509 lines
88 KiB
XML
509 lines
88 KiB
XML
<document id="6C72A0464810394DFA1C6D6A18C09BC4" ID-DOI="10.5281/zenodo.215146" ID-GBIF-Dataset="d839fd88-c35d-45a1-809c-9ed94c8f2013" ID-ISSN="1175-5326" ID-Zenodo-Dep="215146" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460480022206" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2012" docId="039B87C5FF95DF00FF00E0F52BC53964" docLanguage="en" docName="zt03307p061.pdf" docOrigin="Zootaxa 3307" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Athanopsis gotoi Anker, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="60" masterDocId="FFA2FFBDFF92DF0CFF97E22228563B2B" masterDocTitle="Notes on the Indo-West Pacific shrimp genus Athanopsis Coutière, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae)" masterLastPageNumber="61" masterPageNumber="48" pageNumber="55" updateTime="1698307435540" updateUser="plazi">
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<mods:title id="7EF7B6E2D4A709F08C3F0738FE891677">Notes on the Indo-West Pacific shrimp genus Athanopsis Coutière, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae)</mods:title>
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<mods:roleTerm id="919621C668AAE62FC60314DDEE66AA61">Author</mods:roleTerm>
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<mods:namePart id="F36155E3DBDA39B7CEDF5B130FED12FF">Anker, Arthur</mods:namePart>
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<mods:typeOfResource id="A8C66B493AD88531169B4B56E567A683">text</mods:typeOfResource>
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<mods:title id="70E434E564A5F98FC351226030560321">Zootaxa</mods:title>
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<mods:part id="8DB7CD2DFE91AEBF263858F37056A4E3">
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<mods:date id="971BC65C7827275D63BD6B3407CBF24B">2012</mods:date>
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<mods:detail id="29352792CB31D3B8B83D7DC9BD57C15E" type="volume">
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<mods:number id="966AB1952EB48CA9582D647BF498485D">3307</mods:number>
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<mods:classification id="7C558073DD7C814F6F3D5CA72871071C">journal article</mods:classification>
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<mods:identifier id="42EF3D9C9486CA15946D580CDD2F1086" type="DOI">10.5281/zenodo.215146</mods:identifier>
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<mods:identifier id="C9C8AC4046B5C0BDF68872B7D73A810D" type="GBIF-Dataset">d839fd88-c35d-45a1-809c-9ed94c8f2013</mods:identifier>
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<mods:identifier id="036478AC25C79D46B6F7C51C0F874084" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1895A7E20AED5F432938D44021D8192A" type="Zenodo-Dep">215146</mods:identifier>
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<treatment id="039B87C5FF95DF00FF00E0F52BC53964" ID-DOI="http://doi.org/10.5281/zenodo.6178252" ID-GBIF-Taxon="119517049" ID-Zenodo-Dep="6178252" LSID="urn:lsid:plazi:treatment:039B87C5FF95DF00FF00E0F52BC53964" httpUri="http://treatment.plazi.org/id/039B87C5FF95DF00FF00E0F52BC53964" lastPageId="12" lastPageNumber="60" pageId="7" pageNumber="55">
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<subSubSection id="C3286558FF95DF0BFF00E0F529063839" pageId="7" pageNumber="55" type="nomenclature">
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<paragraph id="8B8D36D3FF95DF0BFF00E0F5299039DA" blockId="7.[151,454,727,786]" box="[151,454,727,753]" pageId="7" pageNumber="55">
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<heading id="D0C581BFFF95DF0BFF00E0F5299039DA" bold="true" box="[151,454,727,753]" fontSize="11" level="1" pageId="7" pageNumber="55" reason="1">
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<emphasis id="B946EAC1FF95DF0BFF00E0F5299039DA" bold="true" box="[151,454,727,753]" pageId="7" pageNumber="55">
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<taxonomicName id="4C324D50FF95DF0BFF00E0F5293739DA" ID-CoL="85MDD" box="[151,353,727,753]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="55" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
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<emphasis id="B946EAC1FF95DF0BFF00E0F5293739DA" bold="true" box="[151,353,727,753]" italics="true" pageId="7" pageNumber="55">Athanopsis gotoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27557BAFF95DF0BFEFEE0F5299039DA" box="[361,454,727,753]" pageId="7" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
<paragraph id="8B8D36D3FF95DF0BFF00E0D829063839" blockId="7.[151,454,727,786]" box="[151,336,762,786]" pageId="7" pageNumber="55">
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<figureCitation id="13092A56FF95DF0BFF00E0D828B23839" box="[151,228,762,786]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Figs. 5</figureCitation>
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,
