrops cardamomensis sp. nov. (FMNH 259191) A. lateral, B. dorsal, C. ventral. Small letters identify the following measurements: a (eye width): 3.3mm, b (supraocular width): 1.55mm, c (bottom edge of rostral): 2.5mm, d: (head length): 18.75 mm." figureDoi="http://doi.org/10.5281/zenodo.5291121" httpUri="https://zenodo.org/record/5291121/files/figure.png" pageId="4" pageNumber="5">Figs. 3</figureCitation>
,
<figureCitation id="37EE2A23FFD4FF9AFE684CD9FE4CF93C" box="[451,465,1700,1724]" captionStart="FIGURE 5" captionStartId="10.[151,250,1972,1995]" captionTargetBox="[253,1314,199,1920]" captionTargetId="figure-49@10.[246,1329,193,1928]" captionTargetPageId="10" captionText="FIGURE 5. Line drawings of the head scalation of the holotype of Cryptelytrops rubeus sp. nov. (FMNH 262718). Labelling as in Fig. 3 (a: 4.00mm, b: 2.05mm, c: 3.45mm, d: 24.9mm)." figureDoi="http://doi.org/10.5281/zenodo.5291125" httpUri="https://zenodo.org/record/5291125/files/figure.png" pageId="4" pageNumber="5">5</figureCitation>
and
<figureCitation id="37EE2A23FFD4FF9AFDBB4CD9FDBDF93C" box="[528,544,1700,1724]" captionStart="FIGURE 7" captionStartId="16.[151,250,1976,1999]" captionTargetBox="[307,1289,201,1941]" captionTargetId="figure-49@16.[283,1298,193,1942]" captionTargetPageId="16" captionText="FIGURE 7. Line drawings of the head scalation of the holotype of Cryptelytrops macrops s. s. (MHNG 1400.85). Labelling as in Fig. 3 (a: 4.15mm, b: 2.30mm, c: 3.00mm, d: 23.2mm)." figureDoi="http://doi.org/10.5281/zenodo.5291131" httpUri="https://zenodo.org/record/5291131/files/figure.png" pageId="4" pageNumber="5">7</figureCitation>
), than in either
<emphasis id="9DA1EAB4FFD4FF9AFD494CD9FC12F93C" box="[738,911,1700,1724]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFD494CD9FCFBF93C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[738,870,1700,1724]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFCD94CD8FC12F93C" box="[882,911,1701,1724]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
</emphasis>
, or
<taxonomicName id="68D54D25FFD4FF9AFC6F4CD9FBABF93C" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[964,1078,1700,1724]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD4FF9AFC6F4CD9FBABF93C" box="[964,1078,1700,1724]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
. Furthermore, compared to
<emphasis id="9DA1EAB4FFD4FF9AFA294CD9FE81F960" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="68D54D25FFD4FF9AFA294CD9FF6BF960" baseAuthorityName="Kramer" baseAuthorityYear="1977" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD4FF9AFF544CB4FE81F960" box="[255,284,1737,1760]" pageId="4" pageNumber="5" sensu="stricto">s.s</taxonomicNameLabel>
</emphasis>
,, the scale reduction from 19 to 17 scale rows around the body (VS19TO17) tends to occur further away from the head, it tends to have more scales between the rear edges of the supraoculars (BTWSUPOC2), a higher number of sublabial scales (SUBLAB), and a relatively smaller eye (DEYE). Compared to
<taxonomicName id="68D54D25FFD4FF9AFB4E4D6DFACFF8A8" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[1253,1362,1808,1832]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD4FF9AFB4E4D6DFACFF8A8" box="[1253,1362,1808,1832]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
, it has more keeled body scales at mid-body (BSCK), the scale reduction from 12 to 10 scale rows around the tail (SC12O10) also tends to occur further away from the vent, and there are a larger number of ventral scales in
<taxonomicName id="68D54D25FFD4FF9AFAE54D25FEBFF814" class="Reptilia" family="Viperidae" genus="Crytpelytrops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD4FF9AFAE54D25FEBFF814" italics="true" pageId="4" pageNumber="5">C. cardamomensis</emphasis>
</taxonomicName>
females than in
<taxonomicName id="68D54D25FFD4FF9AFE714D01FDDAF814" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[474,583,1916,1940]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD4FF9AFE714D01FDDAF814" box="[474,583,1916,1940]" italics="true" pageId="4" pageNumber="5">C. rubeus</emphasis>
</taxonomicName>
females.
</paragraph>
</subSubSection>
<caption id="FBAA662EFFD5FF9BFF3C4DEAFCEBF86B" ID-DOI="http://doi.org/10.5281/zenodo.5291121" ID-Zenodo-Dep="5291121" httpUri="https://zenodo.org/record/5291121/files/figure.png" pageId="5" pageNumber="6" startId="5.[151,250,1943,1966]" targetBox="[287,1311,209,1911]" targetPageId="5">
<paragraph id="AF6A36A6FFD5FF9BFF3C4DEAFCEBF86B" blockId="5.[151,1436,1943,2027]" pageId="5" pageNumber="6">
<emphasis id="9DA1EAB4FFD5FF9BFF3C4DEAFE8FF82E" bold="true" box="[151,274,1943,1966]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
Line drawings of the head scalation of the holotype of
<taxonomicName id="68D54D25FFD5FF9BFCEF4DE5FBEFF82E" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[836,1138,1944,1966]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
<emphasis id="9DA1EAB4FFD5FF9BFCEF4DE5FBEFF82E" box="[836,1138,1944,1966]" italics="true" pageId="5" pageNumber="6">Cryptelytrops cardamomensis</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFD5FF9BFBD24DE5FB57F82D" bold="true" box="[1145,1226,1944,1965]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="869257CFFFD5FF9BFBD24DE5FB57F82D" box="[1145,1226,1944,1965]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(FMNH 259191) A. lateral, B. dorsal, C. ventral. Small letters identify the following measurements: a (eye width): 3.3mm, b (supraocular width): 1.55mm, c (bottom edge of rostral): 2.5mm, d: (head length): 18.75 mm.
</paragraph>
</caption>
<caption id="FBAA662EFFD6FF98FF3C4D06FC57F86E" ID-DOI="http://doi.org/10.5281/zenodo.5291123" ID-Zenodo-Dep="5291123" httpUri="https://zenodo.org/record/5291123/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1915,1938]" targetBox="[250,1338,202,1900]" targetPageId="6">
<paragraph id="AF6A36A6FFD6FF98FF3C4D06FC57F86E" blockId="6.[151,1436,1915,2030]" pageId="6" pageNumber="7">
<emphasis id="9DA1EAB4FFD6FF98FF3C4D06FE89F812" bold="true" box="[151,276,1915,1938]" pageId="6" pageNumber="7">FIGURE 4.</emphasis>
<taxonomicName id="68D54D25FFD6FF98FEB74D01FDD0F812" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[284,589,1916,1938]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
<emphasis id="9DA1EAB4FFD6FF98FEB74D01FDD0F812" box="[284,589,1916,1938]" italics="true" pageId="6" pageNumber="7">Cryptelytrops cardamomensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD6FF98FDFE4D00FD00F812" box="[597,669,1917,1938]" pageId="6" pageNumber="7" rank="species">sp.nov.</taxonomicNameLabel>
A. Close-up of head and mid-body region of adult male (left) and female (right) from Khao Kitchakut National Park, Chantaburi Prov., Thailand (photos: A Malhotra). B. Adult male (left) and female (right) from Kâmpóng Saôm Bay, Koh Kong Prov., Cambodia (photos: A Gumprecht); C. and D. Adult specimens from Mount Da Lai in Cardamom Mountains, southwest Cambodia (photos: Jeremy Holden).
</paragraph>
</caption>
<subSubSection id="E7CF652DFFD7FF96FF6E4AEAFC7CFEBF" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="AF6A36A6FFD7FF99FF6E4AEAFEFAFD4C" blockId="7.[151,1436,151,716]" pageId="7" pageNumber="8">
<emphasis id="9DA1EAB4FFD7FF99FF6E4AEAFE79FF2F" bold="true" box="[197,484,151,176]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">
Description of
<typeStatus id="706E8804FFD7FF99FEDD4AE5FE40FF30" box="[374,477,152,176]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
.
</emphasis>
Body cylindrical, head triangular in dorsal aspect and very distinct from neck. Canthus rostralis distinct. Head scales small (no large dorsal shields), smooth except in temporal region, region between the suprabials and temporal region, and on the rear of head, which show weak tubercular keeling. Dorsal body scales weakly keeled. 171 ventral scales, 63 pairs subcaudal scales, 21 dorsal scales at mid
<emphasis id="9DA1EAB4FFD7FF99FB5D4B78FA99FE9C" box="[1270,1284,261,284]" italics="true" pageId="7" pageNumber="8"></emphasis>
body. Rostral scale roughly triangular, upper edge about half the width of lower edge. Pupil vertically elliptical. Loreal pit present. Nostril completely enclosed in nasal scale. Nasal scale partially fused with first supralabial, bearing a suture present behind the nostril, but not anterior to it. Ten and 11 supralabials on the right and left sides respectively, similarly 13/12 sublabials, three postocular scales. Shield bordering anterior edge of pit fused with second supralabial, but separated from nasal by one small scale on either side. Subocular scale touches third supralabial, but separated by one scale from fourth and fifth supralabials. At least 11 scales between supraoculars, with 14 scales between their posterior edges. Five teeth on palatine, 16 on pterygoid, and 16 on dentary. Snout
<emphasis id="9DA1EAB4FFD7FF99FBDD487CFB19FD98" box="[1142,1156,513,536]" italics="true" pageId="7" pageNumber="8"></emphasis>
vent length
<quantity id="682D9B43FFD7FF99FAA74B82FAF7FD97" box="[1292,1386,511,535]" metricMagnitude="-1" metricUnit="m" metricValue="2.91" pageId="7" pageNumber="8" unit="cm" value="29.1">29.1 cm</quantity>
; tail length
<quantity id="682D9B43FFD7FF99FF4D4859FEA5FDBB" box="[230,312,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="8" unit="cm" value="5.6">5.6 cm</quantity>
. Head length measured from tip of snout to posterior edge of lower jawbone
<quantity id="682D9B43FFD7FF99FB314859FA8CFDBB" box="[1178,1297,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.875" pageId="7" pageNumber="8" unit="mm" value="18.75">18.75 mm</quantity>
; head width measured between rear outer edges of supraoculars
<quantity id="682D9B43FFD7FF99FD45483AFCD4FDDF" box="[750,841,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="7" pageNumber="8" unit="mm" value="8.3">8.3 mm</quantity>
, and at widest point of head
<quantity id="682D9B43FFD7FF99FB35483AFA9AFDDF" box="[1182,1287,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="7" pageNumber="8" unit="mm" value="12.9">12.9 mm</quantity>
. Supraocular scale (on right hand side)
<quantity id="682D9B43FFD7FF99FE144811FD88FD03" box="[447,533,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="8" unit="mm" value="3.9">3.9 mm</quantity>
long,
<quantity id="682D9B43FFD7FF99FDF44811FD59FD03" box="[607,708,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="7" pageNumber="8" unit="mm" value="1.55">1.55 mm</quantity>
wide. Distance from eye to nostril
<quantity id="682D9B43FFD7FF99FBFC4811FB23FD03" box="[1111,1214,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="7" pageNumber="8" unit="mm" value="3.85">3.85 mm</quantity>
, eye to pit
<quantity id="682D9B43FFD7FF99FA954811FA0AFD03" box="[1342,1431,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="mm" value="0.9">0.9 mm</quantity>
, posterior edge of pit to anterior edge of nostril
<quantity id="682D9B43FFD7FF99FD0D48F2FC9DFD27" box="[678,768,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="8" unit="mm" value="3.6">3.6 mm</quantity>
. Eye diameter
<quantity id="682D9B43FFD7FF99FC0C48F2FB9DFD27" box="[935,1024,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="8" unit="mm" value="3.3">3.3 mm</quantity>
. Scale reduction formula (derived as in
<bibRefCitation id="CB444B57FFD7FF99FF1848C9FEC7FD4C" author="Thorpe, R. S." box="[179,346,692,716]" pageId="7" pageNumber="8" pagination="27 - 43" refId="ref12721" refString="Thorpe, R. S. (1975) Quantitative handling of characters useful in snake systematics with particular reference to intraspecific variation in the ringed snake Natrix natrix (L.). Biological Journal of the Linnean Society, 7, 27 - 43." type="journal article" year="1975">Thorpe [1975]</bibRefCitation>
):
</paragraph>
<paragraph id="AF6A36A6FFD7FF99FF6E491AFEABFB8F" blockId="7.[151,1437,871,2012]" pageId="7" pageNumber="8">
<emphasis id="9DA1EAB4FFD7FF99FF6E491AFDFEFCFF" bold="true" box="[197,611,871,896]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">
Colour of
<typeStatus id="706E8804FFD7FF99FE954915FE38FC00" box="[318,421,872,896]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dark brownish grey dorsally, the ventral surface somewhat paler than the dorsal surface towards anterior of body, but less distinct posteriorly. Pale lateral stripe well developed, covering approximately one
<emphasis id="9DA1EAB4FFD7FF99FEC149CCFEE5FC48" box="[362,376,945,968]" italics="true" pageId="7" pageNumber="8"></emphasis>
third of first dorsal scale row, increasing in width towards head. Upper and lower labials not distinct in colour from the rest of the head, but chin shields, scales between chin shields and sublabial scales are creamy white.
</paragraph>
<paragraph id="AF6A36A6FFD7FF99FF6E4E66FAAEF987" blockId="7.[151,1437,871,2012]" pageId="7" pageNumber="8">
<emphasis id="9DA1EAB4FFD7FF99FF6E4E66FE59FBB4" box="[197,452,1051,1076]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">
<emphasis id="9DA1EAB4FFD7FF99FF6E4E66FEFFFBB4" bold="true" box="[197,354,1051,1076]" pageId="7" pageNumber="8">Colour in life</emphasis>
(
<figureCitation id="37EE2A23FFD7FF99FED94E61FE2AFBB4" box="[370,439,1052,1076]" captionStart="FIGURE 4" captionStartId="6.[151,250,1915,1938]" captionTargetBox="[250,1338,202,1900]" captionTargetId="figure-82@6.[242,1344,193,1903]" captionTargetPageId="6" captionText="FIGURE 4. Cryptelytrops cardamomensis sp.nov. A. Close-up of head and mid-body region of adult male (left) and female (right) from Khao Kitchakut National Park, Chantaburi Prov., Thailand (photos: A Malhotra). B. Adult male (left) and female (right) from Kâmpóng Saôm Bay, Koh Kong Prov., Cambodia (photos: A Gumprecht); C. and D. Adult specimens from Mount Da Lai in Cardamom Mountains, southwest Cambodia (photos: Jeremy Holden)." figureDoi="http://doi.org/10.5281/zenodo.5291123" httpUri="https://zenodo.org/record/5291123/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
).
</emphasis>
No notes or photographs were taken of the
<typeStatus id="706E8804FFD7FF99FC064E61FB92FBB4" box="[941,1039,1052,1076]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
in life. This description is based on photographs of the male and female specimens from Khao Kitchakut NP,
<collectingCountry id="D7C27636FFD7FF99FC434E42FBD6FBD7" box="[1000,1099,1087,1111]" name="Thailand" pageId="7" pageNumber="8">Thailand</collectingCountry>
as well as photographs on p. 230 Plate II and p. 232 Plate I (adult male from Khao Soi Dao wildlife sanctuary,
<collectingRegion id="6D11F844FFD7FF99FB824E19FA89FBFC" box="[1065,1300,1124,1148]" country="Thailand" name="Chanthaburi" pageId="7" pageNumber="8">Chantaburi Province</collectingRegion>
), and p. 233 Plate II, and p. 235 Pl II (adult male and female respectively, from
<collectingRegion id="6D11F844FFD7FF99FC274EF5FBF2FB20" box="[908,1135,1160,1184]" country="Cambodia" name="Koh Kong" pageId="7" pageNumber="8">Koh Kong Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD7FF99FBD04EF5FB6DFB20" box="[1147,1264,1160,1184]" name="Cambodia" pageId="7" pageNumber="8">Cambodia</collectingCountry>
) of
<bibRefCitation id="CB444B57FFD7FF99FAB54EF5FEB8FB44" author="Gumprecht, A. &amp; Tillack, F. &amp; Orlov, N. L. &amp; Captain, A. &amp; Ryabov, S." pageId="7" pageNumber="8" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. &amp; Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
Gumprecht
<emphasis id="9DA1EAB4FFD7FF99FF3C4ED0FF54FB44" box="[151,201,1196,1220]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
. (2004)
</bibRefCitation>
. Dorsal and ventral surface grass green, very similar in shade. Lateral white stripe more prominent in male, extending onto the second dorsal scale row (confined to the first in female). In female, obvious bright blue dot on the upper rear quadrant of first scale row; present in some males, but less obvious. In male only, lateral white stripe extends onto the head as postocular stripe. Upper lip slightly paler than rest of dorsal surface of head, more obvious in female. Sublabials, and to lesser extent supralabials, also suffused with pale blue, mostly confined to edges of scales. Same pale blue also covers most scales on ventral surface of head to first part of body, more saturated and extensive on underside of head than in
<taxonomicName id="68D54D25FFD7FF99FD144FF9FCA2FA1C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[703,831,1412,1436]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFD7FF99FD144FF9FCA2FA1C" box="[703,831,1412,1436]" italics="true" pageId="7" pageNumber="8">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD7FF99FCED4FF9FCF7FA1B" box="[838,874,1412,1435]" pageId="7" pageNumber="8" sensu="stricto">s.s.</taxonomicNameLabel>
Occasional small patches of white and green pigment on scales of underside of head. Interstitial skin bright blue, black banding sometimes on anterior body, scales here have narrow edging of black pigment at base. Tail solid brick red, to vent on dorsal surface, approximately one-third (male) to half (female) distance to the vent on ventral surface. Eye golden yellow in both sexes.
</paragraph>
<paragraph id="AF6A36A6FFD7FF96FF6E4C6EFC7CFEBF" blockId="7.[151,1437,871,2012]" lastBlockId="8.[151,1436,151,644]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="9DA1EAB4FFD7FF99FF6E4C6EFEA1F9AB" bold="true" box="[197,316,1555,1580]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Variation.</emphasis>
Sexual dimorphism is not very obvious in coloration. Females never have postocular stripes, but they may be only indistinct white streaks in males. The lateral white stripe may be much more prominent in males. Females have relatively shorter tails and fewer subcaudals (
<quantity id="682D9B43FFD7FF99FC844C21FC0EF9F4" box="[815,915,1628,1652]" metricMagnitude="0" metricUnit="m" metricValue="1.5367000000000002" metricValueMax="1.6002000000000003" metricValueMin="1.4731999999999998" pageId="7" pageNumber="8" unit="in" value="60.5" valueMax="63.0" valueMin="58.0">
58
<emphasis id="9DA1EAB4FFD7FF99FCE74C20FCC7F9F4" box="[844,858,1629,1652]" italics="true" pageId="7" pageNumber="8"></emphasis>
63 in
</quantity>
females, versus
<quantity id="682D9B43FFD7FF99FBE64C21FB18F9F4" box="[1101,1157,1628,1652]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" pageId="7" pageNumber="8" unit="in" value="69.0">69 in</quantity>
the single male in which this count could be made). Females also are likely to reach an overall larger size (maximum recorded
<quantity id="682D9B43FFD7FF99FAAC4CFDFAFDF917" box="[1287,1376,1664,1688]" metricMagnitude="-1" metricUnit="m" metricValue="5.36" pageId="7" pageNumber="8" unit="cm" value="53.6">53.6 cm</quantity>
SVL versus
<quantity id="682D9B43FFD7FF99FF4D4CD9FEDFF93B" box="[230,322,1700,1724]" metricMagnitude="-1" metricUnit="m" metricValue="4.99" pageId="7" pageNumber="8" unit="cm" value="49.9">49.9 cm</quantity>
for males), but the sample is small. The number of ventral scales is similar in both sexes, varying between 169 and 173. Keeling of body scales at mid
<emphasis id="9DA1EAB4FFD7FF99FD504CB4FC94F960" box="[763,777,1737,1760]" italics="true" pageId="7" pageNumber="8"></emphasis>
body is usually strong. Temporal scales and scales in the region between the temporal scales and supralabials show weak tubercular keeling, and keeling is strong on the scales covering the rear of the head. The ratio between the lengths of the upper and lower edges of the rostral scale varies between 0.35 and 0.56. Supralabials vary between 10 and 11 and sublabials between 11 and 14. The suture between the nasal scale and first labial scale is always more extensive to the rear of the nostril, usually absent in front of the nostril, but there may be a slight kink in the anterior margin of the fused scale at the likely position. The minimum number of scales between supraoculars varies between 8
<emphasis id="9DA1EAB4FFD7FF99FC294DDCFC0DF838" box="[898,912,1953,1976]" italics="true" pageId="7" pageNumber="8"></emphasis>
12 and there are 13
<emphasis id="9DA1EAB4FFD7FF99FBCE4DDCFBEEF838" box="[1125,1139,1953,1976]" italics="true" pageId="7" pageNumber="8"></emphasis>
16 scales between the posterior edges of the supraoculars. The number of scales between the nasals and shield bordering the anterior of the pit varies between 0
<emphasis id="9DA1EAB4FFD8FF96FED14AE4FE15FF30" box="[378,392,153,176]" italics="true" pageId="8" pageNumber="9"></emphasis>
2, and there are 0
<emphasis id="9DA1EAB4FFD8FF96FDE64AE4FDC6FF30" box="[589,603,153,176]" italics="true" pageId="8" pageNumber="9"></emphasis>
1 scales separating the internasals. There are 0
<emphasis id="9DA1EAB4FFD8FF96FBC24AE4FBEAFF30" box="[1129,1143,153,176]" italics="true" pageId="8" pageNumber="9"></emphasis>
1 scales between the third supralabial and the subocular, one scale between the fourth supralabial, and 1
<emphasis id="9DA1EAB4FFD8FF96FC594AC0FB9DFF54" box="[1010,1024,189,212]" italics="true" pageId="8" pageNumber="9"></emphasis>
2 scales between the fifth supralabial and the subocular scale. Hemipenes were examined in FMNH 259192, in which they were both partly everted, and do not differ notably from those of
<taxonomicName id="68D54D25FFD8FF96FD8F4B79FD3EFE9C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[548,675,260,284]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFD8FF96FD8F4B79FD3EFE9C" box="[548,675,260,284]" italics="true" pageId="8" pageNumber="9">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD8FF96FD004B78FD52FE9B" box="[683,719,260,284]" pageId="8" pageNumber="9" sensu="stricto">
<emphasis id="9DA1EAB4FFD8FF96FD004B78FD5AFE9C" box="[683,711,261,284]" italics="true" pageId="8" pageNumber="9">s.s</emphasis>
.
</taxonomicNameLabel>
Teeth were also counted in this specimen, and were identical to the
<typeStatus id="706E8804FFD8FF96FF6B4B5AFEBFFEBF" box="[192,290,295,319]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
except for the number of pterygoid teeth, which numbered 14.