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<figureCitation id="13092A56FF95DF0BFF66E0D829573839" box="[241,257,762,786]" captionStart="FIGURE 6" captionStartId="9.[151,250,949,972]" captionTargetBox="[188,1367,190,876]" captionTargetId="figure@9.[151,1436,161,928]" captionTargetPageId="9" captionText="FIGURE 6. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, major cheliped, mesial view; B, same, lateral view; C, distal major chela, dorsomesial view; D, coxa and ischium of major cheliped, mesial view; E, minor cheliped, lateral view." httpUri="https://zenodo.org/record/215152/files/figure.png" pageId="7" pageNumber="55">6</figureCitation>
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,
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<figureCitation id="13092A56FF95DF0BFE9AE0D8294D3839" box="[269,283,762,786]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="7" pageNumber="55">7</figureCitation>
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C, D
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</paragraph>
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</subSubSection>
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<subSubSection id="C3286558FF95DF0BFF00E1612ACC3F19" pageId="7" pageNumber="55" type="materials_examined">
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<paragraph id="8B8D36D3FF95DF0BFF00E1612B7A38C1" blockId="7.[151,1437,834,1902]" pageId="7" pageNumber="55">
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||
<emphasis id="B946EAC1FF95DF0BFF00E16129123870" bold="true" box="[151,324,834,859]" pageId="7" pageNumber="55">
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<typeStatus id="54898871FF95DF0BFF00E16128863870" box="[151,208,835,859]" pageId="7" pageNumber="55">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54898871FF95DF0BFED9E16029ED3871" box="[334,443,834,858]" pageId="7" pageNumber="55" type="holotype">Holotype</typeStatus>
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: ovigerous female (cl
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<quantity id="4CCA9B36FF95DF0BFD20E1602B443871" box="[695,786,834,858]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="55" unit="mm" value="5.5">5.5 mm</quantity>
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), MNHN-IU-2011-2010-4101,
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<collectingCountry id="F3257643FF95DF0BFBE3E1612CE13871" box="[1140,1207,835,858]" name="Japan" pageId="7" pageNumber="55">Japan</collectingCountry>
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, Kagoshima Prefecture, Amami-Oshima
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<collectingCountry id="F3257643FF95DF0BFE18E14429803855" box="[399,470,870,894]" name="Iceland" pageId="7" pageNumber="55">Island</collectingCountry>
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, Edateku, 28.17'N, 129.13'E, gravel and coral sand flat exposed at extreme low tide, under rocks, with
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<taxonomicName id="4C324D50FF95DF0BFEF1E1A92AE83888" box="[358,702,907,931]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="7" pageNumber="55" phylum="Echiura" rank="species" species="erythrogrammon">
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<emphasis id="B946EAC1FF95DF0BFEF1E1A92AE83888" box="[358,702,907,931]" italics="true" pageId="7" pageNumber="55">Ochetostoma erythrogrammon</emphasis>
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</taxonomicName>
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, leg. R. Goto,
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<date id="FF8C1013FF95DF0BFCFBE1A82C5E3889" box="[876,1032,906,931]" pageId="7" pageNumber="55" value="2010-05-15">15 May 2010</date>
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||
.
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<typeStatus id="54898871FF95DF0BFB83E1A92C2C3888" box="[1044,1146,907,931]" pageId="7" pageNumber="55" type="paratype">Paratype</typeStatus>
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: ovigerous female (cl
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<quantity id="4CCA9B36FF95DF0BFAEEE1A8289138ED" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="55" unit="mm" value="5.7">5.7 mm</quantity>
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), CBM-ZC 10620,
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<collectingCountry id="F3257643FF95DF0BFE09E18D29B738ED" box="[414,481,943,966]" name="Japan" pageId="7" pageNumber="55">Japan</collectingCountry>
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, Okinawa Prefecture, Okinawa
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<collectingCountry id="F3257643FF95DF0BFCD4E18C2BDC38ED" box="[835,906,942,966]" name="Iceland" pageId="7" pageNumber="55">Island</collectingCountry>
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, Nago, Awa, depth
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<quantity id="4CCA9B36FF95DF0BFBF3E18C2CC638ED" box="[1124,1168,942,966]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="55" unit="m" value="6.0">6 m</quantity>
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, under rocks, with echiuran (Ochetostomatinae), leg.