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFD8FF96FF6E4B36FDBDFE4F" pageId="8" pageNumber="9" type="distribution">
<paragraph id="AF6A36A6FFD8FF96FF6E4B36FDBDFE4F" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="9">
<emphasis id="9DA1EAB4FFD8FF96FF6E4B36FEC3FEE4" bold="true" box="[197,350,331,356]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Distribution.</emphasis>
<taxonomicName id="68D54D25FFD8FF96FEC14B31FD20FEE4" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[362,701,332,356]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD8FF96FEC14B31FD20FEE4" box="[362,701,332,356]" italics="true" pageId="8" pageNumber="9">Cryptelytrops cardamomensis</emphasis>
</taxonomicName>
is currently known from
<collectingRegion id="6D11F844FFD8FF96FC5A4B31FB47FEE4" box="[1009,1242,332,356]" country="Cambodia" name="Koh Kong" pageId="8" pageNumber="9">Koh Kong Province</collectingRegion>
in southwestern
<collectingCountry id="D7C27636FFD8FF96FF3C4B12FE97FE07" box="[151,266,367,391]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
and
<collectingRegion id="6D11F844FFD8FF96FEE94B12FDB4FE07" box="[322,553,367,391]" country="Thailand" name="Chanthaburi" pageId="8" pageNumber="9">Chantaburi Province</collectingRegion>
in southeastern
<collectingCountry id="D7C27636FFD8FF96FD494B12FCD4FE07" box="[738,841,367,391]" name="Thailand" pageId="8" pageNumber="9">Thailand</collectingCountry>
. (
<figureCitation id="37EE2A23FFD8FF96FCF74B12FC03FE07" box="[860,926,367,391]" captionStart="FIGURE 1" captionStartId="1.[151,250,1878,1901]" captionTargetBox="[176,1411,773,1840]" captionTargetId="figure-285@1.[173,1417,770,1842]" captionTargetPageId="1" captionText="FIGURE 1. Map of Southeast Asia, showing the distribution of the two new species and Cryptelytrops macrops s.s. Symbols indicate sampling localities used in the genetic analysis to define species. Grey circles: C. macrops s.s; open diamonds: C. cardamomensis sp. nov.; black triangles: C. rubeus sp. nov. The empty circle and triangle denotes localities of specimens that have either been referred to the corresponding species on the basis of photographs, or for which the locality is uncertain and requires confirmation." figureDoi="http://doi.org/10.5281/zenodo.5291115" httpUri="https://zenodo.org/record/5291115/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
). The species probably occurs in appropriate habitat throughout the Cardamom Mountains in southwestern
<collectingCountry id="D7C27636FFD8FF96FCF14BE9FC50FE2C" box="[858,973,404,428]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
and adjacent southeastern
<collectingCountry id="D7C27636FFD8FF96FAAB4BE9FAFBFE2C" box="[1280,1382,404,428]" name="Thailand" pageId="8" pageNumber="9">Thailand</collectingCountry>
, and offshore islands (e.g., Koh Chang).
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFD8FF96FF6E4BA1FF44FDE0" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="AF6A36A6FFD8FF96FF6E4BA1FF44FDE0" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="9">
<emphasis id="9DA1EAB4FFD8FF96FF6E4BA1FEB7FE73" bold="true" box="[197,298,476,500]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Ecology.</emphasis>
<taxonomicName id="68D54D25FFD8FF96FE9A4BA1FD1DFE74" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[305,640,476,500]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD8FF96FE9A4BA1FD1DFE74" box="[305,640,476,500]" italics="true" pageId="8" pageNumber="9">Cryptelytrops cardamomensis</emphasis>
</taxonomicName>
is currently known from low-lying coastal areas up to
<quantity id="682D9B43FFD8FF96FB4C4BA1FAB3FE73" box="[1255,1326,476,500]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="m" value="800.0">800 m</quantity>
elevation in semi-evergreen, evergreen, and mangrove/melaleuca swamp forest. All specimens were found within a couple of metres above the ground, and most within
<quantity id="682D9B43FFD8FF96FDD64859FD25FDBB" box="[637,696,548,572]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="10.0">10 m</quantity>
of water. No food items were found in the digestive tracts of the
<typeStatus id="706E8804FFD8FF96FF3C4835FF4BFDE0" box="[151,214,584,608]" pageId="8" pageNumber="9">types</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFD8FF96FF6E4811FA83FD03" box="[197,1310,620,644]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="AF6A36A6FFD8FF96FF6E4811FA83FD03" blockId="8.[151,1436,151,644]" box="[197,1310,620,644]" pageId="8" pageNumber="9">
<emphasis id="9DA1EAB4FFD8FF96FF6E4811FED1FD04" bold="true" box="[197,332,620,644]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Etymology.</emphasis>
The name refers to the mountain range wherein the
<typeStatus id="706E8804FFD8FF96FC3E4811FC58FD04" box="[917,965,620,644]" pageId="8" pageNumber="9">type</typeStatus>
locality of this species occurs.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,969 @@
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<mods:title id="93105403119ECDB07B88B869E373C2D8">Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia</mods:title>
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<mods:affiliation id="87AC84ADA568AE90C59EE9C9B6038814">North Carolina Museum of Natural Sciences, USA. E-mail: bryan. stuart @ ncdenr. gov</mods:affiliation>
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<paragraph id="AF6A36A6FFD8FF96FF3C48AAFCBBFD6F" blockId="8.[151,806,727,788]" box="[151,806,727,752]" pageId="8" pageNumber="9">
<taxonomicName id="68D54D25FFD8FF96FF3C48AAFE1AFD70" ID-CoL="8S4BR" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[151,391,727,752]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">Cryptelytrops rubeus</taxonomicName>
<taxonomicNameLabel id="869257CFFFD8FF96FE2548A5FE7AFD6F" box="[398,487,728,752]" pageId="8" pageNumber="9" rank="species">
<emphasis id="9DA1EAB4FFD8FF96FE2548A5FE7DFD70" bold="true" box="[398,480,728,752]" pageId="8" pageNumber="9">
<emphasis id="9DA1EAB4FFD8FF96FE2548A5FE32FD70" box="[398,431,728,752]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">sp.</emphasis>
nov
</emphasis>
.
</taxonomicNameLabel>
(Ruby-eyed Green Pitviper)
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFD8FF96FF3C4881FEE0FC94" box="[151,381,764,788]" pageId="8" pageNumber="9" type="description">
<paragraph id="AF6A36A6FFD8FF96FF3C4881FEE0FC94" blockId="8.[151,806,727,788]" box="[151,381,764,788]" pageId="8" pageNumber="9">
<figureCitation id="37EE2A23FFD8FF96FF3C4881FF44FC94" box="[151,217,764,788]" captionStart="FIGURE 5" captionStartId="10.[151,250,1972,1995]" captionTargetBox="[253,1314,199,1920]" captionTargetId="figure-49@10.[246,1329,193,1928]" captionTargetPageId="10" captionText="FIGURE 5. Line drawings of the head scalation of the holotype of Cryptelytrops rubeus sp. nov. (FMNH 262718). Labelling as in Fig. 3 (a: 4.00mm, b: 2.05mm, c: 3.45mm, d: 24.9mm)." figureDoi="http://doi.org/10.5281/zenodo.5291125" httpUri="https://zenodo.org/record/5291125/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
,
<figureCitation id="37EE2A23FFD8FF96FF4D4881FF6BFC94" box="[230,246,764,788]" captionStart="FIGURE 6" captionStartId="11.[151,250,1898,1921]" captionTargetBox="[235,1346,193,1865]" captionTargetId="figure-72@11.[232,1347,193,1869]" captionTargetPageId="11" captionText="FIGURE 6. Cryptelytrops rubeus sp.nov. A. Close-up of head of male (left) and female (right) from vicinity of Ho Chi Minh city, South Vietnam (photos: A Malhotra); B. Close-up of head of male (left) and female (right) from South Vietnam; C. Closeup of tail of male from B; D. Pair from B (male above) (photos: K Udomritthiruj)." figureDoi="http://doi.org/10.5281/zenodo.5291129" httpUri="https://zenodo.org/record/5291129/files/figure.png" pageId="8" pageNumber="9">6</figureCitation>
;
<tableCitation id="E257031DFFD8FF96FEA84881FEE0FC94" box="[259,381,764,788]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="2.[151,234,1482,1505]" captionStartId-1="9.[151,234,358,381]" captionStartId-2="13.[151,234,1575,1598]" captionTargetBox-2="[156,1155,1675,2028]" captionTargetPageId-0="2" captionTargetPageId-1="9" captionTargetPageId-2="13" captionText-0="TABLE 1. Sample sizes for each sex of the three species in the multivariate analyses." captionText-1="TABLE 2. Mean values and standard deviations of morphological characters that in combination can be used to distinguish C. cardamomensis sp. nov. from both C. macrops s.s. and C. rubeus sp. nov. Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm = C. macrops s.s., Cc = C. cardamomensis, Cr = C. rubeus. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from C. cardamomensis (e.g. BTWSUPOC can be used to discriminate both C. macrops and C. rubeus from C. cardamomensis while DV19to17 is only useful to discriminate C. rubeus from C. cardamomensis)." captionText-2="TABLE 3. Mean and standard deviations of morphological characters that in combination can be used to distinguish C. macrops s.s. and C. rubeus sp. nov. Data are for the maximum number of specimens available for that character. Cm = C. macrops s.s., Cr = C. rubeus. Character abbreviations are explained in Appendix 1." pageId="8" pageNumber="9">Tables 13</tableCitation>
</paragraph>
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<subSubSection id="E7CF652DFFD8FF96FF3C4939FCFEFC17" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="AF6A36A6FFD8FF96FF3C4939FC1AFCF8" blockId="8.[151,1437,835,919]" pageId="8" pageNumber="9">
<taxonomicName id="68D54D25FFD8FF96FF3C4939FD98FCD9" authority="Kramer 1977" authorityName="Kramer" authorityYear="1977" box="[151,517,835,857]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="macrops">
<emphasis id="9DA1EAB4FFD8FF96FF3C4939FEE4FCD9" box="[151,377,836,857]" italics="true" pageId="8" pageNumber="9">Trimeresurus macrops</emphasis>
<bibRefCitation id="CB444B57FFD8FF96FED44939FD98FCD9" author="Kramer, E." box="[383,517,835,857]" pageId="8" pageNumber="9" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer 1977</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="CB444B57FFD8FF96FDB84939FD33FCD9" author="Kramer, E." box="[531,686,835,857]" pageId="8" pageNumber="9" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer (1977)</bibRefCitation>
: 757 (part);
<bibRefCitation id="CB444B57FFD8FF96FC834939FBCFFCD9" author="Regenass, U. &amp; Kramer, E." box="[808,1106,835,857]" pageId="8" pageNumber="9" pagination="163 - 205" refId="ref12317" refString="Regenass, U. &amp; Kramer, E. (1981) Zur Systematik der grunen Grubenottern der Gattung Trimeresurus Serpentes, Crotalidae. Revue Suisse Zoologie, 88, 163 - 205." type="journal article" year="1981">Regenass and Kramer (1981)</bibRefCitation>
: 184 (part);
<bibRefCitation id="CB444B57FFD8FF96FB66493EFA05FCD9" author="Orlov, N. &amp; Ananjeva, N. &amp; Barabanov, A. &amp; Ryabov, S. &amp; Khalikov, R." box="[1229,1432,835,858]" pageId="8" pageNumber="9" pagination="177 - 218" refId="ref12165" refString="Orlov, N., Ananjeva, N., Barabanov, A., Ryabov, S. &amp; Khalikov, R. (2002 a) Diversity of vipers Azemiopinae, Crotalinae in East, Southeast, and South Asia: Annotated checklist and natural history data. Reptilia: Squamata: Serpentes: Viperidae. Faunistische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 23, 177 - 218." type="journal article" year="2002">
Orlov
<emphasis id="9DA1EAB4FFD8FF96FAA54939FAA7FCDA" box="[1294,1338,836,858]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2002a)
</bibRefCitation>
: 193 (part);
<bibRefCitation id="CB444B57FFD8FF96FE92491FFD9AFCF8" author="Orlov, N. &amp; Ananjeva, N. &amp; Khalikov, R." box="[313,519,866,888]" pageId="8" pageNumber="9" pagination="345 - 360" refId="ref12238" refString="Orlov, N., Ananjeva, N. &amp; Khalikov, R. (2002 b) Natural History of pitvipers in Eastern and Southeastern Asia. In: Schuett, G. W., Hoggren, M., Douglas, M. E. &amp; Greene, H. W. (Eds.), Biology of the vipers. Eagle Mountain Publishing, Eagle Mountain, Utah, pp 345 - 360." type="book chapter" year="2002">
Orlov
<emphasis id="9DA1EAB4FFD8FF96FED1491EFE3AFCF8" box="[378,423,866,888]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2002b)
</bibRefCitation>
: 353 (part);
<bibRefCitation id="CB444B57FFD8FF96FD29491FFCCDFCF8" author="Stuart, B. L. &amp; Sok, K. &amp; Neang, T." box="[642,848,866,888]" pageId="8" pageNumber="9" pagination="129 - 155" refId="ref12518" refString="Stuart, B. L., Sok, K. &amp; Neang, T. (2006 a) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology, 54, 129 - 155." type="journal article" year="2006">
Stuart
<emphasis id="9DA1EAB4FFD8FF96FD6F491EFD6CFCF8" box="[708,753,866,888]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2006a)
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: 152.
</paragraph>
<paragraph id="AF6A36A6FFD8FF96FF3C49FFFCFEFC17" blockId="8.[151,1437,835,919]" box="[151,867,897,919]" pageId="8" pageNumber="9">
<taxonomicName id="68D54D25FFD8FF96FF3C49FFFDB7FC17" authority="Kramer 1977" authorityName="Kramer" authorityYear="1977" box="[151,554,897,919]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="macrops">
<emphasis id="9DA1EAB4FFD8FF96FF3C49FFFE80FC17" box="[151,285,898,919]" italics="true" pageId="8" pageNumber="9">Trimeresurus</emphasis>
cf.
<emphasis id="9DA1EAB4FFD8FF96FEEF49FFFE07FC17" box="[324,410,898,919]" italics="true" pageId="8" pageNumber="9">macrops</emphasis>
<bibRefCitation id="CB444B57FFD8FF96FE0B49FFFDB7FC17" author="Kramer, E." box="[416,554,897,919]" pageId="8" pageNumber="9" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer 1977</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="CB444B57FFD8FF96FD9F49FCFCB1FC17" author="Gumprecht, A. &amp; Tillack, F. &amp; Orlov, N. L. &amp; Captain, A. &amp; Ryabov, S." box="[564,812,897,919]" pageId="8" pageNumber="9" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. &amp; Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
Gumprecht
<emphasis id="9DA1EAB4FFD8FF96FD0749FFFD45FC17" box="[684,728,897,919]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2004)
</bibRefCitation>
: 236.
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFD8FF96FF3C49BEFEE9FA54" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="AF6A36A6FFD8FF96FF3C49BEFD67FBA4" blockId="8.[151,1437,963,2021]" pageId="8" pageNumber="9">
<emphasis id="9DA1EAB4FFD8FF96FF3C49BEFE1DFC5C" bold="true" box="[151,384,963,988]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Material examined.</emphasis>
<collectingCountry id="D7C27636FFD8FF96FE2C49B9FE60FC5C" box="[391,509,964,988]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
,
<collectingRegion id="6D11F844FFD8FF96FDA349B9FD69FC5C" box="[520,756,964,988]" country="Cambodia" name="Mondolkiri" pageId="8" pageNumber="9">Mondolkiri Province</collectingRegion>
, Seima Biodiversity Conservation Area, FMNH 262717
<emphasis id="9DA1EAB4FFD8FF96FAC049B8FAE4FC5C" box="[1387,1401,965,988]" italics="true" pageId="8" pageNumber="9"></emphasis>
21.
<collectingCountry id="D7C27636FFD8FF96FF3C499AFF64FC7F" box="[151,249,999,1023]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
, southern part of country (no precise locality), BMNH 2005.1602.
<collectingCountry id="D7C27636FFD8FF96FC52499AFBC6FC7F" box="[1017,1115,999,1023]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
,
<collectingRegion id="6D11F844FFD8FF96FBC34995FAA6FC7F" box="[1128,1339,999,1023]" country="Vietnam" name="Ho Chi Minh City" pageId="8" pageNumber="9">Ho Chi Minh City</collectingRegion>
, NMBA 12461.
<collectingCountry id="D7C27636FFD8FF96FF404E71FED0FBA4" box="[235,333,1036,1060]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
, Cu Chi and Bien Hoa, AFS98.53
<emphasis id="9DA1EAB4FFD8FF96FD624E70FD4AFBA4" box="[713,727,1037,1060]" italics="true" pageId="8" pageNumber="9"></emphasis>
55.
</paragraph>
<paragraph id="AF6A36A6FFD8FF96FF6E4E4DFDC3FB34" blockId="8.[151,1437,963,2021]" pageId="8" pageNumber="9">
<materialsCitation id="1FBD3CFBFFD8FF96FF6E4E4DFDC3FB34" collectingDate="2003-11-01" collectorName="K. Sok &amp; T. Neang." country="Cambodia" elevation="1" latitude="12.326" location="Seima Biodiversity Conservation Area" longLatPrecision="77" longitude="107.092" municipality="O'Rang District" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Mondolkiri" typeStatus="holotype">
<emphasis id="9DA1EAB4FFD8FF96FF6E4E4DFEA4FBC8" bold="true" box="[197,313,1072,1096]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">
<typeStatus id="706E8804FFD8FF96FF6E4E4DFEA9FBC8" box="[197,308,1072,1096]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
.
</emphasis>
FMNH 262718, adult female, collected at 1820 hrs on an herbaceous plant
<quantity id="682D9B43FFD8FF96FB3F4E52FB22FBC7" box="[1172,1215,1071,1095]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="1.0">
<elevation id="24F8D195FFD8FF96FB3F4E52FB22FBC7" box="[1172,1215,1071,1095]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="m" value="1.0">1 m</elevation>
</quantity>
from the
<collectorName id="02205370FFD8FF96FA844E52FF7BFBEC" pageId="8" pageNumber="9">O Kamen Stream</collectorName>
in hilly evergreen forest,
<location id="AA0A607DFFD8FF96FDA44E29FC58FBEC" LSID="urn:lsid:plazi:treatment:277C87B0FFD8FF90FF3C48AAFD51FB75:AA0A607DFFD8FF96FDA44E29FC58FBEC" box="[527,965,1108,1132]" country="Cambodia" latitude="12.326" longLatPrecision="77" longitude="107.092" municipality="O'Rang District" name="Seima Biodiversity Conservation Area" pageId="8" pageNumber="9" stateProvince="Mondolkiri">Seima Biodiversity Conservation Area</location>
(formerly part of Samling Logging Concession),
<collectingMunicipality id="4F0EACDCFFD8FF96FEAA4E0AFE2AFB0F" box="[257,439,1143,1167]" pageId="8" pageNumber="9">ORang District</collectingMunicipality>
,
<collectingRegion id="6D11F844FFD8FF96FE694E0AFD32FB0F" box="[450,687,1143,1167]" country="Cambodia" name="Mondolkiri" pageId="8" pageNumber="9">Mondolkiri Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD8FF96FD114E0AFCB0FB0F" box="[698,813,1143,1167]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
(
<geoCoordinate id="CAE15061FFD8FF96FC974E0AFC34FB0F" box="[828,937,1143,1167]" degrees="12.326" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="55" value="12.326">12.326 N</geoCoordinate>
,
<geoCoordinate id="CAE15061FFD8FF96FC1F4E0AFBACFB0F" box="[948,1073,1143,1167]" degrees="107. 092" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="55" value="107.092">107. 092 E</geoCoordinate>
),
<quantity id="682D9B43FFD8FF96FBEE4E0AFB10FB0F" box="[1093,1165,1143,1167]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="500.0">
<elevation id="24F8D195FFD8FF96FBEE4E0AFB10FB0F" box="[1093,1165,1143,1167]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="500.0">500 m</elevation>
</quantity>
elevation, on
<date id="DB6B1066FFD8FF96FA804E0AFF67FB34" pageId="8" pageNumber="9" value="2003-11-01">
<collectingDate id="CB2FE98EFFD8FF96FA804E0AFF67FB34" pageId="8" pageNumber="9" value="2003-11-01">1 November 2003</collectingDate>
</date>
by BLS,
<collectorName id="02205370FFD8FF96FECE4EE1FE28FB34" box="[357,437,1180,1204]" pageId="8" pageNumber="9">K. Sok</collectorName>
, and
<collectorName id="02205370FFD8FF96FE5B4EE1FDC3FB34" box="[496,606,1180,1204]" pageId="8" pageNumber="9">T. Neang.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="AF6A36A6FFD8FF96FF6E4EBDFC4AFAC4" blockId="8.[151,1437,963,2021]" pageId="8" pageNumber="9">
<materialsCitation id="1FBD3CFBFFD8FF96FF6E4EBDFBB3FB7C" collectingDate="2003-11-01" collectingDateMax="2003-11-06" collectingDateMin="2003-11-01" country="Cambodia" elevation="475" location="Phnom Nam Lyr Wildlife Sanctuary" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Mondolkiri" typeStatus="paratype">
<emphasis id="9DA1EAB4FFD8FF96FF6E4EBDFEDEFB57" bold="true" box="[197,323,1216,1240]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">
<typeStatus id="706E8804FFD8FF96FF6E4EBDFEA0FB58" box="[197,317,1216,1240]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
.
</emphasis>
FMNH 262717, 262719
<emphasis id="9DA1EAB4FFD8FF96FDF34EBCFDFBFB58" box="[600,614,1217,1240]" italics="true" pageId="8" pageNumber="9"></emphasis>
262721 (all males, collected from near the type locality in hilly evergreen forest between
<quantity id="682D9B43FFD8FF96FEE94E99FE5CFB7B" box="[322,449,1252,1276]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="8" pageNumber="9" unit="m" value="475.0" valueMax="500.0" valueMin="450.0">
<elevation id="24F8D195FFD8FF96FEE94E99FE5CFB7B" box="[322,449,1252,1276]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="8" pageNumber="9" unit="m" value="475.0" valueMax="500.0" valueMin="450.0">
450
<emphasis id="9DA1EAB4FFD8FF96FEC74E98FEE7FB7C" box="[364,378,1253,1276]" italics="true" pageId="8" pageNumber="9"></emphasis>
500 m
</elevation>
</quantity>
elevation, near streams between
<date id="DB6B1066FFD8FF96FC9D4E99FBBCFB7C" box="[822,1057,1252,1276]" pageId="8" pageNumber="9" value="2003-11-01" valueMax="2003-11-06" valueMin="2003-11-01">
<collectingDate id="CB2FE98EFFD8FF96FC9D4E99FBBCFB7C" box="[822,1057,1252,1276]" pageId="8" pageNumber="9" value="2003-11-01" valueMax="2003-11-06" valueMin="2003-11-01">
1
<emphasis id="9DA1EAB4FFD8FF96FCEF4E98FCCFFB7C" box="[836,850,1253,1276]" italics="true" pageId="8" pageNumber="9"></emphasis>
6 November 2003
</collectingDate>
</date>
);
</materialsCitation>
<materialsCitation id="1FBD3CFBFFD8FF96FB9E4E99FCEEFAA0" collectingDate="2003-11-01" collectingDateMax="2003-11-06" collectingDateMin="2003-11-01" country="Cambodia" elevation="475" location="Phnom Nam Lyr Wildlife Sanctuary" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Mondolkiri" typeStatus="paratype">
FMNH 259190 (female,
<location id="AA0A607DFFD8FF96FAE74E99FE7EFAA0" LSID="urn:lsid:plazi:treatment:277C87B0FFD8FF90FF3C48AAFD51FB75:AA0A607DFFD8FF96FAE74E99FE7EFAA0" country="Cambodia" name="Phnom Nam Lyr Wildlife Sanctuary" pageId="8" pageNumber="9" stateProvince="Mondolkiri">Phnom Nam Lyr Wildlife Sanctuary</location>
,
<collectingRegion id="6D11F844FFD8FF96FE584F75FD7EFAA0" box="[499,739,1288,1312]" country="Cambodia" name="Mondolkiri" pageId="8" pageNumber="9">Mondolkiri Province</collectingRegion>
,
<collectingCountry id="D7C27636FFD8FF96FD584F75FCF1FAA0" box="[755,876,1288,1312]" name="Cambodia" pageId="8" pageNumber="9">Cambodia</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFD8FF96FC294F75FE75FAC4" pageId="8" pageNumber="9">
BMNH 2005.1602 (female, from an unknown location in southern
<collectingCountry id="D7C27636FFD8FF96FED74F51FE7DFAC4" box="[380,480,1324,1348]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFD8FF96FE584F51FC49FAC4" box="[499,980,1324,1348]" country="Vietnam" location="Saigon" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Ho Chi Minh City" typeStatus="paratype">
NMBA 12461 (
<collectingRegion id="6D11F844FFD8FF96FD084F51FD68FAC4" box="[675,757,1324,1348]" country="Vietnam" name="Ho Chi Minh City" pageId="8" pageNumber="9">Saigon</collectingRegion>
, southern
<collectingCountry id="D7C27636FFD8FF96FCCC4F51FC56FAC4" box="[871,971,1324,1348]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="AF6A36A6FFD8FF96FF6E4F32FEE9FA54" blockId="8.[151,1437,963,2021]" pageId="8" pageNumber="9">
<emphasis id="9DA1EAB4FFD8FF96FF6E4F32FE24FAE7" bold="true" box="[197,441,1359,1384]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Additional material.</emphasis>
<materialsCitation id="1FBD3CFBFFD8FF96FE694F2DFEFCFA0C" country="Vietnam" location="Ho Chi Minh City" pageId="8" pageNumber="9" specimenCount="1" stateProvince="Ho Chi Minh City">
Data for
<emphasis id="9DA1EAB4FFD8FF96FD834F2CFDAEFAE8" box="[552,563,1361,1384]" italics="true" pageId="8" pageNumber="9">s</emphasis>
everal specimens obtained from dealers in the vicinity of
<collectingRegion id="6D11F844FFD8FF96FB6E4F2DFA0AFAE8" box="[1221,1431,1360,1384]" country="Vietnam" name="Ho Chi Minh City" pageId="8" pageNumber="9">Ho Chi Minh City</collectingRegion>
, southern
<collectingCountry id="D7C27636FFD8FF96FEA94F09FEFCFA0C" box="[258,353,1396,1420]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
</materialsCitation>
are included in all analyses.