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<collectingCountry id="F3257643FF95DF0BFE7DE1F129A838C1" box="[490,510,979,1002]" name="Italy" pageId="7" pageNumber="55">Y</collectingCountry>
|
||
. Yamada,
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<date id="FF8C1013FF95DF0BFDE5E1F02B7F38C1" box="[626,809,978,1003]" pageId="7" pageNumber="55" value="2009-08-16">16 August 2009</date>
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||
.
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</paragraph>
|
||
<paragraph id="8B8D36D3FF95DF0BFF50E1D42ACC3F19" blockId="7.[151,1437,834,1902]" pageId="7" pageNumber="55">
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||
<emphasis id="B946EAC1FF95DF0BFF50E1D42A653F24" bold="true" box="[199,563,1014,1039]" pageId="7" pageNumber="55">Additional material examined.</emphasis>
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<specimenCount id="9D34FD5AFF95DF0BFDAEE1D42ACC3F25" box="[569,666,1014,1038]" pageId="7" pageNumber="55" type="female">1 female</specimenCount>
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(cl
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<quantity id="4CCA9B36FF95DF0BFD52E1D42B483F25" box="[709,798,1014,1038]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="7" pageNumber="55" unit="mm" value="4.6">4.6 mm</quantity>
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),
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<collectionCode id="ED23AE16FF95DF0BFCA6E1D52BD33F25" LSID="urn:lsid:biocol.org:col:34992" box="[817,901,1015,1038]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="55">RMNH</collectionCode>
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D49180,
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<collectingCountry id="F3257643FF95DF0BFC64E1D42C353F25" box="[1011,1123,1014,1038]" name="Indonesia" pageId="7" pageNumber="55">Indonesia</collectingCountry>
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, Talaud Islands, Beo
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<collectingCountry id="F3257643FF95DF0BFAC7E1D42DC13F25" box="[1360,1431,1014,1038]" name="Iceland" pageId="7" pageNumber="55">Island</collectingCountry>
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||
, Snellius Expedition, sta.
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<date id="FF8C1013FF95DF0BFE39E6382A053F19" box="[430,595,1050,1074]" pageId="7" pageNumber="55" value="1929-06-14" valueMax="1929-06-21" valueMin="1929-06-14">29, 14-21 June</date>
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1930.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3286558FF95DF07FF50E61C29713A4F" lastPageId="11" lastPageNumber="59" pageId="7" pageNumber="55" type="description">
|
||
<paragraph id="8B8D36D3FF95DF0BFF50E61C2AB43E21" blockId="7.[151,1437,834,1902]" pageId="7" pageNumber="55">
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||
<emphasis id="B946EAC1FF95DF0BFF50E61C290F3F7C" bold="true" box="[199,345,1086,1111]" pageId="7" pageNumber="55">Description.</emphasis>
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Small-sized alpheid shrimp (cl
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<quantity id="4CCA9B36FF95DF0BFD59E61C2B0A3F7D" box="[718,860,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.7" metricValueMin="4.6" pageId="7" pageNumber="55" unit="mm" value="5.15" valueMax="5.7" valueMin="4.6">4.6–5.7 mm</quantity>
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). Carapace glabrous, not setose, markedly domed (
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<figureCitation id="13092A56FF95DF0BFF08E64028B73F51" box="[159,225,1122,1146]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
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C). Rostrum well developed, laterally compressed, straight, as long as broad at base, with shallowly concave lateral margins; tip rounded in lateral view, subacute in dorsal view, overreaching mid-length of second article of antennular peduncle; rostral carina absent (
|
||
<figureCitation id="13092A56FF95DF0BFD17E6882A953FE9" box="[640,707,1194,1218]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
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A, B). Extracorneal teeth relatively small, subtriangular, distally blunt, reaching distinctly beyond anterior margin of cornea (
|
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<figureCitation id="13092A56FF95DF0BFCD5E6EC2BD33FCD" box="[834,901,1230,1254]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
|
||
A, B). Pterygostomial angle slightly protruding anteriorly, rounded; cardiac notch deep (
|
||
<figureCitation id="13092A56FF95DF0BFDCCE6D02AC83E21" box="[603,670,1266,1290]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
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B, C).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF95DF0BFF5EE7342B3C3E95" blockId="7.[151,1437,834,1902]" pageId="7" pageNumber="55">
|
||
Pleura of first to fourth abdominal somites rounded posteroventrally; fifth pleuron with blunt posteroventral angle; sixth somite with large articulated triangular flap. Telson widest at proximal third, distally tapering; dorsal surface with two pairs of small spiniform setae, situated at some distance from lateral margin, at about 0.5 and 0.75 telson length, respectively; posterior margin slightly convex, with two pairs of slender spiniform setae at each lateral angle, mesial about three times as long as lateral (
|
||
<figureCitation id="13092A56FF95DF0BFD62E7842B6E3E95" box="[757,824,1446,1470]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">
|
||
Fig.