<materialsCitation id="1FBD3CFBFFD8FF96FD174F09FEAAFA30" country="Vietnam" county="Cho - Ray Hospital" latitude="10.95" location="These" longLatPrecision="778" longitude="106.82" municipality="Kiem" pageId="8" pageNumber="9" specimenCount="1">
The specimens were stated to originate from Cu Chi (
<geoCoordinate id="CAE15061FFD8FF96FA9D4F09FA05FA0B" box="[1334,1432,1396,1420]" degrees="10.75" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="10.75">10.75 N</geoCoordinate>
,
<geoCoordinate id="CAE15061FFD8FF96FF3C4FE5FF63FA30" box="[151,254,1432,1456]" degrees="106.67" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="106.67">106.67 E</geoCoordinate>
) and
</materialsCitation>
<materialsCitation id="1FBD3CFBFFD8FF96FEEB4FE5FEEDFA54" country="Vietnam" county="Cho - Ray Hospital" latitude="10.95" location="These" longLatPrecision="778" longitude="106.82" municipality="Kiem" pageId="8" pageNumber="9" specimenCount="1">
Bien Hoa (
<geoCoordinate id="CAE15061FFD8FF96FE164FE5FD80FA2F" box="[445,541,1432,1456]" degrees="10.95" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="555" value="10.95">10.95 N</geoCoordinate>
,
<geoCoordinate id="CAE15061FFD8FF96FD814FE5FD09FA30" box="[554,660,1432,1456]" degrees="106.82" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="106.82">106.82 E</geoCoordinate>
).
<location id="AA0A607DFFD8FF96FD014FE5FD70FA30" LSID="urn:lsid:plazi:treatment:277C87B0FFD8FF90FF3C48AAFD51FB75:AA0A607DFFD8FF96FD014FE5FD70FA30" box="[682,749,1432,1456]" country="Vietnam" county="Cho - Ray Hospital" latitude="10.95" longLatPrecision="778" longitude="106.82" municipality="Kiem" name="These" pageId="8" pageNumber="9">These</location>
specimens were left in the care of
<location id="AA0A607DFFD8FF96FB2B4FE5FB03FA2F" LSID="urn:lsid:plazi:treatment:277C87B0FFD8FF90FF3C48AAFD51FB75:AA0A607DFFD8FF96FB2B4FE5FB03FA2F" box="[1152,1182,1432,1455]" country="Vietnam" county="Cho - Ray Hospital" latitude="10.95" longLatPrecision="778" longitude="106.82" municipality="Kiem" name="Dr" pageId="8" pageNumber="9">Dr</location>
T-X.
<collectingMunicipality id="4F0EACDCFFD8FF96FB494FE5FABDFA30" box="[1250,1312,1432,1456]" pageId="8" pageNumber="9">Kiem</collectingMunicipality>
(
<collectingCounty id="460B4E2AFFD8FF96FA9B4FE5FF64FA54" pageId="8" pageNumber="9">
Cho
<emphasis id="9DA1EAB4FFD8FF96FACA4FE4FAF2FA30" box="[1377,1391,1433,1456]" italics="true" pageId="8" pageNumber="9"></emphasis>
Ray Hospital
</collectingCounty>
,
<collectingCountry id="D7C27636FFD8FF96FEAF4FC1FEF5FA54" box="[260,360,1468,1492]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFD8FF97FF6E4FA2FD6AF86E" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="AF6A36A6FFD8FF97FF6E4FA2FF61FEBC" blockId="8.[151,1437,963,2021]" lastBlockId="9.[151,1436,151,316]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="9DA1EAB4FFD8FF96FF6E4FA2FEA2FA77" bold="true" box="[197,319,1503,1528]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Diagnosis.</emphasis>
<taxonomicName id="68D54D25FFD8FF96FEEC4F9DFDA9FA78" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[327,564,1504,1528]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD8FF96FEEC4F9DFDA9FA78" box="[327,564,1504,1528]" italics="true" pageId="8" pageNumber="9">Cryptelytrops rubeus</emphasis>
</taxonomicName>
is distinguished from all other Asian pitviper species which have the typical &quot;green pitviper&quot; colouration (uniform green dorsal colour and a lateral stripe present on the first few dorsal scale rows in one or both sexes), except other species of
<taxonomicName id="68D54D25FFD8FF96FD4B4C5BFCE4F9BE" authorityName="Cope" authorityYear="1860" box="[736,889,1574,1598]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="9DA1EAB4FFD8FF96FD4B4C5BFCE4F9BE" box="[736,889,1574,1598]" italics="true" pageId="8" pageNumber="9">Cryptelytrops</emphasis>
</taxonomicName>
, by the presence of a fused first supralabial and nasal scale. It can be distinguished from
<emphasis id="9DA1EAB4FFD8FF96FDC34C34FC0CF9E1" box="[616,913,1609,1633]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="68D54D25FFD8FF96FDC34C34FC9DF9E1" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[616,768,1609,1633]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
,
<taxonomicName id="68D54D25FFD8FF96FCA64C34FC0CF9E1" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[781,913,1609,1633]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
</emphasis>
and
<taxonomicName id="68D54D25FFD8FF96FC604C34FB0FF9E1" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[971,1170,1609,1633]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="9DA1EAB4FFD8FF96FC604C34FB0FF9E1" box="[971,1170,1609,1633]" italics="true" pageId="8" pageNumber="9">C. septentrionalis</emphasis>
</taxonomicName>
primarily by the larger size of the eye (most obvious in adults), the wider supraoculars, and the shape of the head. The latter is more elongate
<emphasis id="9DA1EAB4FFD8FF96FF6E4CECFF4EF928" box="[197,211,1681,1704]" italics="true" pageId="8" pageNumber="9"></emphasis>
oval in
<emphasis id="9DA1EAB4FFD8FF96FE8C4CEDFDD1F928" box="[295,588,1680,1704]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="68D54D25FFD8FF96FE8C4CEDFE23F928" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[295,446,1680,1704]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
,
<taxonomicName id="68D54D25FFD8FF96FE624CEDFDD1F928" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[457,588,1680,1704]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
</emphasis>
and
<emphasis id="9DA1EAB4FFD8FF96FD294CEDFCCDF928" box="[642,848,1680,1704]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="68D54D25FFD8FF96FD294CEDFCD1F928" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[642,844,1680,1704]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="septentrionalis">C. septentrionalis</taxonomicName>
,
</emphasis>
but widens quite abruptly behind the eyes in
<emphasis id="9DA1EAB4FFD8FF96FAE64CEDFE6FF94B" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="68D54D25FFD8FF96FAE64CEDFEB9F94B" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cardamomensis">C. cardamomensis</taxonomicName>
,
<taxonomicName id="68D54D25FFD8FF96FE9C4CCEFE22F94B" authorityName="Kramer" authorityYear="1977" box="[311,447,1715,1739]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD8FF96FE644CC9FE6FF94B" box="[463,498,1716,1739]" pageId="8" pageNumber="9" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="68D54D25FFD8FF96FD924CCEFD33F94B" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[569,686,1715,1739]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD8FF96FD924CCEFD33F94B" box="[569,686,1715,1739]" italics="true" pageId="8" pageNumber="9">C. rubeus</emphasis>
</taxonomicName>
to give the latter a characteristically triangular shaped head.
<taxonomicName id="68D54D25FFD8FF96FF3C4CABFE18F96E" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[151,389,1750,1774]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD8FF96FF3C4CABFE18F96E" box="[151,389,1750,1774]" italics="true" pageId="8" pageNumber="9">Cryptelytrops rubeus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="68D54D25FFD8FF96FD154CABFC12F96E" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[702,911,1750,1774]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
<emphasis id="9DA1EAB4FFD8FF96FD154CABFC12F96E" box="[702,911,1750,1774]" italics="true" pageId="8" pageNumber="9">C. cardamomensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD8FF96FC334CAAFC7BF96E" box="[920,998,1751,1774]" pageId="8" pageNumber="9" rank="species">sp.nov.</taxonomicNameLabel>
(described above) using the combination of the following characters (further details are given in
<tableCitation id="E257031DFFD8FF96FC9B4C84FC1BF891" box="[816,902,1785,1809]" captionStart="TABLE 2" captionStartId="9.[151,234,358,381]" captionTargetPageId="9" captionText="TABLE 2. Mean values and standard deviations of morphological characters that in combination can be used to distinguish C. cardamomensis sp. nov. from both C. macrops s.s. and C. rubeus sp. nov. Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm = C. macrops s.s., Cc = C. cardamomensis, Cr = C. rubeus. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from C. cardamomensis (e.g. BTWSUPOC can be used to discriminate both C. macrops and C. rubeus from C. cardamomensis while DV19to17 is only useful to discriminate C. rubeus from C. cardamomensis)." pageId="8" pageNumber="9">Table 2</tableCitation>
). In males,
<taxonomicName id="68D54D25FFD8FF96FBAD4C84FBEEF891" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[1030,1139,1785,1809]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD8FF96FBAD4C84FBEEF891" box="[1030,1139,1785,1809]" italics="true" pageId="8" pageNumber="9">C. rubeus</emphasis>
</taxonomicName>
tends to have fewer scales between the rear edges of the supraoculars (BTWSUPOC2), a relatively smaller eye (DEYE), and a less prominent postocular white stripe (SCROC). The scale reduction from 19 to 17 dorsal scale rows (DV19TO17) tends to involve lower scale rows, it tends to have less keeled body scales at mid-body (BSCK), and fewer scales between the last supralabial and the chin shields (VENTEDGE). In females, the scale reduction from 19 to 17 dorsal scale rows (DV19TO17) tends to involve higher scale rows in
<taxonomicName id="68D54D25FFD8FF96FC8E4DD7FC09F842" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[805,916,1962,1986]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD8FF96FC8E4DD7FC09F842" box="[805,916,1962,1986]" italics="true" pageId="8" pageNumber="9">C. rubeus</emphasis>
</taxonomicName>
than in
<taxonomicName id="68D54D25FFD8FF96FC5E4DD7FB5BF842" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[1013,1222,1962,1986]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD8FF96FC5E4DD7FB5BF842" box="[1013,1222,1962,1986]" italics="true" pageId="8" pageNumber="9">C. cardamomensis</emphasis>
</taxonomicName>
, it has a relatively shorter head (LHEAD), a small scale is never present between the nasal scale and the scale bordering the anterior edge of the pit (NASPIT), the lateral white stripe covers a larger proportion of the first dorsal scale row (SCR1), it tends to have fewer (larger) scales bordering the supralabial scales (BORSUPOC), and less keeled body scales at mid-body (BSCK). The scale reduction from 12 to 10 scale rows around the tail (SC12O10) also tends to occur closer to the vent in females of
<taxonomicName id="68D54D25FFD9FF97FDBD4B7CFD76FE99" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[534,747,257,281]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD9FF97FDBD4B7CFD76FE99" box="[534,747,257,281]" italics="true" pageId="9" pageNumber="10">C. cardamomensis</emphasis>
</taxonomicName>
than
<taxonomicName id="68D54D25FFD9FF97FC994B7CFC3EFE99" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[818,931,257,281]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD9FF97FC994B7CFC3EFE99" box="[818,931,257,281]" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
and they also have relatively smaller eyes (DEYE).
</paragraph>
<caption id="FBAA662EFFD9FF97FF3C4B1BFE7EFDC9" pageId="9" pageNumber="10" startId="9.[151,234,358,381]" targetBox="[156,1275,613,1627]" targetIsTable="true" targetPageId="9">
<paragraph id="AF6A36A6FFD9FF97FF3C4B1BFE7EFDC9" blockId="9.[151,1436,358,586]" pageId="9" pageNumber="10">
<emphasis id="9DA1EAB4FFD9FF97FF3C4B1BFE99FEFD" bold="true" box="[151,260,358,381]" pageId="9" pageNumber="10">TABLE 2.</emphasis>
Mean values and standard deviations of morphological characters that in combination can be used to distinguish
<taxonomicName id="68D54D25FFD9FF97FA2F4B1AFEA8FE1A" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
<emphasis id="9DA1EAB4FFD9FF97FA2F4B1AFEA8FE1A" italics="true" pageId="9" pageNumber="10">C. cardamomensis</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFD9FF97FE974BF9FE13FE19" bold="true" box="[316,398,388,409]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="869257CFFFD9FF97FE974BF9FE13FE19" box="[316,398,388,409]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from both
<emphasis id="9DA1EAB4FFD9FF97FDA84BF9FD02FE19" box="[515,671,388,410]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="68D54D25FFD9FF97FDA84BF9FDEAFE19" authorityName="Kramer" authorityYear="1977" box="[515,631,388,410]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD9FF97FDD44BF9FD02FE19" box="[639,671,388,409]" pageId="9" pageNumber="10" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="68D54D25FFD9FF97FD794BF9FCABFE1A" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[722,822,388,410]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
<emphasis id="9DA1EAB4FFD9FF97FD794BF9FCABFE1A" box="[722,822,388,410]" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFD9FF97FC964BF9FC12FE19" bold="true" box="[829,911,388,409]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="869257CFFFD9FF97FC964BF9FC12FE19" box="[829,911,388,409]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm =
<emphasis id="9DA1EAB4FFD9FF97FEB44BA1FE21FE71" box="[287,444,476,498]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="68D54D25FFD9FF97FEB44BA1FE09FE71" authorityName="Kramer" authorityYear="1977" box="[287,404,476,498]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD9FF97FE374BA1FE21FE71" box="[412,444,476,497]" pageId="9" pageNumber="10" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
, Cc =
<taxonomicName id="68D54D25FFD9FF97FDAF4BA1FD74FE71" authority=", Cr" authorityName="Cr" box="[516,745,475,498]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD9FF97FDAF4BA1FD5AFE72" box="[516,711,476,498]" italics="true" pageId="9" pageNumber="10">C. cardamomensis,</emphasis>
Cr
</taxonomicName>
=
<taxonomicName id="68D54D25FFD9FF97FCAE4BA1FCF4FE72" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[773,873,476,498]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD9FF97FCAE4BA1FCF4FE72" box="[773,873,476,498]" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from
<taxonomicName id="68D54D25FFD9FF97FC7B4B84FB11FD8F" box="[976,1164,505,527]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD9FF97FC7B4B84FB11FD8F" box="[976,1164,505,527]" italics="true" pageId="9" pageNumber="10">C. cardamomensis</emphasis>
</taxonomicName>
(e.g. BTWSUPOC can be used to discriminate both
<taxonomicName id="68D54D25FFD9FF97FE36486AFD8CFDAC" authorityName="Kramer" authorityYear="1977" box="[413,529,535,557]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="macrops">
<emphasis id="9DA1EAB4FFD9FF97FE36486AFD8CFDAC" box="[413,529,535,557]" italics="true" pageId="9" pageNumber="10">C. macrops</emphasis>
</taxonomicName>
and
<taxonomicName id="68D54D25FFD9FF97FDE8486AFD3BFDAD" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[579,678,535,557]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD9FF97FDE8486AFD3BFDAD" box="[579,678,535,557]" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
from
<taxonomicName id="68D54D25FFD9FF97FD4F486AFC3DFDAD" box="[740,928,535,557]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD9FF97FD4F486AFC3DFDAD" box="[740,928,535,557]" italics="true" pageId="9" pageNumber="10">C. cardamomensis</emphasis>
</taxonomicName>
while DV19to17 is only useful to discriminate
<taxonomicName id="68D54D25FFD9FF97FA2F486AFF41FDCA" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD9FF97FA2F486AFF41FDCA" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
from
<taxonomicName id="68D54D25FFD9FF97FEB14849FE48FDCA" box="[282,469,564,586]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="cardamomensis">
<emphasis id="9DA1EAB4FFD9FF97FEB14849FE48FDCA" box="[282,469,564,586]" italics="true" pageId="9" pageNumber="10">C. cardamomensis</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="AF6A36A6FFD9FF97FF374818FB7CF9DB" pageId="9" pageNumber="10">
<table id="DDD5C406FFD90061FF374818FB66F9DB" box="[156,1275,613,1627]" gridcols="4" gridrows="25" pageId="9" pageNumber="10">
<tr id="11E534E4FFD90061FF374818FB66FDFB" box="[156,1275,613,635]" gridrow="0" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374818FEAEFDFB" box="[156,307,613,635]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">Character</th>
<th id="52345D98FFD90061FE744818FDE8FDFB" box="[479,629,613,635]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">Cm</th>
<th id="52345D98FFD90061FC8A4818FC24FDFB" box="[801,953,613,635]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">Cc</th>
<th id="52345D98FFD90061FBCF4818FB66FDFB" box="[1124,1275,613,635]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">Cr</th>
</tr>
<tr id="11E534E4FFD90061FF3748F3FB66FD24" box="[156,1275,654,676]" gridrow="1" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF3748F3FB66FD24" box="[156,1275,654,676]" colspan="4" colspanRight="3" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Males</th>
</tr>
<tr id="11E534E4FFD90061FF3748C5FB66FD4E" box="[156,1275,696,718]" gridrow="2" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF3748C5FEAEFD4E" box="[156,307,696,718]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">BTWSUPOC2</th>
<td id="52345D98FFD90061FE7448C5FDE8FD4E" box="[479,629,696,718]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">11.96 (±1.22)</td>
<td id="52345D98FFD90061FC8A48C5FC24FD4E" box="[801,953,696,718]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">14.50 (±2.12)</td>
<td id="52345D98FFD90061FBCF48C5FB66FD4E" box="[1124,1275,696,718]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">10.17 (±1.94)</td>
</tr>
<tr id="11E534E4FFD90061FF37489CFB66FD77" box="[156,1275,737,759]" gridrow="3" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF37489CFEAEFD77" box="[156,307,737,759]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">DV19TO17</th>
<td id="52345D98FFD90061FE74489CFDE8FD77" box="[479,629,737,759]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">4.29 (±0.43)</td>
<td id="52345D98FFD90061FC8A489CFC24FD77" box="[801,953,737,759]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">4.25 (±1.06)</td>
<td id="52345D98FFD90061FBCF489CFB66FD77" box="[1124,1275,737,759]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">3.92 (±0.38)</td>
</tr>
<tr id="11E534E4FFD90061FF374977FB66FCA0" box="[156,1275,778,800]" gridrow="4" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374977FEAEFCA0" box="[156,307,778,800]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">BSCK</th>
<td id="52345D98FFD90061FE744977FDE8FCA0" box="[479,629,778,800]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">0.80 (±0.25)</td>
<td id="52345D98FFD90061FC8A4977FC24FCA0" box="[801,953,778,800]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">1.00 (±0.00)</td>
<td id="52345D98FFD90061FBCF4977FB66FCA0" box="[1124,1275,778,800]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">0.38 (±0.21)</td>
</tr>
<tr id="11E534E4FFD90061FF37494EFB66FCC9" box="[156,1275,819,841]" gridrow="5" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF37494EFEAEFCC9" box="[156,307,819,841]" gridcol="0" gridrow="5" pageId="9" pageNumber="10">VENTEDGE</th>
<td id="52345D98FFD90061FE74494EFDE8FCC9" box="[479,629,819,841]" gridcol="1" gridrow="5" pageId="9" pageNumber="10">7.46 (±0.45)</td>
<td id="52345D98FFD90061FC8A494EFC24FCC9" box="[801,953,819,841]" gridcol="2" gridrow="5" pageId="9" pageNumber="10">7.50 (±0.71)</td>
<td id="52345D98FFD90061FBCF494EFB66FCC9" box="[1124,1275,819,841]" gridcol="3" gridrow="5" pageId="9" pageNumber="10">6.92 (±0.49)</td>
</tr>
<tr id="11E534E4FFD90061FF374920FB66FCF3" box="[156,1275,861,883]" gridrow="6" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374920FEAEFCF3" box="[156,307,861,883]" gridcol="0" gridrow="6" pageId="9" pageNumber="10">DEYE</th>
<td id="52345D98FFD90061FE744920FDE8FCF3" box="[479,629,861,883]" gridcol="1" gridrow="6" pageId="9" pageNumber="10">4.03 (±0.36)</td>
<td id="52345D98FFD90061FC8A4920FC24FCF3" box="[801,953,861,883]" gridcol="2" gridrow="6" pageId="9" pageNumber="10">4.46 (±0.13)</td>
<td id="52345D98FFD90061FBCF4920FB66FCF3" box="[1124,1275,861,883]" gridcol="3" gridrow="6" pageId="9" pageNumber="10">4.08 (± 0.19)</td>
</tr>
<tr id="11E534E4FFD90061FF3749FBFB66FC1C" box="[156,1275,902,924]" gridrow="7" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF3749FBFEAEFC1C" box="[156,307,902,924]" gridcol="0" gridrow="7" pageId="9" pageNumber="10">SCROC</th>
<td id="52345D98FFD90061FE7449FBFDE8FC1C" box="[479,629,902,924]" gridcol="1" gridrow="7" pageId="9" pageNumber="10">0.69 (±0.93)</td>
<td id="52345D98FFD90061FC8A49FBFC24FC1C" box="[801,953,902,924]" gridcol="2" gridrow="7" pageId="9" pageNumber="10">2.00 (±0.00)</td>
<td id="52345D98FFD90061FBCF49FBFB66FC1C" box="[1124,1275,902,924]" gridcol="3" gridrow="7" pageId="9" pageNumber="10">0.33 (±0.82)</td>