|
||
<date id="FF8C1013FF95DF0BFCBEE7842B6E3E95" box="[809,824,1446,1470]" pageId="7" pageNumber="55">5</date>
|
||
</figureCitation>
|
||
I, J).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF95DF0BFF5EE7E92A563D2D" blockId="7.[151,1437,834,1902]" pageId="7" pageNumber="55">
|
||
Eyes completely concealed in dorsal and lateral views; cornea well pigmented, occupying most of terminal portion of eyestalk (
|
||
<figureCitation id="13092A56FF95DF0BFEEFE7CC29EC3D2D" box="[376,442,1518,1542]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF95DF0BFF5EE4302AA53DF5" blockId="7.[151,1437,834,1902]" pageId="7" pageNumber="55">
|
||
Antennular peduncle stout; stylocerite blunt distally, reaching to 0.7–0.8 length of second article (
|
||
<figureCitation id="13092A56FF95DF0BFAA5E4302D213D01" box="[1330,1399,1554,1578]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
|
||
A); ventromesial carina with very strong, anteriorly produced tooth (
|
||
<figureCitation id="13092A56FF95DF0BFCE7E4142BE53D65" box="[880,947,1590,1614]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
|
||
D); second article shorter than wide; lateral flagellum biramous, fused portion composed of three articles; accessory ramus with at least four tufts of aesthetascs (
|
||
<figureCitation id="13092A56FF95DF0BFF08E45C28B23DBD" box="[159,228,1662,1686]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
|
||
B). Antenna with stout basicerite ending in large subacute tooth distoventrally; scaphocerite exceeding end of antennular peduncle, blade ovate, broad; small distolateral tooth exceeding anterior margin of blade; carpocerite about the same length as scaphocerite (
|
||
<figureCitation id="13092A56FF95DF0BFDDBE4E42AD93DF5" box="[588,655,1734,1758]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
|
||
A, B, E).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF95DF0BFF5EE4C82C993C45" blockId="7.[151,1437,834,1902]" pageId="7" pageNumber="55">
|
||
Mouthparts not dissected, typical for genus in external view. Third maxilliped with coxa bearing subacute lateral plate above mastigobranch; antepenultimate article flattened, somewhat broadened distally; penultimate article more than twice as long as wide proximally; ultimate article narrower than penultimate, tapering distally, furnished with transverse rows of thick serrulate setae, tip with at least two stout spiniform setae (
|
||
<figureCitation id="13092A56FF95DF0BFBFAE5742CF93C45" box="[1133,1199,1878,1902]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="7" pageNumber="55">Fig. 5</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
<caption id="DF4D665BFF9ADF04FF00E51E29B73CC7" httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="8" pageNumber="56" targetBox="[171,1398,198,1824]" targetPageId="8">
|
||
<paragraph id="8B8D36D3FF9ADF04FF00E51E29B73CC7" blockId="8.[151,1436,1852,2028]" pageId="8" pageNumber="56">
|
||
<emphasis id="B946EAC1FF9ADF04FF00E51E29423C79" bold="true" box="[151,276,1852,1875]" pageId="8" pageNumber="56">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C324D50FF9ADF04FE8BE51F299E3C78" box="[284,456,1853,1875]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="56" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF9ADF04FE8BE51F299E3C78" box="[284,456,1853,1875]" italics="true" pageId="8" pageNumber="56">Athanopsis gotoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557BAFF9ADF04FE47E51F2A743C79" box="[464,546,1853,1874]" pageId="8" pageNumber="56" rank="species">
|
||
<emphasis id="B946EAC1FF9ADF04FE47E51F2A743C79" bold="true" box="[464,546,1853,1874]" pageId="8" pageNumber="56">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU-2010-4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4D665BFF9BDF05FF00E1972ACD3F23" httpUri="https://zenodo.org/record/215152/files/figure.png" pageId="9" pageNumber="57" targetBox="[188,1367,190,876]" targetPageId="9">
|
||
<paragraph id="8B8D36D3FF9BDF05FF00E1972ACD3F23" blockId="9.[151,1436,949,1032]" pageId="9" pageNumber="57">
|
||