</tr>
<tr id="11E534E4FFD90061FF3749CDFB66FC46" box="[156,1275,944,966]" gridrow="8" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF3749CDFEAEFC46" box="[156,307,944,966]" gridcol="0" gridrow="8" pageId="9" pageNumber="10">BORSUPOC</th>
<td id="52345D98FFD90061FE7449CDFDE8FC46" box="[479,629,944,966]" gridcol="1" gridrow="8" pageId="9" pageNumber="10">9.65 (±0.90)</td>
<td id="52345D98FFD90061FC8A49CDFC24FC46" box="[801,953,944,966]" gridcol="2" gridrow="8" pageId="9" pageNumber="10">8.75 (±0.35)</td>
<td id="52345D98FFD90061FBCF49CDFB66FC46" box="[1124,1275,944,966]" gridcol="3" gridrow="8" pageId="9" pageNumber="10">8.58 (± 0.97)</td>
</tr>
<tr id="11E534E4FFD90061FF3749A4FB66FC6F" box="[156,1275,985,1007]" gridrow="9" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF3749A4FEAEFC6F" box="[156,307,985,1007]" gridcol="0" gridrow="9" pageId="9" pageNumber="10">VS19TO17</th>
<td id="52345D98FFD90061FE7449A4FDE8FC6F" box="[479,629,985,1007]" gridcol="1" gridrow="9" pageId="9" pageNumber="10">68.00 (±1.99)</td>
<td id="52345D98FFD90061FC8A49A4FC24FC6F" box="[801,953,985,1007]" gridcol="2" gridrow="9" pageId="9" pageNumber="10">69.29 (±2.16)</td>
<td id="52345D98FFD90061FBCF49A4FB66FC6F" box="[1124,1275,985,1007]" gridcol="3" gridrow="9" pageId="9" pageNumber="10">69.29 (±1.53)</td>
</tr>
<tr id="11E534E4FFD90061FF374E7FFB66FB98" box="[156,1275,1026,1048]" gridrow="10" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374E7FFEAEFB98" box="[156,307,1026,1048]" gridcol="0" gridrow="10" pageId="9" pageNumber="10">KTEMP</th>
<td id="52345D98FFD90061FE744E7FFDE8FB98" box="[479,629,1026,1048]" gridcol="1" gridrow="10" pageId="9" pageNumber="10">0.67 (±0.31)</td>
<td id="52345D98FFD90061FC8A4E7FFC24FB98" box="[801,953,1026,1048]" gridcol="2" gridrow="10" pageId="9" pageNumber="10">0.50 (±0.00)</td>
<td id="52345D98FFD90061FBCF4E7FFB66FB98" box="[1124,1275,1026,1048]" gridcol="3" gridrow="10" pageId="9" pageNumber="10">0.46 (±0.29)</td>
</tr>
<tr id="11E534E4FFD90061FF374E56FB66FBC1" box="[156,1275,1067,1089]" gridrow="11" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374E56FB66FBC1" box="[156,1275,1067,1089]" colspan="4" colspanRight="3" gridcol="0" gridrow="11" pageId="9" pageNumber="10">b. Females</th>
</tr>
<tr id="11E534E4FFD90061FF374E28FB66FBEB" box="[156,1275,1109,1131]" gridrow="12" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374E28FEAEFBEB" box="[156,307,1109,1131]" gridcol="0" gridrow="12" pageId="9" pageNumber="10">STRIPE</th>
<td id="52345D98FFD90061FE744E28FDE8FBEB" box="[479,629,1109,1131]" gridcol="1" gridrow="12" pageId="9" pageNumber="10">0.70 (±0.57)</td>
<td id="52345D98FFD90061FC8A4E28FC24FBEB" box="[801,953,1109,1131]" gridcol="2" gridrow="12" pageId="9" pageNumber="10">2.00 (±0.00)</td>
<td id="52345D98FFD90061FBCF4E28FB66FBEB" box="[1124,1275,1109,1131]" gridcol="3" gridrow="12" pageId="9" pageNumber="10">1.00 (±0.71)</td>
</tr>
<tr id="11E534E4FFD90061FF374E03FB66FB14" box="[156,1275,1150,1172]" gridrow="13" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374E03FEAEFB14" box="[156,307,1150,1172]" gridcol="0" gridrow="13" pageId="9" pageNumber="10">SCR1</th>
<td id="52345D98FFD90061FE744E03FDE8FB14" box="[479,629,1150,1172]" gridcol="1" gridrow="13" pageId="9" pageNumber="10">0.15 (±0.13)</td>
<td id="52345D98FFD90061FC8A4E03FC24FB14" box="[801,953,1150,1172]" gridcol="2" gridrow="13" pageId="9" pageNumber="10">0.25 (±0.07)</td>
<td id="52345D98FFD90061FBCF4E03FB66FB14" box="[1124,1275,1150,1172]" gridcol="3" gridrow="13" pageId="9" pageNumber="10">0.29 (±0.22)</td>
</tr>
<tr id="11E534E4FFD90061FF374ED5FB66FB3E" box="[156,1275,1192,1214]" gridrow="14" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374ED5FEAEFB3E" box="[156,307,1192,1214]" gridcol="0" gridrow="14" pageId="9" pageNumber="10">SC12TO10</th>
<td id="52345D98FFD90061FE744ED5FDE8FB3E" box="[479,629,1192,1214]" gridcol="1" gridrow="14" pageId="9" pageNumber="10">5.81 (±1.62)</td>
<td id="52345D98FFD90061FC8A4ED5FC24FB3E" box="[801,953,1192,1214]" gridcol="2" gridrow="14" pageId="9" pageNumber="10">5.37 (±0.28)</td>
<td id="52345D98FFD90061FBCF4ED5FB66FB3E" box="[1124,1275,1192,1214]" gridcol="3" gridrow="14" pageId="9" pageNumber="10">4.65 (±1.71)</td>
</tr>
<tr id="11E534E4FFD90061FF374EACFB66FB67" box="[156,1275,1233,1255]" gridrow="15" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374EACFEAEFB67" box="[156,307,1233,1255]" gridcol="0" gridrow="15" pageId="9" pageNumber="10">SUBLAB</th>
<td id="52345D98FFD90061FE744EACFDE8FB67" box="[479,629,1233,1255]" gridcol="1" gridrow="15" pageId="9" pageNumber="10">11.75 (±0.60)</td>
<td id="52345D98FFD90061FC8A4EACFC24FB67" box="[801,953,1233,1255]" gridcol="2" gridrow="15" pageId="9" pageNumber="10">13.00 (±0.71)</td>
<td id="52345D98FFD90061FBCF4EACFB66FB67" box="[1124,1275,1233,1255]" gridcol="3" gridrow="15" pageId="9" pageNumber="10">12.00 (±1.28)</td>
</tr>
<tr id="11E534E4FFD90061FF374E87FB66FA90" box="[156,1275,1274,1296]" gridrow="16" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374E87FEAEFA90" box="[156,307,1274,1296]" gridcol="0" gridrow="16" pageId="9" pageNumber="10">DV19TO17</th>
<td id="52345D98FFD90061FE744E87FDE8FA90" box="[479,629,1274,1296]" gridcol="1" gridrow="16" pageId="9" pageNumber="10">4.38 (±0.58)</td>
<td id="52345D98FFD90061FC8A4E87FC24FA90" box="[801,953,1274,1296]" gridcol="2" gridrow="16" pageId="9" pageNumber="10">3.50 (±0.00)</td>
<td id="52345D98FFD90061FBCF4E87FB66FA90" box="[1124,1275,1274,1296]" gridcol="3" gridrow="16" pageId="9" pageNumber="10">4.20 (±0.27)</td>
</tr>
<tr id="11E534E4FFD90061FF374F5EFB66FAB9" box="[156,1275,1315,1337]" gridrow="17" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374F5EFEAEFAB9" box="[156,307,1315,1337]" gridcol="0" gridrow="17" pageId="9" pageNumber="10">VS19TO17</th>
<td id="52345D98FFD90061FE744F5EFDE8FAB9" box="[479,629,1315,1337]" gridcol="1" gridrow="17" pageId="9" pageNumber="10">67.64 (±2.99)</td>
<td id="52345D98FFD90061FC8A4F5EFC24FAB9" box="[801,953,1315,1337]" gridcol="2" gridrow="17" pageId="9" pageNumber="10">68.38 (±1.30)</td>
<td id="52345D98FFD90061FBCF4F5EFB66FAB9" box="[1124,1275,1315,1337]" gridcol="3" gridrow="17" pageId="9" pageNumber="10">69.26 (±2.74)</td>
</tr>
<tr id="11E534E4FFD90061FF374F30FB66FAE3" box="[156,1275,1357,1379]" gridrow="18" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374F30FEAEFAE3" box="[156,307,1357,1379]" gridcol="0" gridrow="18" pageId="9" pageNumber="10">NASPIT</th>
<td id="52345D98FFD90061FE744F30FDE8FAE3" box="[479,629,1357,1379]" gridcol="1" gridrow="18" pageId="9" pageNumber="10">0.13 (±0.39)</td>
<td id="52345D98FFD90061FC8A4F30FC24FAE3" box="[801,953,1357,1379]" gridcol="2" gridrow="18" pageId="9" pageNumber="10">1.00 (±0.00)</td>
<td id="52345D98FFD90061FBCF4F30FB66FAE3" box="[1124,1275,1357,1379]" gridcol="3" gridrow="18" pageId="9" pageNumber="10">0.00 (±0.00)</td>
</tr>
<tr id="11E534E4FFD90061FF374F0BFB66FA0C" box="[156,1275,1398,1420]" gridrow="19" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374F0BFEAEFA0C" box="[156,307,1398,1420]" gridcol="0" gridrow="19" pageId="9" pageNumber="10">BORSUPOC</th>
<td id="52345D98FFD90061FE744F0BFDE8FA0C" box="[479,629,1398,1420]" gridcol="1" gridrow="19" pageId="9" pageNumber="10">9.70 (±1.19)</td>
<td id="52345D98FFD90061FC8A4F0BFC24FA0C" box="[801,953,1398,1420]" gridcol="2" gridrow="19" pageId="9" pageNumber="10">9.00 (±0.71)</td>
<td id="52345D98FFD90061FBCF4F0BFB66FA0C" box="[1124,1275,1398,1420]" gridcol="3" gridrow="19" pageId="9" pageNumber="10">8.50 (±0.35)</td>
</tr>
<tr id="11E534E4FFD90061FF374FDDFB66FA36" box="[156,1275,1440,1462]" gridrow="20" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374FDDFEAEFA36" box="[156,307,1440,1462]" gridcol="0" gridrow="20" pageId="9" pageNumber="10">BSCK</th>
<td id="52345D98FFD90061FE744FDDFDE8FA36" box="[479,629,1440,1462]" gridcol="1" gridrow="20" pageId="9" pageNumber="10">0.75 (±0.30)</td>
<td id="52345D98FFD90061FC8A4FDDFC24FA36" box="[801,953,1440,1462]" gridcol="2" gridrow="20" pageId="9" pageNumber="10">0.75 (±0.35)</td>
<td id="52345D98FFD90061FBCF4FDDFB66FA36" box="[1124,1275,1440,1462]" gridcol="3" gridrow="20" pageId="9" pageNumber="10">0.50 (±0.35)</td>
</tr>
<tr id="11E534E4FFD90061FF374FB4FB66FA5F" box="[156,1275,1481,1503]" gridrow="21" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374FB4FEAEFA5F" box="[156,307,1481,1503]" gridcol="0" gridrow="21" pageId="9" pageNumber="10">BTWSUPOC2</th>
<td id="52345D98FFD90061FE744FB4FDE8FA5F" box="[479,629,1481,1503]" gridcol="1" gridrow="21" pageId="9" pageNumber="10">12.75 (±1.33)</td>
<td id="52345D98FFD90061FC8A4FB4FC24FA5F" box="[801,953,1481,1503]" gridcol="2" gridrow="21" pageId="9" pageNumber="10">15.00 (±1.41)</td>
<td id="52345D98FFD90061FBCF4FB4FB66FA5F" box="[1124,1275,1481,1503]" gridcol="3" gridrow="21" pageId="9" pageNumber="10">13.40 (±3.78)</td>
</tr>
<tr id="11E534E4FFD90061FF374F8FFB66F988" box="[156,1275,1522,1544]" gridrow="22" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374F8FFEAEF988" box="[156,307,1522,1544]" gridcol="0" gridrow="22" pageId="9" pageNumber="10">VSC</th>
<td id="52345D98FFD90061FE744F8FFDE8F988" box="[479,629,1522,1544]" gridcol="1" gridrow="22" pageId="9" pageNumber="10">169.73 (±4.18)</td>
<td id="52345D98FFD90061FC8A4F8FFC24F988" box="[801,953,1522,1544]" gridcol="2" gridrow="22" pageId="9" pageNumber="10">170.00 (±1.41)</td>
<td id="52345D98FFD90061FBCF4F8FFB66F988" box="[1124,1275,1522,1544]" gridcol="3" gridrow="22" pageId="9" pageNumber="10">165.30 (±4.55)</td>
</tr>
<tr id="11E534E4FFD90061FF374C66FB66F9B1" box="[156,1275,1563,1585]" gridrow="23" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374C66FEAEF9B1" box="[156,307,1563,1585]" gridcol="0" gridrow="23" pageId="9" pageNumber="10">LABNAS</th>
<td id="52345D98FFD90061FE744C66FDE8F9B1" box="[479,629,1563,1585]" gridcol="1" gridrow="23" pageId="9" pageNumber="10">0.59 (±0.14)</td>
<td id="52345D98FFD90061FC8A4C66FC24F9B1" box="[801,953,1563,1585]" gridcol="2" gridrow="23" pageId="9" pageNumber="10">0.75 (±0.00)</td>
<td id="52345D98FFD90061FBCF4C66FB66F9B1" box="[1124,1275,1563,1585]" gridcol="3" gridrow="23" pageId="9" pageNumber="10">0.66 (±0.22)</td>
</tr>
<tr id="11E534E4FFD90061FF374C38FB66F9DB" box="[156,1275,1605,1627]" gridrow="24" pageId="9" pageNumber="10">
<th id="52345D98FFD90061FF374C38FEAEF9DB" box="[156,307,1605,1627]" gridcol="0" gridrow="24" pageId="9" pageNumber="10">DEYE</th>
<td id="52345D98FFD90061FE744C38FDE8F9DB" box="[479,629,1605,1627]" gridcol="1" gridrow="24" pageId="9" pageNumber="10">4.24 (±0.46)</td>
<td id="52345D98FFD90061FC8A4C38FC24F9DB" box="[801,953,1605,1627]" gridcol="2" gridrow="24" pageId="9" pageNumber="10">3.69 (±0.75)</td>
<td id="52345D98FFD90061FBCF4C38FB66F9DB" box="[1124,1275,1605,1627]" gridcol="3" gridrow="24" pageId="9" pageNumber="10">3.91 (±0.67)</td>
</tr>
</table>
</paragraph>
<paragraph id="AF6A36A6FFD9FF97FF6E4CE4FD6AF86E" blockId="9.[151,1436,1689,2030]" pageId="9" pageNumber="10">
Compared to
<emphasis id="9DA1EAB4FFD9FF97FEC94CE4FD94F931" box="[354,521,1689,1713]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="68D54D25FFD9FF97FEC94CE4FE79F931" authorityName="Kramer" authorityYear="1977" box="[354,484,1689,1713]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFD9FF97FE464CE7FD94F931" box="[493,521,1690,1713]" pageId="9" pageNumber="10" sensu="stricto">s.s</taxonomicNameLabel>
</emphasis>
(
<tableCitation id="E257031DFFD9FF97FDB04CE4FDEFF931" box="[539,626,1689,1713]" captionStart="TABLE 3" captionStartId="13.[151,234,1575,1598]" captionTargetBox="[156,1155,1675,2028]" captionTargetPageId="13" captionText="TABLE 3. Mean and standard deviations of morphological characters that in combination can be used to distinguish C. macrops s.s. and C. rubeus sp. nov. Data are for the maximum number of specimens available for that character. Cm = C. macrops s.s., Cr = C. rubeus. Character abbreviations are explained in Appendix 1." pageId="9" pageNumber="10">Table 3</tableCitation>
), in both sexes
<taxonomicName id="68D54D25FFD9FF97FC874CE4FC07F931" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[812,922,1689,1713]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD9FF97FC874CE4FC07F931" box="[812,922,1689,1713]" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
has a more prominent lateral stripe with the white area covering a larger proportion of the first scale row (SCR1), less keeled body scales at mid-body (BSCK), and the scale reduction from 19 to 17 dorsal scale rows (VS19TO17) occurs further away from the head. In addition, there are never any small scales present between the nasal scale and the scale bordering the anterior edge of the pit (NASPIT) in
<taxonomicName id="68D54D25FFD9FF97FED04D5BFE75F8BE" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[379,488,1830,1854]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD9FF97FED04D5BFE75F8BE" box="[379,488,1830,1854]" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
, while they sometimes occur in
<taxonomicName id="68D54D25FFD9FF97FCFB4D5BFC4DF8BE" authorityName="Kramer" authorityYear="1977" box="[848,976,1830,1854]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFD9FF97FCFB4D5BFC4DF8BE" box="[848,976,1830,1854]" italics="true" pageId="9" pageNumber="10">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD9FF97FC7C4D5AFC66F8BE" box="[983,1019,1831,1854]" pageId="9" pageNumber="10" sensu="stricto">
<emphasis id="9DA1EAB4FFD9FF97FC7C4D5AFC6EF8BE" box="[983,1011,1831,1854]" italics="true" pageId="9" pageNumber="10">s.s</emphasis>
.
</taxonomicNameLabel>
In addition, males tend to have a less prominent postocular stripe (SCROC), fewer sublabial scales (SUBLAB), fewer scales between the rear edges of the supraocular scales (BTWSUPOC2), fewer scales between the last supralabial and the chin shields (VENTEDGE), and less keeled temporal scales (KTEMP). In females,
<taxonomicName id="68D54D25FFD9FF97FCCA4DEDFC53F828" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[865,974,1936,1960]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFD9FF97FCCA4DEDFC53F828" box="[865,974,1936,1960]" italics="true" pageId="9" pageNumber="10">C. rubeus</emphasis>
</taxonomicName>
tends to have fewer ventral scales, fewer (larger) scales bordering the supraoculars (BORSUPOC), and a larger number of scales between the rear edges of the supraoculars (BTWSUPOC2) than
<taxonomicName id="68D54D25FFD9FF97FDE74DABFD56F86E" authorityName="Kramer" authorityYear="1977" box="[588,715,2006,2030]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFD9FF97FDE74DABFD56F86E" box="[588,715,2006,2030]" italics="true" pageId="9" pageNumber="10">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFD9FF97FD784DAAFD6AF86E" box="[723,759,2007,2030]" pageId="9" pageNumber="10" sensu="stricto">
<emphasis id="9DA1EAB4FFD9FF97FD784DAAFD72F86E" box="[723,751,2007,2030]" italics="true" pageId="9" pageNumber="10">s.s</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="FBAA662EFFDAFF94FF3C4DC9FD62F869" ID-DOI="http://doi.org/10.5281/zenodo.5291125" ID-Zenodo-Dep="5291125" httpUri="https://zenodo.org/record/5291125/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1972,1995]" targetBox="[253,1314,199,1920]" targetPageId="10">
<paragraph id="AF6A36A6FFDAFF94FF3C4DC9FD62F869" blockId="10.[151,1436,1972,2025]" pageId="10" pageNumber="11">
<emphasis id="9DA1EAB4FFDAFF94FF3C4DC9FE91F84A" bold="true" box="[151,268,1972,1995]" pageId="10" pageNumber="11">FIGURE 5</emphasis>
. Line drawings of the head scalation of the holotype of
<taxonomicName id="68D54D25FFDAFF94FCE04DC9FBBCF84A" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[843,1057,1972,1994]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
<emphasis id="9DA1EAB4FFDAFF94FCE04DC9FBBCF84A" box="[843,1057,1972,1994]" italics="true" pageId="10" pageNumber="11">Cryptelytrops rubeus</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFDAFF94FB834DC8FBE4F84A" bold="true" box="[1064,1145,1973,1994]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="869257CFFFDAFF94FB834DC8FBE4F84A" box="[1064,1145,1973,1994]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(FMNH 262718). Labelling as in Fig. 3 (a: 4.00mm, b: 2.05mm, c: 3.45mm, d: 24.9mm).
</paragraph>
</caption>
<caption id="FBAA662EFFDBFF95FF3C4D17FB8CF83F" ID-DOI="http://doi.org/10.5281/zenodo.5291129" ID-Zenodo-Dep="5291129" httpUri="https://zenodo.org/record/5291129/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1898,1921]" targetBox="[235,1346,193,1865]" targetPageId="11">
<paragraph id="AF6A36A6FFDBFF95FF3C4D17FA01F81C" blockId="11.[151,1436,1898,1983]" pageId="11" pageNumber="12">
<emphasis id="9DA1EAB4FFDBFF95FF3C4D17FE8FF801" bold="true" box="[151,274,1898,1921]" pageId="11" pageNumber="12">FIGURE 6.</emphasis>
<taxonomicName id="68D54D25FFDBFF95FE8E4D16FE66F801" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[293,507,1899,1921]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
<emphasis id="9DA1EAB4FFDBFF95FE8E4D16FE66F801" box="[293,507,1899,1921]" italics="true" pageId="11" pageNumber="12">Cryptelytrops rubeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFDBFF95FDAA4D11FDD7F801" box="[513,586,1900,1921]" pageId="11" pageNumber="12" rank="species">sp.nov.</taxonomicNameLabel>
A. Close-up of head of male (left) and female (right) from vicinity of Ho Chi Minh city, South Vietnam (photos: A Malhotra); B. Close-up of head of male (left) and female (right) from South Vietnam; C.
</paragraph>
<paragraph id="AF6A36A6FFDBFF95FF3C4DD4FB8CF83F" blockId="11.[151,1436,1898,1983]" box="[151,1041,1961,1983]" pageId="11" pageNumber="12">Closeup of tail of male from B; D. Pair from B (male above) (photos: K Udomritthiruj).</paragraph>
</caption>
<caption id="FBAA662EFFDCFF92FF3C4E15FAD0FB49" pageId="12" pageNumber="13" startId="12.[151,229,1128,1153]" targetBox="[159,1427,198,1104]" targetPageId="12">
<paragraph id="AF6A36A6FFDCFF92FF3C4E15FAD0FB49" blockId="12.[151,1436,1128,1226]" pageId="12" pageNumber="13">
<emphasis id="9DA1EAB4FFDCFF92FF3C4E15FEF3FB01" bold="true" box="[151,366,1128,1153]" pageId="12" pageNumber="13">
<figureCitation id="37EE2A23FFDCFF92FF3C4E15FF62FB01" box="[151,255,1128,1153]" captionStart="FIGURE 6" captionStartId="11.[151,250,1898,1921]" captionTargetBox="[235,1346,193,1865]" captionTargetId="figure-72@11.[232,1347,193,1869]" captionTargetPageId="11" captionText="FIGURE 6. Cryptelytrops rubeus sp.nov. A. Close-up of head of male (left) and female (right) from vicinity of Ho Chi Minh city, South Vietnam (photos: A Malhotra); B. Close-up of head of male (left) and female (right) from South Vietnam; C. Closeup of tail of male from B; D. Pair from B (male above) (photos: K Udomritthiruj)." figureDoi="http://doi.org/10.5281/zenodo.5291129" httpUri="https://zenodo.org/record/5291129/files/figure.png" pageId="12" pageNumber="13">Figure 6</figureCitation>
(contd.).
</emphasis>
<taxonomicName id="68D54D25FFDCFF92FED24E14FDF5FB01" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[377,616,1129,1153]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">Cryptelytrops rubeus</taxonomicName>
<taxonomicNameLabel id="869257CFFFDCFF92FDDF4E17FD52FB01" box="[628,719,1130,1153]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
E. Adult specimen from
<collectingRegion id="6D11F844FFDCFF92FBAB4E14FB58FB01" box="[1024,1221,1129,1153]" country="Cambodia" name="Mondolkiri" pageId="12" pageNumber="13">Mondolkiri Prov.</collectingRegion>
,
<collectingCountry id="D7C27636FFDCFF92FB624E14FAA2FB01" box="[1225,1343,1129,1153]" name="Cambodia" pageId="12" pageNumber="13">Cambodia</collectingCountry>
(photo: Bryan L Stuart); F: Adult male from Nam Cat Tien National Park,
<collectingCountry id="D7C27636FFDCFF92FC2C4EF0FC7BFB25" box="[903,998,1165,1189]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
(photo: Peter Paul van Dijk); G: Adult male from O Reang, Ko Seima Protected Forest.
<collectingRegion id="6D11F844FFDCFF92FD154ECCFCDDFB49" box="[702,832,1201,1225]" country="Cambodia" name="Mondolkiri" pageId="12" pageNumber="13">Mondolkiri</collectingRegion>
, northwest
<collectingCountry id="D7C27636FFDCFF92FC6B4ECCFBAEFB49" box="[960,1075,1201,1225]" name="Cambodia" pageId="12" pageNumber="13">Cambodia</collectingCountry>
(photo: Jeremy Holden).
</paragraph>
</caption>
<subSubSection id="E7CF652DFFDCFF93FF6E4E85FC1BFA78" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="AF6A36A6FFDCFF92FF6E4E85FBF8F965" blockId="12.[151,1436,1272,1766]" pageId="12" pageNumber="13">
<emphasis id="9DA1EAB4FFDCFF92FF6E4E85FE79FA90" bold="true" box="[197,484,1272,1297]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">
Description of
<typeStatus id="706E8804FFDCFF92FEDC4E84FE40FA91" box="[375,477,1273,1297]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
.
</emphasis>
Body cylindrical; head triangular in dorsal aspect, very distinct from neck. Canthus rostralis distinct. Head scales small (no large dorsal shields), smooth except for weakly keeled scales in temporal region, region above suprabials, and rear of head. Dorsal body scales weakly keeled. 163 ventral scales, 58 pairs subcaudal scales, 21 dorsal scales at mid
<emphasis id="9DA1EAB4FFDCFF92FDC74F1BFDE7FAFD" box="[620,634,1382,1405]" italics="true" pageId="12" pageNumber="13"></emphasis>
body. Rostral scale roughly triangular, upper edge about half width of lower edge. Pupil vertically elliptical. Loreal pit present. Nostril completely enclosed in nasal scale. Nasal scale partially fused with first supralabial, suture extending part way towards nostril on both sides. Eleven and 12 supralabials on right and left sides respectively, 12 sublabials, two postocular scales. Shield bordering anterior edge of pit fused with second supralabial, lacking any small scales between it and nasal. Subocular scale separated by one scale from third, fourth, and fifth supralabials. At least 8 scales between supraoculars, 14 scales between their posterior edges. Five teeth on palatine, 16 on pterygoid, and 16 on dentary. Snout
<emphasis id="9DA1EAB4FFDCFF92FB864C43FBA6F9D5" box="[1069,1083,1598,1621]" italics="true" pageId="12" pageNumber="13"></emphasis>
vent length
<quantity id="682D9B43FFDCFF92FB6B4C40FA82F9D5" box="[1216,1311,1597,1621]" metricMagnitude="-1" metricUnit="m" metricValue="4.99" pageId="12" pageNumber="13" unit="cm" value="49.9">49.9 cm</quantity>
; tail length
<quantity id="682D9B43FFDCFF92FF3C4C1CFF75F9F9" box="[151,232,1633,1657]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="12" pageNumber="13" unit="cm" value="9.1">9.1 cm</quantity>
. Head length measured from tip of snout to posterior edge of lower jawbone
<quantity id="682D9B43FFDCFF92FB924C1CFB3CF9F9" box="[1081,1185,1633,1657]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="12" pageNumber="13" unit="mm" value="24.9">24.9 mm</quantity>
; head width measured between rear outer edges of supraoculars
<quantity id="682D9B43FFDCFF92FDC64CF8FD4EF91D" box="[621,723,1669,1693]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="12" pageNumber="13" unit="mm" value="11.4">11.4 mm</quantity>
, at widest point of head
<quantity id="682D9B43FFDCFF92FC414CF8FBC2F91D" box="[1002,1119,1669,1693]" metricMagnitude="-2" metricUnit="m" metricValue="1.685" pageId="12" pageNumber="13" unit="mm" value="16.85">16.85 mm</quantity>
. Supraocular scale (on right hand side)
<quantity id="682D9B43FFDCFF92FEB84CD4FEF5F941" box="[275,360,1705,1729]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="12" pageNumber="13" unit="mm" value="5.7">5.7 mm</quantity>
long,
<quantity id="682D9B43FFDCFF92FE1A4CD4FD89F941" box="[433,532,1705,1729]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="13" unit="mm" value="2.05">2.05 mm</quantity>
wide. Distance from eye to nostril 5.0 mm, eye to pit 1.0 mm, posterior edge of pit to anterior edge of nostril
<quantity id="682D9B43FFDCFF92FE744CB0FDDBF965" box="[479,582,1741,1765]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" pageId="12" pageNumber="13" unit="mm" value="4.85">4.85 mm</quantity>
. Eye diameter 4.0 mm. Scale reduction formula:
</paragraph>
<paragraph id="AF6A36A6FFDCFF93FF6E4DFDFACFFF54" blockId="12.[151,1436,1920,2017]" lastBlockId="13.[151,1436,151,1528]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="9DA1EAB4FFDCFF92FF6E4DFDFDFBF818" bold="true" box="[197,614,1920,1945]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">
Colour of
<typeStatus id="706E8804FFDCFF92FE944DFCFE3BF819" box="[319,422,1921,1945]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dark brownish grey on dorsal surface of head, fading to steely blue grey laterally. Ventral scales are very similar in colour to the first few dorsal scale rows. Pale lateral stripe present, covering approximately one
<emphasis id="9DA1EAB4FFDCFF92FE004DB7FE24F861" box="[427,441,1994,2017]" italics="true" pageId="12" pageNumber="13"></emphasis>
quarter of first scale row at mid-body, fading towards the head. Upper and lower labials not distinct in colour from rest of head. Paired chin shields, scales between the chin shields creamy white in colour, although a single pair of chinshields and the occasional scale blue
<emphasis id="9DA1EAB4FFDDFF93FCD34AC0FC1BFF54" box="[888,902,189,212]" italics="true" pageId="13" pageNumber="14"></emphasis>
grey. Ventral scales uniformly blue
<emphasis id="9DA1EAB4FFDDFF93FAA64AC0FA86FF54" box="[1293,1307,189,212]" italics="true" pageId="13" pageNumber="14"></emphasis>
grey.
</paragraph>
<paragraph id="AF6A36A6FFDDFF93FF6E4AA2FF59FE2C" blockId="13.[151,1436,151,1528]" pageId="13" pageNumber="14">
<emphasis id="9DA1EAB4FFDDFF93FF6E4AA2FEF4FF78" bold="true" box="[197,361,223,248]" inLineHeading="true" pageId="13" pageNumber="14" reason="1">Colour in life.</emphasis>
Photographs of a red
<emphasis id="9DA1EAB4FFDDFF93FDFD4A9CFDF9FF78" box="[598,612,225,248]" italics="true" pageId="13" pageNumber="14"></emphasis>
eyed green pitviper which is almost certainly referable to
<taxonomicName id="68D54D25FFDDFF93FB4E4A9DFACFFF78" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[1253,1362,224,248]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFDDFF93FB4E4A9DFACFFF78" box="[1253,1362,224,248]" italics="true" pageId="13" pageNumber="14">C. rubeus</emphasis>
</taxonomicName>
, taken in Nam Cat Tien National Park in
<date id="DB6B1066FFDDFF93FDB54B79FD08FE9C" box="[542,661,260,284]" pageId="13" pageNumber="14" value="2000-05">May 2000</date>
, were supplied by P-P. van Dijk. This specimen, together with those pictured in
<bibRefCitation id="CB444B57FFDDFF93FEB04B5AFDB2FEBF" author="Gumprecht, A. &amp; Tillack, F. &amp; Orlov, N. L. &amp; Captain, A. &amp; Ryabov, S." box="[283,559,295,320]" pageId="13" pageNumber="14" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. &amp; Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
Gumprecht
<emphasis id="9DA1EAB4FFDDFF93FE0A4B54FE49FEC0" box="[417,468,296,320]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. (2004)
</bibRefCitation>
(p. 236
<emphasis id="9DA1EAB4FFDDFF93FD214B54FD05FEC0" box="[650,664,297,320]" italics="true" pageId="13" pageNumber="14"></emphasis>
238), photographs taken by AM of the specimens from Cu Chi and Bien Hoa above, of FMNH 262717 by BLS in
<collectingCountry id="D7C27636FFDDFF93FD644B31FCDFFEE4" box="[719,834,332,356]" name="Cambodia" pageId="13" pageNumber="14">Cambodia</collectingCountry>
(described in
<bibRefCitation id="CB444B57FFDDFF93FC5B4B31FB45FEE4" author="Stuart, B. L. &amp; Sok, K. &amp; Neang, T." box="[1008,1240,332,356]" pageId="13" pageNumber="14" pagination="129 - 155" refId="ref12518" refString="Stuart, B. L., Sok, K. &amp; Neang, T. (2006 a) A collection of amphibians and reptiles from hilly eastern Cambodia. Raffles Bulletin of Zoology, 54, 129 - 155." type="journal article" year="2006">
Stuart
<emphasis id="9DA1EAB4FFDDFF93FB954B30FBEEFEE4" box="[1086,1139,332,356]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. [2006a
</bibRefCitation>
), and of BMNH 2005.1602, provided by K. Udomritthiruj (
<figureCitation id="37EE2A23FFDDFF93FDDF4B12FD25FE07" box="[628,696,367,391]" captionStart="FIGURE 6" captionStartId="11.[151,250,1898,1921]" captionTargetBox="[235,1346,193,1865]" captionTargetId="figure-72@11.[232,1347,193,1869]" captionTargetPageId="11" captionText="FIGURE 6. Cryptelytrops rubeus sp.nov. A. Close-up of head of male (left) and female (right) from vicinity of Ho Chi Minh city, South Vietnam (photos: A Malhotra); B. Close-up of head of male (left) and female (right) from South Vietnam; C. Closeup of tail of male from B; D. Pair from B (male above) (photos: K Udomritthiruj)." figureDoi="http://doi.org/10.5281/zenodo.5291129" httpUri="https://zenodo.org/record/5291129/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
), were used to write this description of the colour of the species in life.
</paragraph>
<paragraph id="AF6A36A6FFDDFF93FF6E4BCAFE79FCB2" blockId="13.[151,1436,151,1528]" pageId="13" pageNumber="14">Posterior half of the body dark to grass green, similar on ventral and dorsal surface. Lateral white stripe more prominent and may extend onto the second dorsal scale row in males (confined to first row in females). In males, lateral white stripe may be edged below with dull reddishbrown more obvious on the rear half of the body. White stripe may also extend onto head as postocular stripe, passing underneath the eye and fading out on lower preocular scale. Upper lip (supralabials and 12 scale rows above) suffused with pale blue and differentiated from rest of head; mostly confined to scale edges, but may be more widespread towards rear of mouth. Pale blue also largely covering sublabials, genials and paired chinshields, extending onto central part of the ventral scales in anterior of body. Scales between the chinshields and the sublabials creamy white. Patches of other colours including deep yellow also present on these scales. Interstitial skin black. Tail solid orangered, this extending to vent dorsally and ventrally (although in males forming distinct pattern of blotches on the distal half of the tail). Eye bright or deep reddishorange in both sexes.</paragraph>
<paragraph id="AF6A36A6FFDDFF93FF6E4943FC1BFA78" blockId="13.[151,1436,151,1528]" pageId="13" pageNumber="14">
<emphasis id="9DA1EAB4FFDDFF93FF6E4943FEA1FCD6" box="[197,316,830,855]" inLineHeading="true" pageId="13" pageNumber="14" reason="1">
<emphasis id="9DA1EAB4FFDDFF93FF6E4943FEA8FCD7" bold="true" box="[197,309,830,855]" pageId="13" pageNumber="14">Variation</emphasis>
.
</emphasis>
Sexual dimorphism is not very obvious although females lack postocular stripes and have a less prominent lateral stripe, never edged with brick red. Eye colour appears to be the same in both sexes, with the exception of
<specimenCount id="B9D3FD2FFFDDFF93FE8349F8FE3EFC1D" box="[296,419,901,925]" pageId="13" pageNumber="14" type="female">one female</specimenCount>
from southern
<collectingCountry id="D7C27636FFDDFF93FDE649F8FD31FC1D" box="[589,684,901,925]" name="Vietnam" pageId="13" pageNumber="14">Vietnam</collectingCountry>
(BMNH 2005.1602), in which the eye was golden yellow. Females have relatively shorter tails and fewer subcaudals (52
<emphasis id="9DA1EAB4FFDDFF93FD4549D4FD61FC40" box="[750,764,937,960]" italics="true" pageId="13" pageNumber="14"></emphasis>
63, versus
<quantity id="682D9B43FFDDFF93FCDE49D5FC44FC40" box="[885,985,936,960]" metricMagnitude="0" metricUnit="m" metricValue="1.7399" metricValueMax="1.8796" metricValueMin="1.6002" pageId="13" pageNumber="14" unit="in" value="68.5" valueMax="74.0" valueMin="63.0">
63
<emphasis id="9DA1EAB4FFDDFF93FC3A49D4FC02FC40" box="[913,927,937,960]" italics="true" pageId="13" pageNumber="14"></emphasis>
74 in
</quantity>
males). Sexual dimorphism in size may also be low/absent in this species (maximum recorded
<quantity id="682D9B43FFDDFF93FCAE49B1FCFDFC63" box="[773,864,972,996]" metricMagnitude="-1" metricUnit="m" metricValue="5.029999999999999" pageId="13" pageNumber="14" unit="cm" value="50.3">50.3 cm</quantity>
SVL versus
<quantity id="682D9B43FFDDFF93FC5E49B1FBCDFC63" box="[1013,1104,972,996]" metricMagnitude="-1" metricUnit="m" metricValue="5.05" pageId="13" pageNumber="14" unit="cm" value="50.5">50.5 cm</quantity>
for males), but the sample is small. The number of ventral scales is similar in both sexes (159 and 172). Body scales smooth to strongly keeled at mid
<emphasis id="9DA1EAB4FFDDFF93FF764E69FF76FBAB" box="[221,235,1044,1067]" italics="true" pageId="13" pageNumber="14"></emphasis>
body. Temporal scales show weak to strong tubercular keeling, and keeling is also weak to strongly present on the scales covering the rear of the head. In some specimens, the scales in the region between the temporal scales and supralabials are also keeled. The ratio between the lengths of the upper and lower edges of the rostral scale varies between 0.39 and 0.53. Supralabials vary between 8 and 12 and sublabials between 9 and 13. The suture that partially separates the first supralabial and nasal scale is more obliquely inclined than in the other species, where it tends to be more horizontal, and tends to be confined to the rear of the nostril. The minimum number of scales between supraoculars varies between 7 and 12 and there are 7 and 19 scales between the posterior edges of the supraoculars. There are never any small scales between the nasals and shield bordering the anterior of the pit, and exceptionally there was a single scale between the internasals in
<specimenCount id="B9D3FD2FFFDDFF93FCD24F52FB8AFAC6" box="[889,1047,1326,1350]" pageId="13" pageNumber="14" type="generic">one specimen</specimenCount>
. There are 0
<emphasis id="9DA1EAB4FFDDFF93FB0F4F52FB2FFAC6" box="[1188,1202,1327,1350]" italics="true" pageId="13" pageNumber="14"></emphasis>
1 scales between the third and fourth supralabials and the subocular, and 1
<emphasis id="9DA1EAB4FFDDFF93FCAD4F2EFC89FAEA" box="[774,788,1363,1386]" italics="true" pageId="13" pageNumber="14"></emphasis>
2 scales between the fifth supralabial and the subocular scale. Hemipenes do not differ notably from those of
<taxonomicName id="68D54D25FFDDFF93FD564F0BFCE2FA0D" authorityName="Kramer" authorityYear="1977" box="[765,895,1398,1422]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFDDFF93FD564F0BFCE2FA0D" box="[765,895,1398,1422]" italics="true" pageId="13" pageNumber="14">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFDDFF93FC234F0BFC36FA0D" box="[904,939,1398,1421]" pageId="13" pageNumber="14" sensu="stricto">
<emphasis id="9DA1EAB4FFDDFF93FC234F0BFC39FA0D" box="[904,932,1398,1421]" italics="true" pageId="13" pageNumber="14">s.s</emphasis>
.
</taxonomicNameLabel>
There are always five teeth on the palatine bone (as in most other pitvipers of the
<taxonomicName id="68D54D25FFDDFF93FDE34FE7FD41FA31" box="[584,732,1434,1457]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="9DA1EAB4FFDDFF93FDE34FE7FD41FA31" box="[584,732,1434,1457]" italics="true" pageId="13" pageNumber="14">Trimeresurus</emphasis>
</taxonomicName>
radiation), while those on the pterygoid vary between and 15
<emphasis id="9DA1EAB4FFDDFF93FA254FE7FA01FA31" box="[1422,1436,1434,1457]" italics="true" pageId="13" pageNumber="14"></emphasis>
18 (in
<specimenCount id="B9D3FD2FFFDDFF93FF484FC0FE1CFA54" box="[227,385,1468,1492]" pageId="13" pageNumber="14" type="generic">one specimen</specimenCount>
, a count of 12 was made on one side, but the other side had 16, a count that was within the range seen in other specimens), and on the dentary between 14
<emphasis id="9DA1EAB4FFDDFF93FCFE4F9CFCFEFA78" box="[853,867,1505,1528]" italics="true" pageId="13" pageNumber="14"></emphasis>
17.
</paragraph>
</subSubSection>
<caption id="FBAA662EFFDDFF93FF3C4C5AFC7DF9FA" pageId="13" pageNumber="14" startId="13.[151,234,1575,1598]" targetBox="[156,1155,1675,2028]" targetIsTable="true" targetPageId="13">
<paragraph id="AF6A36A6FFDDFF93FF3C4C5AFC7DF9FA" blockId="13.[151,1436,1575,1659]" pageId="13" pageNumber="14">
<emphasis id="9DA1EAB4FFDDFF93FF3C4C5AFE95F9BE" bold="true" box="[151,264,1575,1598]" pageId="13" pageNumber="14">TABLE 3.</emphasis>
Mean and standard deviations of morphological characters that in combination can be used to distinguish
<taxonomicName id="68D54D25FFDDFF93FA2F4C5AFF70F9DC" authorityName="Kramer" authorityYear="1977" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFDDFF93FA2F4C5AFF70F9DC" italics="true" pageId="13" pageNumber="14">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFDDFF93FF5D4C3AFE8BF9DC" box="[246,278,1607,1628]" pageId="13" pageNumber="14" sensu="stricto">s.s.</taxonomicNameLabel>
and
<taxonomicName id="68D54D25FFDDFF93FEE74C3BFE2CF9DC" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[332,433,1606,1628]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
<emphasis id="9DA1EAB4FFDDFF93FEE74C3BFE2CF9DC" box="[332,433,1606,1628]" italics="true" pageId="13" pageNumber="14">C. rubeus</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFDDFF93FE104C3BFD93F9DB" bold="true" box="[443,526,1606,1627]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="869257CFFFDDFF93FE104C3BFD93F9DB" box="[443,526,1606,1627]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Data are for the maximum number of specimens available for that character. Cm =
<emphasis id="9DA1EAB4FFDDFF93FA2F4C3BFE89F9FA" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="68D54D25FFDDFF93FA2F4C3BFF70F9FA" authorityName="Kramer" authorityYear="1977" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFDDFF93FF584C18FE89F9FA" box="[243,276,1637,1658]" pageId="13" pageNumber="14" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
, Cr =
<taxonomicName id="68D54D25FFDDFF93FEFF4C18FE2AF9FB" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[340,439,1637,1659]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFDDFF93FEFF4C18FE2AF9FB" box="[340,439,1637,1659]" italics="true" pageId="13" pageNumber="14">C. rubeus</emphasis>
</taxonomicName>
. Character abbreviations are explained in Appendix 1.