<emphasis id="B946EAC1FF9BDF05FF00E197294238E1" bold="true" box="[151,276,949,972]" pageId="9" pageNumber="57">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C324D50FF9BDF05FE8BE197299E38E0" box="[284,456,949,971]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="57" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF9BDF05FE8BE197299E38E0" box="[284,456,949,971]" italics="true" pageId="9" pageNumber="57">Athanopsis gotoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557BAFF9BDF05FE47E1942A7438E0" box="[464,546,950,971]" pageId="9" pageNumber="57" rank="species">
|
||
<emphasis id="B946EAC1FF9BDF05FE47E1942A7438E0" bold="true" box="[464,546,950,971]" pageId="9" pageNumber="57">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU-2010-4101); A, major cheliped, mesial view; B, same, lateral view; C, distal major chela, dorsomesial view; D, coxa and ischium of major cheliped, mesial view; E, minor cheliped, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D36D3FF9BDF05FF5EE6162B253E8D" blockId="9.[151,1437,1076,2036]" pageId="9" pageNumber="57">
|
||
Chelipeds very asymmetrical in shape and unequal in size; carried folded when not in use (
|
||
<figureCitation id="13092A56FF9BDF05FB51E6162D5D3F67" box="[1222,1291,1076,1100]" captionStart="FIGURE 6" captionStartId="9.[151,250,949,972]" captionTargetBox="[188,1367,190,876]" captionTargetId="figure@9.[151,1436,161,928]" captionTargetPageId="9" captionText="FIGURE 6. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, major cheliped, mesial view; B, same, lateral view; C, distal major chela, dorsomesial view; D, coxa and ischium of major cheliped, mesial view; E, minor cheliped, lateral view." httpUri="https://zenodo.org/record/215152/files/figure.png" pageId="9" pageNumber="57">Fig. 6</figureCitation>
|
||
). Major cheliped greatly enlarged, robust; basis with distodorsal spiniform seta; ischium stout, with one distodorsal spiniform seta, mesial side flattened, with one spiniform seta; merus long, slightly inflated, widening distally, ventrally excavated, ventrolateral and ventromesial margins smooth; carpus short, cup-shaped, with broad blunt lobe ventromesially; chela somewhat swollen, subcylindrical; palm depressed on ventrolateral (flexor) surface, distolateral surface smooth; ventromesial surface with about five small tubercles; fingers about 0.4 palm length, somewhat twisted, curved, not gaping, with a few scarce setae or setal tufts, without dense setal brush; cutting edges of pollex and dactylus serrated, with six-seven rounded teeth becoming smaller distally (
|
||
<figureCitation id="13092A56FF9BDF05FC51E7042C5C3E15" box="[966,1034,1318,1342]" captionStart="FIGURE 6" captionStartId="9.[151,250,949,972]" captionTargetBox="[188,1367,190,876]" captionTargetId="figure@9.[151,1436,161,928]" captionTargetPageId="9" captionText="FIGURE 6. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, major cheliped, mesial view; B, same, lateral view; C, distal major chela, dorsomesial view; D, coxa and ischium of major cheliped, mesial view; E, minor cheliped, lateral view." httpUri="https://zenodo.org/record/215152/files/figure.png" pageId="9" pageNumber="57">Fig. 6</figureCitation>
|
||
A–D). Minor cheliped significantly smaller than major cheliped; ischium unarmed on distodorsal margin; merus about four times as long as wide, slightly depressed ventrally; carpus vase-shaped, smooth; chela simple, slender, with fingers slightly longer than palm, cutting edges unarmed, with row of stiff setae (
|
||
<figureCitation id="13092A56FF9BDF05FD7AE7AC2B793E8D" box="[749,815,1422,1446]" captionStart="FIGURE 6" captionStartId="9.[151,250,949,972]" captionTargetBox="[188,1367,190,876]" captionTargetId="figure@9.[151,1436,161,928]" captionTargetPageId="9" captionText="FIGURE 6. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, major cheliped, mesial view; B, same, lateral view; C, distal major chela, dorsomesial view; D, coxa and ischium of major cheliped, mesial view; E, minor cheliped, lateral view." httpUri="https://zenodo.org/record/215152/files/figure.png" pageId="9" pageNumber="57">Fig. 6</figureCitation>
|
||