</paragraph>
</caption>
<paragraph id="AF6A36A6FFDDFF93FF374CF6FB1FF86C" pageId="13" pageNumber="14">
<table id="DDD5C406FFDD0061FF374CF6FB1EF86C" box="[156,1155,1675,2028]" colsContinueIn="14.[156,1167,160,678]" gridcols="3" gridrows="9" pageId="13" pageNumber="14">
<tr id="11E534E4FFDD0061FF374CF6FB1EF921" box="[156,1155,1675,1697]" gridrow="0" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374CF6FEAEF921" box="[156,307,1675,1697]" gridcol="0" gridrow="0" pageId="13" pageNumber="14">Character</th>
<th id="52345D98FFDD0061FDE14CF6FD49F921" box="[586,724,1675,1697]" gridcol="1" gridrow="0" pageId="13" pageNumber="14">Cm</th>
<th id="52345D98FFDD0061FC534CF6FB1EF921" box="[1016,1155,1675,1697]" gridcol="2" gridrow="0" pageId="13" pageNumber="14">Cr</th>
</tr>
<tr id="11E534E4FFDD0061FF374CC9FB1EF94A" box="[156,1155,1716,1738]" gridrow="1" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374CC9FB1EF94A" box="[156,1155,1716,1738]" colspan="3" colspanRight="2" gridcol="0" gridrow="1" pageId="13" pageNumber="14">a. Males</th>
</tr>
<tr id="11E534E4FFDD0061FF374CA3FB1EF974" box="[156,1155,1758,1780]" gridrow="2" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374CA3FEAEF974" box="[156,307,1758,1780]" gridcol="0" gridrow="2" pageId="13" pageNumber="14">SCROC</th>
<td id="52345D98FFDD0061FDE14CA3FD49F974" box="[586,724,1758,1780]" gridcol="1" gridrow="2" pageId="13" pageNumber="14">0.69 (±0.93)</td>
<td id="52345D98FFDD0061FC534CA3FB1EF974" box="[1016,1155,1758,1780]" gridcol="2" gridrow="2" pageId="13" pageNumber="14">0.33 (±0.82)</td>
</tr>
<tr id="11E534E4FFDD0061FF374D7AFB1EF89D" box="[156,1155,1799,1821]" gridrow="3" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374D7AFEAEF89D" box="[156,307,1799,1821]" gridcol="0" gridrow="3" pageId="13" pageNumber="14">SCR1</th>
<td id="52345D98FFDD0061FDE14D7AFD49F89D" box="[586,724,1799,1821]" gridcol="1" gridrow="3" pageId="13" pageNumber="14">0.19 (±0.19)</td>
<td id="52345D98FFDD0061FC534D7AFB1EF89D" box="[1016,1155,1799,1821]" gridcol="2" gridrow="3" pageId="13" pageNumber="14">0.32 (±0.08)</td>
</tr>
<tr id="11E534E4FFDD0061FF374D4DFB1EF8C6" box="[156,1155,1840,1862]" gridrow="4" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374D4DFEAEF8C6" box="[156,307,1840,1862]" gridcol="0" gridrow="4" pageId="13" pageNumber="14">BSCK</th>
<td id="52345D98FFDD0061FDE14D4DFD49F8C6" box="[586,724,1840,1862]" gridcol="1" gridrow="4" pageId="13" pageNumber="14">0.8 (±0.25)</td>
<td id="52345D98FFDD0061FC534D4DFB1EF8C6" box="[1016,1155,1840,1862]" gridcol="2" gridrow="4" pageId="13" pageNumber="14">0.38 (±0.21)</td>
</tr>
<tr id="11E534E4FFDD0061FF374D27FB1EF8F0" box="[156,1155,1882,1904]" gridrow="5" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374D27FEAEF8F0" box="[156,307,1882,1904]" gridcol="0" gridrow="5" pageId="13" pageNumber="14">VS19TO17</th>
<td id="52345D98FFDD0061FDE14D27FD49F8F0" box="[586,724,1882,1904]" gridcol="1" gridrow="5" pageId="13" pageNumber="14">68.00 (±1.99)</td>
<td id="52345D98FFDD0061FC534D27FB1EF8F0" box="[1016,1155,1882,1904]" gridcol="2" gridrow="5" pageId="13" pageNumber="14">69.29 (±1.53)</td>
</tr>
<tr id="11E534E4FFDD0061FF374DFEFB1EF819" box="[156,1155,1923,1945]" gridrow="6" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374DFEFEAEF819" box="[156,307,1923,1945]" gridcol="0" gridrow="6" pageId="13" pageNumber="14">SUPLAB</th>
<td id="52345D98FFDD0061FDE14DFEFD49F819" box="[586,724,1923,1945]" gridcol="1" gridrow="6" pageId="13" pageNumber="14">9.65 (±0.49)</td>
<td id="52345D98FFDD0061FC534DFEFB1EF819" box="[1016,1155,1923,1945]" gridcol="2" gridrow="6" pageId="13" pageNumber="14">9.58 (± 1.02)</td>
</tr>
<tr id="11E534E4FFDD0061FF374DD1FB1EF842" box="[156,1155,1964,1986]" gridrow="7" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374DD1FEAEF842" box="[156,307,1964,1986]" gridcol="0" gridrow="7" pageId="13" pageNumber="14">SUBLAB</th>
<td id="52345D98FFDD0061FDE14DD1FD49F842" box="[586,724,1964,1986]" gridcol="1" gridrow="7" pageId="13" pageNumber="14">11.62 (±0.54)</td>
<td id="52345D98FFDD0061FC534DD1FB1EF842" box="[1016,1155,1964,1986]" gridcol="2" gridrow="7" pageId="13" pageNumber="14">10.92 (±0.74)</td>
</tr>
<tr id="11E534E4FFDD0061FF374DABFB1EF86C" box="[156,1155,2006,2028]" gridrow="8" pageId="13" pageNumber="14">
<th id="52345D98FFDD0061FF374DABFEAEF86C" box="[156,307,2006,2028]" gridcol="0" gridrow="8" pageId="13" pageNumber="14">BTWSUPOC2</th>
<td id="52345D98FFDD0061FDE14DABFD49F86C" box="[586,724,2006,2028]" gridcol="1" gridrow="8" pageId="13" pageNumber="14">11.96 (±1.22)</td>
<td id="52345D98FFDD0061FC534DABFB1EF86C" box="[1016,1155,2006,2028]" gridcol="2" gridrow="8" pageId="13" pageNumber="14">10.17 (±1.94)</td>
</tr>
</table>
</paragraph>
<paragraph id="AF6A36A6FFDEFF90FF374ADDFBEBFD26" pageId="14" pageNumber="15">
<table id="DDD5C406FFDE0061FF374ADDFB12FD26" box="[156,1167,160,678]" colsContinueFrom="13.[156,1155,1675,2028]" gridcols="3" gridrows="13" pageId="14" pageNumber="15">
<tr id="11E534E4FFDE0061FF374ADDFB12FF36" box="[156,1167,160,182]" gridrow="0" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374ADDFEAEFF36" box="[156,307,160,182]" gridcol="0" gridrow="0" pageId="14" pageNumber="15">VENTEDGE</th>
<th id="52345D98FFDE0061FDE14ADDFD7CFF36" box="[586,737,160,182]" gridcol="1" gridrow="0" pageId="14" pageNumber="15">7.46 (±0.45)</th>
<th id="52345D98FFDE0061FC534ADDFB12FF36" box="[1016,1167,160,182]" gridcol="2" gridrow="0" pageId="14" pageNumber="15">6.92 (±0.49)</th>
</tr>
<tr id="11E534E4FFDE0061FF374AB4FB12FF5F" box="[156,1167,201,223]" gridrow="1" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374AB4FEAEFF5F" box="[156,307,201,223]" gridcol="0" gridrow="1" pageId="14" pageNumber="15">SC12TO10</th>
<td id="52345D98FFDE0061FDE14AB4FD7CFF5F" box="[586,737,201,223]" gridcol="1" gridrow="1" pageId="14" pageNumber="15">5.15 (±1.52)</td>
<td id="52345D98FFDE0061FC534AB4FB12FF5F" box="[1016,1167,201,223]" gridcol="2" gridrow="1" pageId="14" pageNumber="15">5.14 (±1.24)</td>
</tr>
<tr id="11E534E4FFDE0061FF374A8FFB12FE88" box="[156,1167,242,264]" gridrow="2" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374A8FFEAEFE88" box="[156,307,242,264]" gridcol="0" gridrow="2" pageId="14" pageNumber="15">NASPIT</th>
<td id="52345D98FFDE0061FDE14A8FFD7CFE88" box="[586,737,242,264]" gridcol="1" gridrow="2" pageId="14" pageNumber="15">0.19 (±0.47)</td>
<td id="52345D98FFDE0061FC534A8FFB12FE88" box="[1016,1167,242,264]" gridcol="2" gridrow="2" pageId="14" pageNumber="15">0.00 (±0.0)</td>
</tr>
<tr id="11E534E4FFDE0061FF374B66FB12FEB1" box="[156,1167,283,305]" gridrow="3" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374B66FEAEFEB1" box="[156,307,283,305]" gridcol="0" gridrow="3" pageId="14" pageNumber="15">KTEMP</th>
<td id="52345D98FFDE0061FDE14B66FD7CFEB1" box="[586,737,283,305]" gridcol="1" gridrow="3" pageId="14" pageNumber="15">0.67 (±0.31)</td>
<td id="52345D98FFDE0061FC534B66FB12FEB1" box="[1016,1167,283,305]" gridcol="2" gridrow="3" pageId="14" pageNumber="15">0.46 (±0.29)</td>
</tr>
<tr id="11E534E4FFDE0061FF374B38FB12FEDB" box="[156,1167,325,347]" gridrow="4" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374B38FB12FEDB" box="[156,1167,325,347]" colspan="3" colspanRight="2" gridcol="0" gridrow="4" pageId="14" pageNumber="15">b. Females</th>
</tr>
<tr id="11E534E4FFDE0061FF374B13FB12FE04" box="[156,1167,366,388]" gridrow="5" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374B13FEAEFE04" box="[156,307,366,388]" gridcol="0" gridrow="5" pageId="14" pageNumber="15">SCR1</th>
<td id="52345D98FFDE0061FDE14B13FD7CFE04" box="[586,737,366,388]" gridcol="1" gridrow="5" pageId="14" pageNumber="15">0.15 (±0.13)</td>
<td id="52345D98FFDE0061FC534B13FB12FE04" box="[1016,1167,366,388]" gridcol="2" gridrow="5" pageId="14" pageNumber="15">0.29 (±0.23)</td>
</tr>
<tr id="11E534E4FFDE0061FF374BE5FB12FE2E" box="[156,1167,408,430]" gridrow="6" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374BE5FEAEFE2E" box="[156,307,408,430]" gridcol="0" gridrow="6" pageId="14" pageNumber="15">VSC</th>
<td id="52345D98FFDE0061FDE14BE5FD7CFE2E" box="[586,737,408,430]" gridcol="1" gridrow="6" pageId="14" pageNumber="15">169.73 (±4.18)</td>
<td id="52345D98FFDE0061FC534BE5FB12FE2E" box="[1016,1167,408,430]" gridcol="2" gridrow="6" pageId="14" pageNumber="15">165.30 (±4.55)</td>
</tr>
<tr id="11E534E4FFDE0061FF374BBCFB12FE57" box="[156,1167,449,471]" gridrow="7" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374BBCFEAEFE57" box="[156,307,449,471]" gridcol="0" gridrow="7" pageId="14" pageNumber="15">VS19TO17</th>
<td id="52345D98FFDE0061FDE14BBCFD7CFE57" box="[586,737,449,471]" gridcol="1" gridrow="7" pageId="14" pageNumber="15">67.64 (±2.99)</td>
<td id="52345D98FFDE0061FC534BBCFB12FE57" box="[1016,1167,449,471]" gridcol="2" gridrow="7" pageId="14" pageNumber="15">69.26 (±2.74)</td>
</tr>
<tr id="11E534E4FFDE0061FF374B97FB12FD80" box="[156,1167,490,512]" gridrow="8" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374B97FEAEFD80" box="[156,307,490,512]" gridcol="0" gridrow="8" pageId="14" pageNumber="15">BORSUPOC</th>
<td id="52345D98FFDE0061FDE14B97FD7CFD80" box="[586,737,490,512]" gridcol="1" gridrow="8" pageId="14" pageNumber="15">9.70 (±1.19)</td>
<td id="52345D98FFDE0061FC534B97FB12FD80" box="[1016,1167,490,512]" gridcol="2" gridrow="8" pageId="14" pageNumber="15">8.50 (±0.35)</td>
</tr>
<tr id="11E534E4FFDE0061FF37486EFB12FDA9" box="[156,1167,531,553]" gridrow="9" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF37486EFEAEFDA9" box="[156,307,531,553]" gridcol="0" gridrow="9" pageId="14" pageNumber="15">NASPIT</th>
<td id="52345D98FFDE0061FDE1486EFD7CFDA9" box="[586,737,531,553]" gridcol="1" gridrow="9" pageId="14" pageNumber="15">0.13 (±0.39)</td>
<td id="52345D98FFDE0061FC53486EFB12FDA9" box="[1016,1167,531,553]" gridcol="2" gridrow="9" pageId="14" pageNumber="15">0.00 (±0.00)</td>
</tr>
<tr id="11E534E4FFDE0061FF374840FB12FDD3" box="[156,1167,573,595]" gridrow="10" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF374840FEAEFDD3" box="[156,307,573,595]" gridcol="0" gridrow="10" pageId="14" pageNumber="15">STRIPE</th>
<td id="52345D98FFDE0061FDE14840FD7CFDD3" box="[586,737,573,595]" gridcol="1" gridrow="10" pageId="14" pageNumber="15">0.70 (±0.57)</td>
<td id="52345D98FFDE0061FC534840FB12FDD3" box="[1016,1167,573,595]" gridcol="2" gridrow="10" pageId="14" pageNumber="15">1.00 (±0.71)</td>
</tr>
<tr id="11E534E4FFDE0061FF37481BFB12FDFC" box="[156,1167,614,636]" gridrow="11" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF37481BFEAEFDFC" box="[156,307,614,636]" gridcol="0" gridrow="11" pageId="14" pageNumber="15">BTWSUPOC2</th>
<td id="52345D98FFDE0061FDE1481BFD7CFDFC" box="[586,737,614,636]" gridcol="1" gridrow="11" pageId="14" pageNumber="15">12.75 (±1.33)</td>
<td id="52345D98FFDE0061FC53481BFB12FDFC" box="[1016,1167,614,636]" gridcol="2" gridrow="11" pageId="14" pageNumber="15">13.40 (±3.78)</td>
</tr>
<tr id="11E534E4FFDE0061FF3748EDFB12FD26" box="[156,1167,656,678]" gridrow="12" pageId="14" pageNumber="15">
<th id="52345D98FFDE0061FF3748EDFEAEFD26" box="[156,307,656,678]" gridcol="0" gridrow="12" pageId="14" pageNumber="15">BSCK</th>
<td id="52345D98FFDE0061FDE148EDFD7CFD26" box="[586,737,656,678]" gridcol="1" gridrow="12" pageId="14" pageNumber="15">0.75 (±0.30)</td>
<td id="52345D98FFDE0061FC5348EDFB12FD26" box="[1016,1167,656,678]" gridcol="2" gridrow="12" pageId="14" pageNumber="15">0.50 (±0.35)</td>
</tr>
</table>
</paragraph>
<subSubSection id="E7CF652DFFDEFF90FF6E4898FF77FC0D" pageId="14" pageNumber="15" type="distribution">
<paragraph id="AF6A36A6FFDEFF90FF6E4898FF77FC0D" blockId="14.[151,1436,741,1270]" pageId="14" pageNumber="15">
<emphasis id="9DA1EAB4FFDEFF90FF6E4898FEC3FD7E" bold="true" box="[197,350,741,766]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Distribution.</emphasis>
<taxonomicName id="68D54D25FFDEFF90FEC34898FDCAFD7D" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[360,599,741,765]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rubeus">
<emphasis id="9DA1EAB4FFDEFF90FEC34898FDCAFD7D" box="[360,599,741,765]" italics="true" pageId="14" pageNumber="15">Cryptelytrops rubeus</emphasis>
</taxonomicName>
is currently known from southern
<collectingCountry id="D7C27636FFDEFF90FC5B4898FBD2FD7D" box="[1008,1103,741,765]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
from the vicinity of
<collectingRegion id="6D11F844FFDEFF90FAEC489BFE93FCA1" country="Vietnam" name="Ho Chi Minh City" pageId="14" pageNumber="15">Ho Chi Minh City</collectingRegion>
and the low-elevation hills on the southern and western edges of the Langbian (= Dalat) Plateau in southern
<collectingCountry id="D7C27636FFDEFF90FF554950FEC0FCC5" box="[254,349,813,837]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
and eastern
<collectingCountry id="D7C27636FFDEFF90FE404950FDC3FCC5" box="[491,606,813,837]" name="Cambodia" pageId="14" pageNumber="15">Cambodia</collectingCountry>
(
<figureCitation id="37EE2A23FFDEFF90FDC54950FD2DFCC5" box="[622,688,813,837]" captionStart="FIGURE 1" captionStartId="1.[151,250,1878,1901]" captionTargetBox="[176,1411,773,1840]" captionTargetId="figure-285@1.[173,1417,770,1842]" captionTargetPageId="1" captionText="FIGURE 1. Map of Southeast Asia, showing the distribution of the two new species and Cryptelytrops macrops s.s. Symbols indicate sampling localities used in the genetic analysis to define species. Grey circles: C. macrops s.s; open diamonds: C. cardamomensis sp. nov.; black triangles: C. rubeus sp. nov. The empty circle and triangle denotes localities of specimens that have either been referred to the corresponding species on the basis of photographs, or for which the locality is uncertain and requires confirmation." figureDoi="http://doi.org/10.5281/zenodo.5291115" httpUri="https://zenodo.org/record/5291115/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
). The occurrence of this species in Nam Cat Tien National Park is supported by a photographic record of a redeyed green pitviper (
<figureCitation id="37EE2A23FFDEFF90FCD6492CFC5CFCE9" box="[893,961,849,873]" captionStart="FIGURE 6" captionStartId="11.[151,250,1898,1921]" captionTargetBox="[235,1346,193,1865]" captionTargetId="figure-72@11.[232,1347,193,1869]" captionTargetPageId="11" captionText="FIGURE 6. Cryptelytrops rubeus sp.nov. A. Close-up of head of male (left) and female (right) from vicinity of Ho Chi Minh city, South Vietnam (photos: A Malhotra); B. Close-up of head of male (left) and female (right) from South Vietnam; C. Closeup of tail of male from B; D. Pair from B (male above) (photos: K Udomritthiruj)." figureDoi="http://doi.org/10.5281/zenodo.5291129" httpUri="https://zenodo.org/record/5291129/files/figure.png" pageId="14" pageNumber="15">Fig. 6</figureCitation>
), which is almost certainly referable to
<emphasis id="9DA1EAB4FFDEFF90FA29492CFF77FC0D" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="68D54D25FFDEFF90FA29492CFF7AFC0D" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rubeus">C. rubeus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFDEFF90FF6E49E4FB78FB0A" pageId="14" pageNumber="15" type="biology_ecology">
<paragraph id="AF6A36A6FFDEFF90FF6E49E4FB78FB0A" blockId="14.[151,1436,741,1270]" pageId="14" pageNumber="15">
<emphasis id="9DA1EAB4FFDEFF90FF6E49E4FEB7FC31" bold="true" box="[197,298,921,945]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Ecology.</emphasis>
Specimens from
<collectingCountry id="D7C27636FFDEFF90FE5649E4FDEDFC31" box="[509,624,921,945]" name="Cambodia" pageId="14" pageNumber="15">Cambodia</collectingCountry>
were collected at night (1810
<emphasis id="9DA1EAB4FFDEFF90FC7E49E7FC7EFC31" box="[981,995,922,945]" italics="true" pageId="14" pageNumber="15"></emphasis>
2040 h) on the soil bank, herbaceous plants, bushes, tree roots, and tree branches. All were found within
<quantity id="682D9B43FFDEFF90FC2049C0FC59FC55" box="[907,964,957,981]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="10.0">10 m</quantity>
of swift, rocky streams (often near waterfalls) at an elevation of
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450
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. The species was found in sympatry with
<taxonomicName id="68D54D25FFDEFF90FBA0499CFB79FC79" box="[1035,1252,993,1017]" class="Reptilia" family="Viperidae" genus="Viridovipera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="vogeli">
<emphasis id="9DA1EAB4FFDEFF90FBA0499CFB79FC79" box="[1035,1252,993,1017]" italics="true" pageId="14" pageNumber="15">Viridovipera vogeli</emphasis>
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at this location. FMNH 262720 contained mammal hairs in its hind gut (stomach contents were too digested to allow more accurate identification of prey). The specimen in Nam Cat Tien was found near a small temporary pond, c.
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northeast of the park headquarters. It was first noted at 2050 hrs moving on grass adjacent to the pond towards tangled poolside vegetation, and later found at 2230 hrs coiled in bamboo/shrubby undergrowth a few meters away.
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFDEFF90FF6E4EE8FD51FB75" pageId="14" pageNumber="15" type="etymology">
<paragraph id="AF6A36A6FFDEFF90FF6E4EE8FD51FB75" blockId="14.[151,1436,741,1270]" pageId="14" pageNumber="15">
<emphasis id="9DA1EAB4FFDEFF90FF6E4EE8FED6FB2D" bold="true" box="[197,331,1173,1197]" inLineHeading="true" pageId="14" pageNumber="15" reason="1">Etymology.</emphasis>
Rubeus, from Latin, meaning reddish, refers to the reddish eye colouration in both sexes, the presence of at least a partial red lateral stripe in males, and the bright and prominent red coloration on the tail, which differentiates it from other macrops group species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,558 @@
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<mods:title id="C008C852483293E3B7E8665F656A4949">Two new species of pitviper of the genus Cryptelytrops Cope 1860 (Squamata: Viperidae: Crotalinae) from Southeast Asia</mods:title>
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<taxonomicNameLabel id="869257CFFFDEFF90FE0A4F42FDB3FAD6" box="[417,558,1343,1366]" pageId="14" pageNumber="15" sensu="stricto">sensu stricto</taxonomicNameLabel>
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(
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)
</paragraph>
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<paragraph id="AF6A36A6FFDEFF90FD494F43FB9BFAD6" blockId="14.[151,1030,1342,1402]" box="[738,1030,1342,1366]" pageId="14" pageNumber="15">(Big-eyed Green Pitviper)</paragraph>
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<subSubSection id="E7CF652DFFDEFF90FF3C4F1FFEE0FAFA" box="[151,381,1378,1402]" pageId="14" pageNumber="15" type="description">
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<figureCitation id="37EE2A23FFDEFF90FF3C4F1FFF47FAFA" box="[151,218,1378,1402]" captionStart="FIGURE 7" captionStartId="16.[151,250,1976,1999]" captionTargetBox="[307,1289,201,1941]" captionTargetId="figure-49@16.[283,1298,193,1942]" captionTargetPageId="16" captionText="FIGURE 7. Line drawings of the head scalation of the holotype of Cryptelytrops macrops s. s. (MHNG 1400.85). Labelling as in Fig. 3 (a: 4.15mm, b: 2.30mm, c: 3.00mm, d: 23.2mm)." figureDoi="http://doi.org/10.5281/zenodo.5291131" httpUri="https://zenodo.org/record/5291131/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
,
<figureCitation id="37EE2A23FFDEFF90FF4D4F1FFF6BFAFA" box="[230,246,1378,1402]" captionStart="FIGURE 8" captionStartId="17.[151,250,1829,1852]" captionTargetBox="[239,1355,180,1807]" captionTargetId="figure-125@17.[234,1359,171,1824]" captionTargetPageId="17" captionText="FIGURE 8. Cryptelytrops macrops sensu stricto. A. Juvenile male from Jae Sorn National Park, Lampang Prov., Thailand (photo: A Malhotra); B. Adult male from Sakaerat Experimental Station, Nakhon Ratchasima Prov., Thailand (photo: A Malhotra); C. Adult female from the vicinity of Bangkok, Thailand (photo: A Gumprecht); D. Adult male (left) and female (right) from north of Khao Yai National park, Nakhon Ratchasima Prov., Thailand (photos: A Gumprecht); E. Juvenile male from Nakhon Ratchasima Prov., Thailand (photo: A Gumprecht). F. Cryptelytrops albolabris from southern Thailand, for comparison. Note the more elongated head, relatively smaller eye, yellower eye and lack of any blue pigmentation (photo: A. Malhotra)." figureDoi="http://doi.org/10.5281/zenodo.5291135" httpUri="https://zenodo.org/record/5291135/files/figure.png" pageId="14" pageNumber="15">8</figureCitation>
;
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4
</paragraph>
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<taxonomicName id="68D54D25FFDEFF90FF3C4FDBFD8EFA3B" authority="Kramer 1977" authorityName="Kramer" authorityYear="1977" box="[151,531,1445,1467]" class="Reptilia" family="Viperidae" genus="Trimeresurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="macrops">
<emphasis id="9DA1EAB4FFDEFF90FF3C4FDBFEE3FA3B" box="[151,382,1446,1467]" italics="true" pageId="14" pageNumber="15">Trimeresurus macrops</emphasis>
<bibRefCitation id="CB444B57FFDEFF90FE224FDBFD8EFA3B" author="Kramer, E." box="[393,531,1445,1467]" pageId="14" pageNumber="15" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer 1977</bibRefCitation>
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:
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: 757 (part);
<bibRefCitation id="CB444B57FFDEFF90FCF84FDBFB17FA3B" author="Regenass, U. &amp; Kramer, E." box="[851,1162,1445,1467]" pageId="14" pageNumber="15" pagination="163 - 205" refId="ref12317" refString="Regenass, U. &amp; Kramer, E. (1981) Zur Systematik der grunen Grubenottern der Gattung Trimeresurus Serpentes, Crotalidae. Revue Suisse Zoologie, 88, 163 - 205." type="journal article" year="1981">Regenass and Kramer (1981)</bibRefCitation>
:184 (part);
<bibRefCitation id="CB444B57FFDEFF90FAA14FD8FE93FA59" author="Viravan, C. &amp; Looareesuwan, S. &amp; Kosakarn, W. &amp; Wuthiekanun, V. &amp; McCarthy, C. J. &amp; Stimson, A. F. &amp; Bunnag, D. &amp; Harinasuta, T. &amp; Warrell, D. A." pageId="14" pageNumber="15" pagination="100 - 106" refId="ref12855" refString="Viravan, C., Looareesuwan, S., Kosakarn, W., Wuthiekanun, V., McCarthy, C. J., Stimson, A. F., Bunnag, D., Harinasuta, T. &amp; Warrell, D. A. (1992) A national hospital-based survey of snakes responsible for bites in Thailand. Transactions of the Royal Society of Tropical Medicine and Hygiene, 86, 100 - 106." type="journal article" year="1992">
Viravan
<emphasis id="9DA1EAB4FFDEFF90FACE4FDBFA0BFA3B" box="[1381,1430,1445,1467]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (1992)
</bibRefCitation>
:102;
<bibRefCitation id="CB444B57FFDEFF90FEEE4FBEFD98FA59" author="Gumprecht, A." box="[325,517,1475,1497]" pageId="14" pageNumber="15" pagination="25 - 36" refId="ref11920" refString="Gumprecht, A. (1998) Die Bambusottern der Gattung Trimeresurus Lacepede. Teil II: Die Grossaugen-bambusotter Trimeresurus macrops Kramer, 1977. Sauria, 20, 25 - 36." type="journal article" year="1998">Gumprecht (1998)</bibRefCitation>
: 25 (part);
<bibRefCitation id="CB444B57FFDEFF90FDDD4FBEFCD8FA59" author="Orlov, N. &amp; Ananjeva, N. &amp; Barabanov, A. &amp; Ryabov, S. &amp; Khalikov, R." box="[630,837,1475,1498]" pageId="14" pageNumber="15" pagination="177 - 218" refId="ref12165" refString="Orlov, N., Ananjeva, N., Barabanov, A., Ryabov, S. &amp; Khalikov, R. (2002 a) Diversity of vipers Azemiopinae, Crotalinae in East, Southeast, and South Asia: Annotated checklist and natural history data. Reptilia: Squamata: Serpentes: Viperidae. Faunistische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 23, 177 - 218." type="journal article" year="2002">
Orlov
<emphasis id="9DA1EAB4FFDEFF90FD134FB9FD7BFA5A" box="[696,742,1476,1498]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2002a)
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: 193 (part);
<bibRefCitation id="CB444B57FFDEFF90FC684FBEFB0EFA59" author="Orlov, N. &amp; Ananjeva, N. &amp; Khalikov, R." box="[963,1171,1475,1498]" pageId="14" pageNumber="15" pagination="345 - 360" refId="ref12238" refString="Orlov, N., Ananjeva, N. &amp; Khalikov, R. (2002 b) Natural History of pitvipers in Eastern and Southeastern Asia. In: Schuett, G. W., Hoggren, M., Douglas, M. E. &amp; Greene, H. W. (Eds.), Biology of the vipers. Eagle Mountain Publishing, Eagle Mountain, Utah, pp 345 - 360." type="book chapter" year="2002">
Orlov
<emphasis id="9DA1EAB4FFDEFF90FBAE4FB9FBAFFA5A" box="[1029,1074,1476,1498]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2002b)
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: 353 (part);
<bibRefCitation id="CB444B57FFDEFF90FABA4FBEFEB3FA78" author="Gumprecht, A. &amp; Tillack, F. &amp; Orlov, N. L. &amp; Captain, A. &amp; Ryabov, S." pageId="14" pageNumber="15" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. &amp; Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
Gumprecht
<emphasis id="9DA1EAB4FFDEFF90FA214FB9FF46FA78" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2004)
</bibRefCitation>
: 230 (part),
<bibRefCitation id="CB444B57FFDEFF90FE024F9FFDE9FA78" author="Teynie, A. &amp; David, P. &amp; Ohler, A. &amp; Luanglath, K." box="[425,628,1506,1528]" pageId="14" pageNumber="15" pagination="33 - 62" refId="ref12667" refString="Teynie, A., David, P., Ohler, A. &amp; Luanglath, K. (2004) Notes on a collection of amphibians and reptiles from southern Laos, with a discussion of the occurrence of Indo - Malayan species. Hamadryad, 29, 33 - 62." type="journal article" year="2004">
Teynié
<emphasis id="9DA1EAB4FFDEFF90FE5F4F9EFDBDFA78" box="[500,544,1506,1528]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2004)
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: 47.
</paragraph>
</subSubSection>
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<materialsCitation id="1FBD3CFBFFDEFF90FE3E4C62FBCEF9B7" box="[405,1107,1567,1591]" country="Thailand" location="Bangkok" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="holotype">
MHNG 1400.85,
<typeStatus id="706E8804FFDEFF90FDC24C62FD53F9B7" box="[617,718,1567,1591]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
, adult male,
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,
<collectingCountry id="D7C27636FFDEFF90FC474C62FBCEF9B7" box="[1004,1107,1567,1591]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
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;
<materialsCitation id="1FBD3CFBFFDEFF90FBCD4C62FCFBF9DC" country="Thailand" location="Bangkok" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paratype">
MHNG 1400.73, 1400.78, 1400.79, 1400.87, 1400.96 100,
<typeStatus id="706E8804FFDEFF90FDB94C39FD1DF9DC" box="[530,640,1604,1628]" pageId="14" pageNumber="15" type="paratype">paratypes</typeStatus>
,
<collectingRegion id="6D11F844FFDEFF90FD274C39FD69F9DC" box="[652,756,1604,1628]" country="Thailand" name="Krung Thep Mahanakhon" pageId="14" pageNumber="15">Bangkok</collectingRegion>
,
<collectingCountry id="D7C27636FFDEFF90FD544C39FCFBF9DC" box="[767,870,1604,1628]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFDEFF90FCDA4C39FC79F9FF" collectorName="Dong Hua Sao National Biodiversity" country="Laos" location="Bolaven Plateau" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Champasak" typeStatus="paratype">
FMNH 25895458,
<collectorName id="02205370FFDEFF90FBFD4C39FF64F9FF" pageId="14" pageNumber="15">Dong Hua Sao National Biodiversity</collectorName>
Conservation Area,
<location id="AA0A607DFFDEFF90FE4E4C1AFD02F9FF" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FE4E4C1AFD02F9FF" box="[485,671,1639,1663]" country="Laos" name="Bolaven Plateau" pageId="14" pageNumber="15" stateProvince="Champasak">Bolaven Plateau</location>
,
<collectingRegion id="6D11F844FFDEFF90FD074C1AFC00F9FF" box="[684,925,1639,1663]" country="Laos" name="Champasak" pageId="14" pageNumber="15">Champasak Province</collectingRegion>
,
<collectingCountry id="D7C27636FFDEFF90FC014C15FC79F9FF" box="[938,996,1640,1663]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFDEFF90FC5B4C15FB32F923" country="Laos" location="Dong Khanthung National Biodiversity Conservation Area" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Champasak" typeStatus="paratype">
FMNH 255249, 25525152, 255256 57,
<location id="AA0A607DFFDEFF90FF684CF1FCFAF924" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FF684CF1FCFAF924" box="[195,871,1676,1700]" country="Laos" name="Dong Khanthung National Biodiversity Conservation Area" pageId="14" pageNumber="15" stateProvince="Champasak">Dong Khanthung National Biodiversity Conservation Area</location>
,
<collectingRegion id="6D11F844FFDEFF90FCDE4CF1FBFAF924" box="[885,1127,1676,1700]" country="Laos" name="Champasak" pageId="14" pageNumber="15">Champasak Province</collectingRegion>
,
<collectingCountry id="D7C27636FFDEFF90FBDE4CF1FB32F923" box="[1141,1199,1676,1699]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFDEFF90FB164CF1FB3AF947" country="Laos" location="Khao Khouay National Biodiversity Conservation Area" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Bolikhamxai" typeStatus="paratype">
FMNH 25525355, Phou
<location id="AA0A607DFFDEFF90FF724CD2FCC9F947" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FF724CD2FCC9F947" box="[217,852,1711,1735]" country="Laos" name="Khao Khouay National Biodiversity Conservation Area" pageId="14" pageNumber="15" stateProvince="Bolikhamxai">Khao Khouay National Biodiversity Conservation Area</location>
,
<collectingRegion id="6D11F844FFDEFF90FCCA4CD2FBFDF947" box="[865,1120,1711,1735]" country="Laos" name="Bolikhamxai" pageId="14" pageNumber="15">Bolikhamxai Province</collectingRegion>
,
<collectingCountry id="D7C27636FFDEFF90FBC64CCDFB3AF947" box="[1133,1191,1712,1735]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFDEFF90FB1E4CCDFBA1F96B" country="Laos" location="Nam No National Biodiversity Conservation Area" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
FMNH 255248, Hin
<location id="AA0A607DFFDEFF90FF3C4CA9FD7DF96C" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FF3C4CA9FD7DF96C" box="[151,736,1748,1772]" country="Laos" name="Nam No National Biodiversity Conservation Area" pageId="14" pageNumber="15" stateProvince="Khammouan">Nam No National Biodiversity Conservation Area</location>
,
<collectingRegion id="6D11F844FFDEFF90FD5B4CA9FC6FF96C" box="[752,1010,1748,1772]" country="Laos" name="Khammouan" pageId="14" pageNumber="15">Khammouan Province</collectingRegion>
,
<collectingCountry id="D7C27636FFDEFF90FBA94CA9FBA1F96B" box="[1026,1084,1748,1771]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFDEFF90FBE04CA9FC24F88F" country="Cambodia" location="Virachey National Park" municipality="Siem Pang District" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Stung Treng" typeStatus="paratype">
FMNH 26271516,
<location id="AA0A607DFFDEFF90FA9C4CA9FEA5F88F" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FA9C4CA9FEA5F88F" country="Cambodia" municipality="Siem Pang District" name="Virachey National Park" pageId="14" pageNumber="15" stateProvince="Stung Treng">Virachey National Park</location>
,
<collectingMunicipality id="4F0EACDCFFDEFF90FEEC4C8AFDB8F88F" box="[327,549,1783,1808]" pageId="14" pageNumber="15">Siem Pang District</collectingMunicipality>
,
<collectingRegion id="6D11F844FFDEFF90FD9F4C8AFCAFF88F" box="[564,818,1783,1808]" country="Cambodia" name="Stung Treng" pageId="14" pageNumber="15">Stung Treng Province</collectingRegion>
,
<collectingCountry id="D7C27636FFDEFF90FCE94C8AFC24F88F" box="[834,953,1783,1807]" name="Cambodia" pageId="14" pageNumber="15">Cambodia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFDEFF90FC6C4C85FD16F8B4" accessionNumber="NMW 23899" country="Thailand" location="Dong Rek Mountains" pageId="14" pageNumber="15" specimenCount="1" typeStatus="paratype">
NMW 23899:4, NMW 23899:10, NMW 23899:13,
<location id="AA0A607DFFDEFF90FEBF4D61FD8EF8B4" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FEBF4D61FD8EF8B4" box="[276,531,1820,1844]" country="Thailand" name="Dong Rek Mountains" pageId="14" pageNumber="15">Dong Rek Mountains</location>
,
<collectingCountry id="D7C27636FFDEFF90FD8F4D61FD16F8B4" box="[548,651,1820,1844]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
</materialsCitation>
;
<materialsCitation id="1FBD3CFBFFDEFF90FD304D61FB15F8D7" accessionNumber="NMW 23897" country="Thailand" location="Don Pia Fei Mountains" pageId="14" pageNumber="15" specimenCount="1" typeStatus="paratype">
NMW 23897:1, NMW 23897:3, NMW 23897:4, NMW 23897:7, NMW 23897:13, NMW 23897:15, NMW 23897:17,
<location id="AA0A607DFFDEFF90FCAB4D3DFB8EF8D7" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FCAB4D3DFB8EF8D7" box="[768,1043,1855,1879]" country="Thailand" name="Don Pia Fei Mountains" pageId="14" pageNumber="15">Don Pia Fei Mountains</location>
,
<collectingCountry id="D7C27636FFDEFF90FB8A4D42FB15F8D7" box="[1057,1160,1855,1879]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
</materialsCitation>
.
<collectorName id="02205370FFDEFF90FB3C4D42FB5FF8D7" box="[1175,1218,1855,1879]" pageId="14" pageNumber="15">The</collectorName>
senior author also examined a number of live
<collectorName id="02205370FFDEFF90FE724D19FD91F8FC" box="[473,524,1892,1916]" pageId="14" pageNumber="15">Thai</collectorName>
specimens under anaesthesia; these included a juvenile male found behind the headquarters of
<location id="AA0A607DFFDEFF90FEE64DF6FDCEF822" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FEE64DF6FDCEF822" box="[333,595,1930,1954]" country="Thailand" name="Jae Sorn National Park" pageId="14" pageNumber="15" stateProvince="Lampang">Jae Sorn National Park</location>
,
<collectingRegion id="6D11F844FFDEFF90FDF44DF6FCA8F822" box="[607,821,1930,1955]" country="Thailand" name="Lampang" pageId="14" pageNumber="15">Lampang province</collectingRegion>
, on
<date id="DB6B1066FFDEFF90FCCE4DF7FB98F822" box="[869,1029,1927,1954]" pageId="14" pageNumber="15" value="1996-07-24">
<collectingDate id="CB2FE98EFFDEFF90FCCE4DF7FB98F822" box="[869,1029,1927,1954]" pageId="14" pageNumber="15" value="1996-07-24">
24
<superScript id="58A09BEEFFDEFF90FC2B4DFAFC11F815" attach="left" box="[896,908,1927,1941]" fontSize="6" pageId="14" pageNumber="15">th</superScript>
July 1996
</collectingDate>
</date>
,
<specimenCount id="B9D3FD2FFFDEFF90FBBA4DF6FB25F822" box="[1041,1208,1930,1954]" pageId="14" pageNumber="15" type="generic">two specimens</specimenCount>
from the vicinity of
<location id="AA0A607DFFDEFF90FF3C4DD2FE71F847" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFDEFF90FF3C4DD2FE71F847" box="[151,492,1967,1991]" country="Thailand" name="Sakaerat Experimental Station" pageId="14" pageNumber="15" stateProvince="Nakhon Ratchasima">Sakaerat Experimental Station</location>
,
<collectingRegion id="6D11F844FFDEFF90FE5C4DD2FCDFF847" box="[503,834,1967,1991]" country="Thailand" name="Nakhon Ratchasima" pageId="14" pageNumber="15">Nakhon Ratchasima Province</collectingRegion>
in
<date id="DB6B1066FFDEFF90FCCE4DD2FC47F847" box="[869,986,1967,1991]" pageId="14" pageNumber="15" value="1999-06">June 1999</date>
, and
<specimenCount id="B9D3FD2FFFDEFF90FBB84DD2FB27F847" box="[1043,1210,1967,1991]" pageId="14" pageNumber="15" type="generic">five specimens</specimenCount>
obtained from dealers through the Queen Savoabha Memorial Institute (QSMI),
<collectingRegion id="6D11F844FFDEFF90FCE84DAFFC35F86A" box="[835,936,2002,2026]" country="Thailand" name="Krung Thep Mahanakhon" pageId="14" pageNumber="15">Bangkok</collectingRegion>
from
<collectingRegion id="6D11F844FFDEFF90FC474DAFFBF6F86A" box="[1004,1131,2002,2026]" country="Thailand" name="Nonthaburi" pageId="14" pageNumber="15">Nonthaburi</collectingRegion>
and
<collectingRegion id="6D11F844FFDEFF90FB0B4DAFFABDF86A" box="[1184,1312,2002,2026]" country="Thailand" name="Ang Thong" pageId="14" pageNumber="15">Ang Thong</collectingRegion>
Provinces, as well as the outskirts of the city itself, in 1998. Although morphological data from these specimens have been used in all analyses, under the conditions of permission, these specimens could not be taken as vouchers and were returned unharmed to the site of capture, or retained by the QSMI for further study.
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFDFFF91FF6E4B7EFAEBFDBC" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="AF6A36A6FFDFFF91FF6E4B7EFAEBFDBC" blockId="15.[151,1436,151,1148]" pageId="15" pageNumber="16">
<emphasis id="9DA1EAB4FFDFFF91FF6E4B7EFE03FE9B" bold="true" box="[197,414,259,284]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Revised diagnosis.</emphasis>
<emphasis id="9DA1EAB4FFDFFF91FE0F4B79FD51FE9C" box="[420,716,260,284]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="68D54D25FFDFFF91FE0F4B79FD3EFE9C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[420,675,260,284]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" sensu="stricto" species="macrops">Cryptelytrops macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFDFFF91FD024B78FD51FE9C" box="[681,716,261,284]" pageId="15" pageNumber="16" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
is distinguished from all other Asian pitviper species which also have the typical &quot;green pitviper&quot; colouration (uniform green dorsal colour and a lateral stripe present on the first few dorsal scale rows in one or both sexes), except other species of
<taxonomicName id="68D54D25FFDFFF91FC334B31FBACFEE4" authorityName="Cope" authorityYear="1860" box="[920,1073,332,356]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="9DA1EAB4FFDFFF91FC334B31FBACFEE4" box="[920,1073,332,356]" italics="true" pageId="15" pageNumber="16">Cryptelytrops</emphasis>
</taxonomicName>
, by the presence of a fused first supralabial and nasal scale. It can be distinguished from
<emphasis id="9DA1EAB4FFDFFF91FCA64B0DFBADFE08" box="[781,1072,368,392]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="68D54D25FFDFFF91FCA64B0DFC3EFE08" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[781,931,368,392]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
,
<taxonomicName id="68D54D25FFDFFF91FC054B0DFBADFE08" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[942,1072,368,392]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
</emphasis>
and
<taxonomicName id="68D54D25FFDFFF91FBCD4B0DFAB0FE08" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[1126,1325,368,392]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="9DA1EAB4FFDFFF91FBCD4B0DFAB0FE08" box="[1126,1325,368,392]" italics="true" pageId="15" pageNumber="16">C. septentrionalis</emphasis>
</taxonomicName>
primarily by the relatively larger size of the eye (most obvious in adults), the relatively wider supraoculars, and the shape of the head, which is more elongate
<emphasis id="9DA1EAB4FFDFFF91FDB54BC4FDB1FE50" box="[542,556,441,464]" italics="true" pageId="15" pageNumber="16"></emphasis>
oval in
<emphasis id="9DA1EAB4FFDFFF91FD224BC5FC26FE50" box="[649,955,440,464]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="68D54D25FFDFFF91FD224BC5FCB9FE50" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[649,804,440,464]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="albolabris">C. albolabris</taxonomicName>
,
<taxonomicName id="68D54D25FFDFFF91FC9F4BC5FC26FE50" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[820,955,440,464]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="insularis">C. insularis</taxonomicName>
</emphasis>
and
<emphasis id="9DA1EAB4FFDFFF91FC504BC5FB53FE50" box="[1019,1230,440,464]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="68D54D25FFDFFF91FC504BC5FB57FE50" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[1019,1226,440,464]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="septentrionalis">C. septentrionalis</taxonomicName>
,
</emphasis>
but widens quite abruptly behind the eyes in
<taxonomicName id="68D54D25FFDFFF91FE654BA1FD03FE74" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[462,670,476,500]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
<emphasis id="9DA1EAB4FFDFFF91FE654BA1FD03FE74" box="[462,670,476,500]" italics="true" pageId="15" pageNumber="16">C. cardamomensis</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFDFFF91FD0E4BA1FD62FE74" bold="true" box="[677,767,476,500]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="869257CFFFDFFF91FD0E4BA1FD62FE74" box="[677,767,476,500]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="9DA1EAB4FFDFFF91FCA64BA1FC25FE74" box="[781,952,476,500]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="68D54D25FFDFFF91FCA64BA1FC10FE74" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[781,909,476,500]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFDFFF91FC3F4BA0FC25FE74" box="[916,952,477,500]" pageId="15" pageNumber="16" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="68D54D25FFDFFF91FC444BA1FBC1FE74" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[1007,1116,476,500]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
<emphasis id="9DA1EAB4FFDFFF91FC444BA1FBC1FE74" box="[1007,1116,476,500]" italics="true" pageId="15" pageNumber="16">C. rubeus</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFDFFF91FBCF4BA1FB20FE74" bold="true" box="[1124,1213,476,500]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="869257CFFFDFFF91FBCF4BA1FB20FE74" box="[1124,1213,476,500]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
to give a characteristically triangular shaped head.
<emphasis id="9DA1EAB4FFDFFF91FDAD487DFCAFFD98" box="[518,818,512,536]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="68D54D25FFDFFF91FDAD487DFC9BFD98" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[518,774,512,536]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" sensu="stricto" species="macrops">Cryptelytrops macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFDFFF91FCA4487CFCAFFD98" box="[783,818,513,536]" pageId="15" pageNumber="16" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
can be distinguished from both
<taxonomicName id="68D54D25FFDFFF91FB09487DFAEEFD98" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[1186,1395,512,536]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="cardamomensis" status="sp. nov.">
<emphasis id="9DA1EAB4FFDFFF91FB09487DFAEEFD98" box="[1186,1395,512,536]" italics="true" pageId="15" pageNumber="16">C. cardamomensis</emphasis>
</taxonomicName>
<emphasis id="9DA1EAB4FFDFFF91FAD1487DFF55FDBC" bold="true" pageId="15" pageNumber="16">
<taxonomicNameLabel id="869257CFFFDFFF91FAD1487DFF55FDBC" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="68D54D25FFDFFF91FF554859FEF6FDBC" authority="Malhotra &amp; Thorpe &amp; Stuart, 2011" authorityName="Malhotra &amp; Thorpe &amp; Stuart" authorityYear="2011" box="[254,363,548,572]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="rubeus" status="sp. nov.">
<emphasis id="9DA1EAB4FFDFFF91FF554859FEF6FDBC" box="[254,363,548,572]" italics="true" pageId="15" pageNumber="16">C. rubeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFDFFF91FED94859FE5EFDBB" box="[370,451,548,571]" pageId="15" pageNumber="16" rank="species">sp. nov</taxonomicNameLabel>
using the characters detailed above (further details are also given in
<tableCitation id="E257031DFFDFFF91FB6E4859FABFFDBC" box="[1221,1314,548,572]" captionStart="TABLE 2" captionStartId="9.[151,234,358,381]" captionTargetPageId="9" captionText="TABLE 2. Mean values and standard deviations of morphological characters that in combination can be used to distinguish C. cardamomensis sp. nov. from both C. macrops s.s. and C. rubeus sp. nov. Observed values have a 95% probability of falling within ± 1.96 x SD of the mean, hence this is a more useful measure than the range. LHEAD is adjusted to the grand mean size of SVL (48.72 cm) in females and DEYE to the grand mean size of LHEAD (22.45, 25.45 mm) in males and females respectively. Cm = C. macrops s.s., Cc = C. cardamomensis, Cr = C. rubeus. Character abbreviations are explained in Appendix 1. Entries in bold identify the taxa which that character helps to discriminate from C. cardamomensis (e.g. BTWSUPOC can be used to discriminate both C. macrops and C. rubeus from C. cardamomensis while DV19to17 is only useful to discriminate C. rubeus from C. cardamomensis)." pageId="15" pageNumber="16">Tables 2</tableCitation>
and
<tableCitation id="E257031DFFDFFF91FAF24859FAF4FDBC" box="[1369,1385,548,572]" captionStart="TABLE 3" captionStartId="13.[151,234,1575,1598]" captionTargetBox="[156,1155,1675,2028]" captionTargetPageId="13" captionText="TABLE 3. Mean and standard deviations of morphological characters that in combination can be used to distinguish C. macrops s.s. and C. rubeus sp. nov. Data are for the maximum number of specimens available for that character. Cm = C. macrops s.s., Cr = C. rubeus. Character abbreviations are explained in Appendix 1." pageId="15" pageNumber="16">3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFDFFF8DFF6C483AFF74FEBF" lastPageId="19" lastPageNumber="20" pageId="15" pageNumber="16" type="description">
<paragraph id="AF6A36A6FFDFFF91FF6C483AFEF4FBFC" blockId="15.[151,1436,151,1148]" pageId="15" pageNumber="16">
<emphasis id="9DA1EAB4FFDFFF91FF6C483AFD98FDE0" bold="true" box="[199,517,583,608]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">
Redescription of
<typeStatus id="706E8804FFDFFF91FE3C4835FE60FDE0" box="[407,509,584,608]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
.
</emphasis>
We supplement
<bibRefCitation id="CB444B57FFDFFF91FD6C483AFC19FDDF" author="Kramer, E." box="[711,900,583,607]" pageId="15" pageNumber="16" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramers (1977)</bibRefCitation>
original description (Appendix 2) of the
<typeStatus id="706E8804FFDFFF91FACA483AFF57FD04" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, as follows. Body cylindrical, head triangular in dorsal aspect and very distinct from neck. Canthus rostralis distinct. Head scales small (no large dorsal shields), smooth except for weak tubercular keeling in temporal region and region between suprabials and temporal region and strong keeling on rear of head. Body scales also strongly keeled dorsally. 168 ventral scales, 68 pairs of subcaudals, 21 dorsal scales at mid
<emphasis id="9DA1EAB4FFDFFF91FB9648A4FBD6FD70" box="[1085,1099,729,752]" italics="true" pageId="15" pageNumber="16"></emphasis>
body. Rostral scale somewhat triangular, upper edge more than half the width of lower edge (ratio between the lengths of the upper to lower edge 0.58). Pupil vertically elliptical. Loreal pit present. Nostril completely enclosed in nasal scale. Nasal scale partially fused with first supralabial, partial suture present behind the nostril, but not to anterior on one side, on both sides of the nostril on the other. Ten supralabials, 13 sublabials, two postocular scales. Shield bordering anterior edge of pit fused with second supralabial, in contact with nasal scale. Subocular scale touches third supralabial, but separated by one scale from fourth and fifth supralabials. At least 7 scales between supraoculars, 13 scales between their posterior edges. Snout
<emphasis id="9DA1EAB4FFDFFF91FEC649A8FEE6FC6C" box="[365,379,981,1004]" italics="true" pageId="15" pageNumber="16"></emphasis>
vent length
<quantity id="682D9B43FFDFFF91FDA949A9FDFFFC6B" box="[514,610,980,1004]" metricMagnitude="-1" metricUnit="m" metricValue="5.06" pageId="15" pageNumber="16" unit="cm" value="50.6">50.6 cm</quantity>
; tail length
<quantity id="682D9B43FFDFFF91FD4149A9FCD7FC6B" box="[746,842,980,1004]" metricMagnitude="-1" metricUnit="m" metricValue="1.33" pageId="15" pageNumber="16" unit="cm" value="13.3">13.3 cm</quantity>
. Head length measured from tip of snout to posterior edge of lower jawbone
<quantity id="682D9B43FFDFFF91FE3D498AFE60FB8F" box="[406,509,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="15" pageNumber="16" unit="mm" value="23.2">23.2 mm</quantity>
; head width measured between rear outer edges of supraoculars
<quantity id="682D9B43FFDFFF91FB65498AFADCFB8F" box="[1230,1345,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.155" pageId="15" pageNumber="16" unit="mm" value="11.55">11.55 mm</quantity>
, at widest point of head
<quantity id="682D9B43FFDFFF91FEF04E61FE5FFBB3" box="[347,450,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="15" pageNumber="16" unit="mm" value="16.9">16.9 mm</quantity>
. Supraocular scale (on right)
<quantity id="682D9B43FFDFFF91FCBB4E61FCE8FBB3" box="[784,885,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" pageId="15" pageNumber="16" unit="mm" value="5.05">5.05 mm</quantity>
long,
<quantity id="682D9B43FFDFFF91FC164E61FB8EFBB3" box="[957,1043,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="16" unit="mm" value="2.3">2.3 mm</quantity>
wide. Distance from eye to nostril
<quantity id="682D9B43FFDFFF91FF3C4E3DFF60FBD7" box="[151,253,1088,1112]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" pageId="15" pageNumber="16" unit="mm" value="4.95">4.95 mm</quantity>
, eye to pit
<quantity id="682D9B43FFDFFF91FED24E3DFE4CFBD7" box="[377,465,1088,1112]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="1.1">1.1 mm</quantity>
, posterior edge of pit to anterior edge of nostril 5.0 mm. Eye diameter
<quantity id="682D9B43FFDFFF91FB464E3DFAC9FBD7" box="[1261,1364,1088,1112]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" pageId="15" pageNumber="16" unit="mm" value="4.15">4.15 mm</quantity>
. Scale reduction formula:
</paragraph>
<paragraph id="AF6A36A6FFDFFF91FF3C4F77FBE2FA0E" blockId="15.[151,1436,1290,2035]" pageId="15" pageNumber="16">
<emphasis id="9DA1EAB4FFDFFF91FF3C4F77FDA9FAA2" bold="true" box="[151,564,1290,1315]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">
Colour of
<typeStatus id="706E8804FFDFFF91FEA44F76FEE8FAA3" box="[271,373,1291,1315]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dorsal surface of head and body brownish grey; sublabials, last three supralabials, and lateral edges of ventral scales steely blue. Apart from mental, sublabials and a few scale rows below sublabials posterior to eighth sublabial, scales on the underside of the head creamy white. Central part and rear edge of ventral scales same creamy white colour. Postocular stripe and lateral stripe not apparent.
</paragraph>
<paragraph id="AF6A36A6FFDFFF91FF3C4FE7FBF0F872" blockId="15.[151,1436,1290,2035]" pageId="15" pageNumber="16">
<emphasis id="9DA1EAB4FFDFFF91FF3C4FE7FEA0FA32" bold="true" box="[151,317,1434,1459]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Colour in life.</emphasis>
This description is based on colour plates on p. 230
<emphasis id="9DA1EAB4FFDFFF91FC214FE6FC05FA32" box="[906,920,1435,1458]" italics="true" pageId="15" pageNumber="16"></emphasis>
235 (with the exception of those listed above) of
<bibRefCitation id="CB444B57FFDFFF91FF1D4FC3FE55FA56" author="Gumprecht, A. &amp; Tillack, F. &amp; Orlov, N. L. &amp; Captain, A. &amp; Ryabov, S." box="[182,456,1470,1494]" pageId="15" pageNumber="16" refId="ref11953" refString="Gumprecht, A., Tillack, F., Orlov, N. L., Captain, A. &amp; Ryabov, S. (2004) Asian Pitvipers. GeitjeBooks, Berlin, 368 pp." type="book" year="2004">
Gumprecht
<emphasis id="9DA1EAB4FFDFFF91FE974FC2FEF3FA56" box="[316,366,1470,1494]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. (2004)
</bibRefCitation>
of specimens from
<collectingRegion id="6D11F844FFDFFF91FD004FC2FC8AFA57" box="[683,791,1471,1495]" country="Thailand" name="Lampang" pageId="15" pageNumber="16">Lampang</collectingRegion>
,
<collectingRegion id="6D11F844FFDFFF91FC8F4FC3FB95FA56" box="[804,1032,1470,1494]" country="Thailand" name="Nakhon Ratchasima" pageId="15" pageNumber="16">Nakhon Ratchasima</collectingRegion>
(Plate 3), and the vicinity of
<collectingRegion id="6D11F844FFDFFF91FAFC4FC2FF59FA7A" country="Thailand" name="Krung Thep Mahanakhon" pageId="15" pageNumber="16">Bangkok</collectingRegion>
,
<collectingCountry id="D7C27636FFDFFF91FF654F9FFEA9FA7A" box="[206,308,1506,1530]" name="Thailand" pageId="15" pageNumber="16">Thailand</collectingCountry>
, as well as macro photographs of the dorsal, lateral, and ventral aspects of the head and body of specimens from
<collectingRegion id="6D11F844FFDFFF91FE8F4C7BFD91F99E" box="[292,524,1542,1566]" country="Thailand" name="Nakhon Ratchasima" pageId="15" pageNumber="16">Nakhon Ratchasima</collectingRegion>
,
<collectingRegion id="6D11F844FFDFFF91FDB14C7AFD1BF99F" box="[538,646,1543,1567]" country="Thailand" name="Lampang" pageId="15" pageNumber="16">Lampang</collectingRegion>
,
<collectingRegion id="6D11F844FFDFFF91FD3E4C7BFC89F99E" box="[661,788,1542,1566]" country="Thailand" name="Nonthaburi" pageId="15" pageNumber="16">Nonthaburi</collectingRegion>
and
<collectingRegion id="6D11F844FFDFFF91FCFB4C7BFC4EF99E" box="[848,979,1542,1566]" country="Thailand" name="Ang Thong" pageId="15" pageNumber="16">Ang Thong</collectingRegion>
provinces of
<collectingCountry id="D7C27636FFDFFF91FBDE4C7BFB45F99E" box="[1141,1240,1542,1566]" name="Thailand" pageId="15" pageNumber="16">Thailand</collectingCountry>
by AM. Ground colour varable, bluish green, grass green or yellow
<emphasis id="9DA1EAB4FFDFFF91FD7F4C56FD7FF9C2" box="[724,738,1579,1602]" italics="true" pageId="15" pageNumber="16"></emphasis>
green, ventral surface slightly paler, often with bluer hue than dorsal surface especially on anterior of body and excepting lateral edges of ventral scales. Prominent lateral white stripe in males (sometimes encroaching onto second dorsal scale row), usually extending onto head as postocular stripe passing below the eye and fading out on lower preocular, below pit. In some specimens, lateral stripe pale blue rather than white. Scales below postocular stripe lighter in colour than rest of head. Females lack obvious postocular stripe, but may have indistinct pale or blue streak. Usually little distinction between colour of supralabials and upper surface of head. Females occasionally have lateral stripes, although covering less than one
<emphasis id="9DA1EAB4FFDFFF91FAAD4D7EFA89F89A" box="[1286,1300,1795,1818]" italics="true" pageId="15" pageNumber="16"></emphasis>
third of first scale row at mid
<emphasis id="9DA1EAB4FFDFFF91FEFF4D5AFEFFF8BE" box="[340,354,1831,1854]" italics="true" pageId="15" pageNumber="16"></emphasis>
body and not extending onto second scale row. Tail dull brick red, dorsally extending as far as vent, with clear margin on lateral side of tail. Eye golden yellow to light orange in both sexes. Males may have small whitish flecks mid
<emphasis id="9DA1EAB4FFDFFF91FE034D12FE2BF806" box="[424,438,1903,1926]" italics="true" pageId="15" pageNumber="16"></emphasis>
dorsally, more prominent in juveniles, sometimes retained in adults. Interstitial skin blue
<emphasis id="9DA1EAB4FFDFFF91FA254D12FA01F806" box="[1422,1436,1903,1926]" italics="true" pageId="15" pageNumber="16"></emphasis>
grey, with black banding. Sublabial scales often suffused with pale blue; rest of the scales on the underside of the head (genials, chin shields, and scales between these and sublabials) are white. Scattered blue pigment on some dorsal scales on head may be present, but not as well developed as in other two species.
</paragraph>
<caption id="FBAA662EFFC0FF8EFF3C4DC5FD79F86C" ID-DOI="http://doi.org/10.5281/zenodo.5291131" ID-Zenodo-Dep="5291131" httpUri="https://zenodo.org/record/5291131/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1976,1999]" targetBox="[307,1289,201,1941]" targetPageId="16">
<paragraph id="AF6A36A6FFC0FF8EFF3C4DC5FD79F86C" blockId="16.[151,1436,1976,2028]" pageId="16" pageNumber="17">
<emphasis id="9DA1EAB4FFC0FF8EFF3C4DC5FE91F84D" bold="true" box="[151,268,1976,1999]" pageId="16" pageNumber="17">FIGURE 7</emphasis>
. Line drawings of the head scalation of the holotype of
<emphasis id="9DA1EAB4FFC0FF8EFCE94DC5FBCBF84E" box="[834,1110,1976,1998]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="68D54D25FFC0FF8EFCE94DC5FBB4F84E" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[834,1065,1976,1998]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" sensu="stricto" species="macrops">Cryptelytrops macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFC0FF8EFB844DC4FBCBF84E" box="[1071,1110,1977,1998]" pageId="16" pageNumber="17" sensu="stricto">s. s.</taxonomicNameLabel>
</emphasis>
(MHNG 1400.85). Labelling as in Fig. 3 (a: 4.15mm, b: 2.30mm, c: 3.00mm, d: 23.2mm).
</paragraph>
</caption>
<caption id="FBAA662EFFC1FF8FFF3C4D58FF5CF873" ID-DOI="http://doi.org/10.5281/zenodo.5291135" ID-Zenodo-Dep="5291135" httpUri="https://zenodo.org/record/5291135/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1829,1852]" targetBox="[239,1355,180,1807]" targetPageId="17">
<paragraph id="AF6A36A6FFC1FF8FFF3C4D58FF5CF873" blockId="17.[151,1436,1829,2035]" pageId="17" pageNumber="18">
<emphasis id="9DA1EAB4FFC1FF8FFF3C4D58FE92F8BC" bold="true" box="[151,271,1829,1852]" pageId="17" pageNumber="18">FIGURE 8</emphasis>
.
<taxonomicName id="68D54D25FFC1FF8FFEB54D58FD95F8BB" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[286,520,1829,1851]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFC1FF8FFEB54D58FD95F8BB" box="[286,520,1829,1851]" italics="true" pageId="17" pageNumber="18">Cryptelytrops macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFC1FF8FFDBA4D5BFD09F8BB" box="[529,660,1829,1851]" pageId="17" pageNumber="18" sensu="stricto">sensu stricto</taxonomicNameLabel>
. A. Juvenile male from Jae Sorn National Park, Lampang Prov., Thailand (photo: A Malhotra); B. Adult male from Sakaerat Experimental Station, Nakhon Ratchasima Prov., Thailand (photo: A Malhotra); C. Adult female from the vicinity of Bangkok, Thailand (photo: A Gumprecht); D. Adult male (left) and female (right) from north of Khao Yai National park, Nakhon Ratchasima Prov., Thailand (photos: A Gumprecht); E. Juvenile male from Nakhon Ratchasima Prov., Thailand (photo: A Gumprecht). F.
<taxonomicName id="68D54D25FFC1FF8FFCB74DDDFB88F836" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[796,1045,1952,1974]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="albolabris">
<emphasis id="9DA1EAB4FFC1FF8FFCB74DDDFB88F836" box="[796,1045,1952,1974]" italics="true" pageId="17" pageNumber="18">Cryptelytrops albolabris</emphasis>
</taxonomicName>
from southern Thailand, for comparison. Note the more elongated head, relatively smaller eye, yellower eye and lack of any blue pigmentation (photo: A. Malhotra).
</paragraph>
</caption>
<paragraph id="AF6A36A6FFC3FF8DFF6E4AEAFF74FEBF" blockId="19.[151,1436,151,1076]" pageId="19" pageNumber="20">
<emphasis id="9DA1EAB4FFC3FF8DFF6E4AEAFEA1FF2F" bold="true" box="[197,316,151,176]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Variation.</emphasis>
There is geographic variation between different populations of
<emphasis id="9DA1EAB4FFC3FF8DFB864AE5FB7CFF30" box="[1069,1249,152,176]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="68D54D25FFC3FF8DFB864AE5FB2FFF30" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[1069,1202,152,176]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" sensu="stricto" species="macrops">C. macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFC3FF8DFB164AE4FB7CFF30" box="[1213,1249,153,176]" pageId="19" pageNumber="20" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
, summarised in Table 4. Within each population, sexual dimorphism is not pronounced, and is most obvious in the prominence of postocular and lateral stripes in males compared to females, and the relatively shorter tails and more massive girth of females. Females also reach an overall larger size (maximum recorded
<quantity id="682D9B43FFC3FF8DFC4A4B79FBA1FE9B" box="[993,1084,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="6.29" pageId="19" pageNumber="20" unit="cm" value="62.9">62.9 cm</quantity>
SVL compared to
<quantity id="682D9B43FFC3FF8DFABC4B79FAEEFE9B" box="[1303,1395,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="5.64" pageId="19" pageNumber="20" unit="cm" value="56.4">56.4 cm</quantity>
for males).
</paragraph>
</subSubSection>
<subSubSection id="E7CF652DFFC3FF8DFF6E4B36FE86FBB4" pageId="19" pageNumber="20" type="distribution">
<paragraph id="AF6A36A6FFC3FF8DFF6E4B36FC23FC94" blockId="19.[151,1436,151,1076]" pageId="19" pageNumber="20">
<emphasis id="9DA1EAB4FFC3FF8DFF6E4B36FEC3FEE3" bold="true" box="[197,350,331,356]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Distribution.</emphasis>
<bibRefCitation id="CB444B57FFC3FF8DFECF4B31FD96FEE4" author="Kramer, E." box="[356,523,332,356]" pageId="19" pageNumber="20" pagination="721 - 761" refId="ref11992" refString="Kramer, E. (1977) Zur Sclangenfauna von Nepal. Rev. Suisse Zoologie, 84, 721 - 761." type="journal article" year="1977">Kramer (1977)</bibRefCitation>
reported the species from
<collectingCountry id="D7C27636FFC3FF8DFC984B31FC0BFEE4" box="[819,918,332,356]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
(
<collectingRegion id="6D11F844FFC3FF8DFC0E4B31FB90FEE4" box="[933,1037,332,356]" country="Thailand" name="Krung Thep Mahanakhon" pageId="19" pageNumber="20">Bangkok</collectingRegion>
) and
<collectingCountry id="D7C27636FFC3FF8DFBE04B31FB30FEE4" box="[1099,1197,332,356]" name="Vietnam" pageId="19" pageNumber="20">Vietnam</collectingCountry>
, and it was presumed to occur in the intervening area. However, it is apparent that the species has a wider distribution (
<figureCitation id="37EE2A23FFC3FF8DFB7B4B12FA8CFE07" box="[1232,1297,367,391]" captionStart="FIGURE 1" captionStartId="1.[151,250,1878,1901]" captionTargetBox="[176,1411,773,1840]" captionTargetId="figure-285@1.[173,1417,770,1842]" captionTargetPageId="1" captionText="FIGURE 1. Map of Southeast Asia, showing the distribution of the two new species and Cryptelytrops macrops s.s. Symbols indicate sampling localities used in the genetic analysis to define species. Grey circles: C. macrops s.s; open diamonds: C. cardamomensis sp. nov.; black triangles: C. rubeus sp. nov. The empty circle and triangle denotes localities of specimens that have either been referred to the corresponding species on the basis of photographs, or for which the locality is uncertain and requires confirmation." figureDoi="http://doi.org/10.5281/zenodo.5291115" httpUri="https://zenodo.org/record/5291115/files/figure.png" pageId="19" pageNumber="20">Fig. 1</figureCitation>
).
<bibRefCitation id="CB444B57FFC3FF8DFA824B12FE95FE2C" author="Viravan, C. &amp; Looareesuwan, S. &amp; Kosakarn, W. &amp; Wuthiekanun, V. &amp; McCarthy, C. J. &amp; Stimson, A. F. &amp; Bunnag, D. &amp; Harinasuta, T. &amp; Warrell, D. A." pageId="19" pageNumber="20" pagination="100 - 106" refId="ref12855" refString="Viravan, C., Looareesuwan, S., Kosakarn, W., Wuthiekanun, V., McCarthy, C. J., Stimson, A. F., Bunnag, D., Harinasuta, T. &amp; Warrell, D. A. (1992) A national hospital-based survey of snakes responsible for bites in Thailand. Transactions of the Royal Society of Tropical Medicine and Hygiene, 86, 100 - 106." type="journal article" year="1992">
Viravan
<emphasis id="9DA1EAB4FFC3FF8DFA234B0CFF30FE2C" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. (1992)
</bibRefCitation>
reported it from western, northern and northeastern
<collectingCountry id="D7C27636FFC3FF8DFCC14BE9FC4DFE2C" box="[874,976,404,428]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
, and although the present study has not been able to confirm its presence in western
<collectingCountry id="D7C27636FFC3FF8DFD0E4BCAFC95FE4F" box="[677,776,439,463]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
(unfortunately, the majority of specimens resulting from Viravan
<emphasis id="9DA1EAB4FFC3FF8DFF5D4BA0FEB5FE74" box="[246,296,476,500]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
.s study, now deposited in the Natural History Museum, London, have since lost their labels carrying the locality information), it is very likely to be present there. One adult female at the QSMI was said to have been obtained from southern
<collectingCountry id="D7C27636FFC3FF8DFE024859FD91FDBC" box="[425,524,548,572]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
via a dealer based in Thung Song. Mitochondrial DNA analysis showed that it was highly similar to
<taxonomicName id="68D54D25FFC3FF8DFE014835FDAFFDE0" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[426,562,584,608]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" sensu="stricto" species="macrops">
<emphasis id="9DA1EAB4FFC3FF8DFE014835FDAFFDE0" box="[426,562,584,608]" italics="true" pageId="19" pageNumber="20">C. macrops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="869257CFFFC3FF8DFDEA4834FDF8FDDF" box="[577,613,584,608]" pageId="19" pageNumber="20" sensu="stricto">
<emphasis id="9DA1EAB4FFC3FF8DFDEA4834FDC0FDE0" box="[577,605,585,608]" italics="true" pageId="19" pageNumber="20">s.s</emphasis>
.
</taxonomicNameLabel>
from the
<collectingRegion id="6D11F844FFC3FF8DFD40483AFC3AFDDF" box="[747,935,583,607]" country="Thailand" name="Krung Thep Mahanakhon" pageId="19" pageNumber="20">Bangkok region</collectingRegion>
(Genbank accession numbers
<accessionNumber id="B086AB45FFC3FF8DFAB6483AFA05FDDF" box="[1309,1432,583,607]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF517184" pageId="19" pageNumber="20">AF517184</accessionNumber>
,
<accessionNumber id="B086AB45FFC3FF8DFF3C4811FE8CFD04" box="[151,273,620,644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF517219" pageId="19" pageNumber="20">AF517219</accessionNumber>
,
<accessionNumber id="B086AB45FFC3FF8DFE8B4811FE07FD04" box="[288,410,620,644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF517163" pageId="19" pageNumber="20">AF517163</accessionNumber>
, and
<accessionNumber id="B086AB45FFC3FF8DFE714811FDC5FD04" box="[474,600,620,644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF517176" pageId="19" pageNumber="20">AF517176</accessionNumber>
). Nevertheless, the occurrence of this species in southern
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must remain speculative at present until it can be confirmed by the collection of additional specimens in the field.
<bibRefCitation id="CB444B57FFC3FF8DFAF148F2FEB2FD4C" author="Orlov, N. &amp; Ananjeva, N. &amp; Barabanov, A. &amp; Ryabov, S. &amp; Khalikov, R." pageId="19" pageNumber="20" pagination="177 - 218" refId="ref12165" refString="Orlov, N., Ananjeva, N., Barabanov, A., Ryabov, S. &amp; Khalikov, R. (2002 a) Diversity of vipers Azemiopinae, Crotalinae in East, Southeast, and South Asia: Annotated checklist and natural history data. Reptilia: Squamata: Serpentes: Viperidae. Faunistische Abhandlungen des Staatlichen Museums fur Tierkunde Dresden, 23, 177 - 218." type="journal article" year="2002">
Orlov
<emphasis id="9DA1EAB4FFC3FF8DFF3C48C8FF54FD4C" box="[151,201,692,716]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. (2002a)
</bibRefCitation>
also illustrate a specimen (Fig. 20: Pg 191) of
<taxonomicName id="68D54D25FFC3FF8DFCEE48C9FC58FD4C" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[837,965,692,716]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="macrops">
<emphasis id="9DA1EAB4FFC3FF8DFCEE48C9FC58FD4C" box="[837,965,692,716]" italics="true" pageId="19" pageNumber="20">C. macrops</emphasis>
</taxonomicName>
said to be from southern
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. However, this specimen seems to be a misidentified male
<taxonomicName id="68D54D25FFC3FF8DFD4648A5FC1CFD70" authorityName="Malhotra &amp; Thorpe" authorityYear="2004" box="[749,897,728,752]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="albolabris">
<emphasis id="9DA1EAB4FFC3FF8DFD4648A5FC1CFD70" box="[749,897,728,752]" italics="true" pageId="19" pageNumber="20">C. albolabris</emphasis>
</taxonomicName>
. The species is also present in southern (
<bibRefCitation id="CB444B57FFC3FF8DFAE548AAFE8DFC94" author="Teynie, A. &amp; David, P. &amp; Ohler, A. &amp; Luanglath, K." pageId="19" pageNumber="20" pagination="33 - 62" refId="ref12667" refString="Teynie, A., David, P., Ohler, A. &amp; Luanglath, K. (2004) Notes on a collection of amphibians and reptiles from southern Laos, with a discussion of the occurrence of Indo - Malayan species. Hamadryad, 29, 33 - 62." type="journal article" year="2004">
Teynié
<emphasis id="9DA1EAB4FFC3FF8DFF3C4880FF55FC94" box="[151,200,764,788]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 2004
</bibRefCitation>
) and central
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(see also “Material examined”, this paper).
</paragraph>
<paragraph id="AF6A36A6FFC3FF8DFF6E495DFE86FBB4" blockId="19.[151,1436,151,1076]" pageId="19" pageNumber="20">
<emphasis id="9DA1EAB4FFC3FF8DFF6E495DFEB7FCB7" bold="true" box="[197,298,800,824]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">Ecology.</emphasis>
<emphasis id="9DA1EAB4FFC3FF8DFE9A495DFDC1FCB8" box="[305,604,800,824]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="68D54D25FFC3FF8DFE9A495DFDADFCB8" baseAuthorityName="Kramer" baseAuthorityYear="1977" box="[305,560,800,824]" class="Reptilia" family="Viperidae" genus="Cryptelytrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" sensu="stricto" species="macrops">Cryptelytrops macrops</taxonomicName>
<taxonomicNameLabel id="869257CFFFC3FF8DFD93495CFDC1FCB8" box="[568,604,801,824]" pageId="19" pageNumber="20" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
occupies hilly areas up to c.
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<quantity id="682D9B43FFC3FF8DFC0F4962FC73FCB7" box="[932,1006,799,823]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="m" value="600.0">
<elevation id="24F8D195FFC3FF8DFC0F4962FC73FCB7" box="[932,1006,799,823]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="m" value="600.0">600 m</elevation>
</quantity>
.
</specimenCount>
It has mainly been found in dry evergreen and mixed deciduous forest mixed with bamboo forest, but also in dry dipterocarp/grassland vegetation. Most specimens were found within a couple of metres of the ground, but occasionally up to
<quantity id="682D9B43FFC3FF8DFB05491AFB47FCFF" box="[1198,1242,871,895]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="4.0">
<elevation id="24F8D195FFC3FF8DFB05491AFB47FCFF" box="[1198,1242,871,895]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="4.0">4 m</elevation>
</quantity>
high. Specimens were frequently found in the vicinity of water. Food items found in the digestive tracts of specimens examined included mammals (NMW 23899:4 from the
<location id="AA0A607DFFC3FF8DFD0B49CDFC0FFC47" LSID="urn:lsid:plazi:treatment:277C87B0FFDEFF8DFF3C4F43FE86FBB4:AA0A607DFFC3FF8DFD0B49CDFC0FFC47" box="[672,914,943,968]" country="Thailand" name="Dong Rek mountains" pageId="19" pageNumber="20" stateProvince="Lampang">Dong Rek mountains</location>
and AFS96.5 from
<collectingRegion id="6D11F844FFC3FF8DFBD249CDFACFFC47" box="[1145,1362,943,968]" country="Thailand" name="Lampang" pageId="19" pageNumber="20">Lampang Province</collectingRegion>
,
<collectingCountry id="D7C27636FFC3FF8DFAF449D2FF56FC6C" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
), frogs (FMNH 255252 and 255254 from
<collectingRegion id="6D11F844FFC3FF8DFD0C49A9FCB7FC6C" box="[679,810,980,1004]" country="Laos" name="Champasak" pageId="19" pageNumber="20">Champasak</collectingRegion>
and
<collectingRegion id="6D11F844FFC3FF8DFCCA49A9FC6CFC6C" box="[865,1009,980,1004]" country="Laos" name="Bolikhamxai" pageId="19" pageNumber="20">Bolikhamxai</collectingRegion>
Provinces,
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respectively; FMNH 262715 and 262716 from Stung Treng Province,
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) and reptiles (FMNH 255253 from
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,
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).
</paragraph>
</subSubSection>
</treatment>
</document>