D, E).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF9BDF05FF5EE7932A5E3CDF" blockId="9.[151,1437,1076,2036]" pageId="9" pageNumber="57">
|
||
Second pereiopod comparatively small, with ischium slightly wider and much shorter than merus, both unarmed; carpus with five articles, first longer than remaining four articles combined, ratio of carpal articles (proximal to distal) about 7: 1: 1: 1: 2; chela simple, longer than most-distal carpal article, with fingers shorter than palm (
|
||
<figureCitation id="13092A56FF9BDF05FF49E43B29493D1A" box="[222,287,1561,1585]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="9" pageNumber="57">Fig. 5</figureCitation>
|
||
G, H). Third pereiopod relatively slender, with strongly compressed ischium, merus, carpus and propodus; ischium with two spiniform setae on ventrolateral surface and one spiniform seta on distodorsal margin; merus almost four times as long as wide, with small, subacute tooth on distolateral margin, without spiniform setae; carpus more slender than merus, about half-length of merus, with stout spiniform seta on distoventral margin; propodus with five spiniform setae along ventral margin and three spiniform setae adjacent to dactylar base; dactylus about 0.4 length of propodus, simple, conical, gradually curving distally, dorsal margin faintly notched at about 0.5 and 0.7 dactylar length, furnished with some setae (
|
||
<figureCitation id="13092A56FF9BDF05FD43E4CB2B403C2A" box="[724,790,1769,1793]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="9" pageNumber="57">
|
||
Fig.
|
||
<date id="FF8C1013FF9BDF05FC90E4CB2B403C2A" box="[775,790,1769,1793]" pageId="9" pageNumber="57">5</date>
|
||
</figureCitation>
|
||
I). Fourth pereiopod similar to third pereiopod, more slen- der; ischium with two spiniform setae on ventrolateral surface, distodorsal margin unarmed; merus about four times as long as wide, wider than merus of third pereiopod; propodus with four spiniform setae; carpus and dactylus as in third pereiopod (
|
||
<figureCitation id="13092A56FF9BDF05FE2BE5732A573C42" box="[444,513,1873,1897]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="9" pageNumber="57">Fig. 5</figureCitation>
|
||
J). Fifth pereiopod more slender than third or fourth pereiopods; ischium without spiniform seta on ventrolateral surface; merus with small, acute tooth on distolateral margin; carpus with two spiniform setae distoventrally; propodus with at least two small spiniform setae along ventral margin and one pair of spiniform setae adjacent to dactylus; distolateral surface with three rows of thick serrulate setae; dactylus similar to that of third pereiopod (
|
||
<figureCitation id="13092A56FF9BDF05FE36E5FE29B23CDF" box="[417,484,2012,2036]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="9" pageNumber="57">Fig. 5</figureCitation>
|
||
K).
|
||
</paragraph>
|
||
<caption id="DF4D665BFF98DF06FF00E41A2BD43C68" httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="10" pageNumber="58" targetBox="[151,1436,193,1569]" targetPageId="10">
|
||
<paragraph id="8B8D36D3FF98DF06FF00E41A2BD43C68" blockId="10.[151,1436,1592,1859]" pageId="10" pageNumber="58">
|
||
<emphasis id="B946EAC1FF98DF06FF00E41A29453D65" bold="true" box="[151,275,1592,1615]" pageId="10" pageNumber="58">FIGURE 7.</emphasis>
|
||
Colour patterns of five species of
|
||
<taxonomicName id="4C324D50FF98DF06FDE3E41A2BD53D65" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[628,899,1592,1614]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B946EAC1FF98DF06FDE3E41A2AB23D65" box="[628,740,1592,1614]" italics="true" pageId="10" pageNumber="58">Athanopsis</emphasis>
|
||
Coutière, 1897
|
||
</taxonomicName>
|
||
A, B—
|
||
<taxonomicName id="4C324D50FF98DF06FC44E41B2D363D65" authority="Banner & Banner, 1981" authorityName="Banner & Banner" authorityYear="1981" box="[979,1376,1592,1614]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="58" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="B946EAC1FF98DF06FC44E41B2C323D65" box="[979,1124,1592,1614]" italics="true" pageId="10" pageNumber="58">A. brevirostris</emphasis>
|
||
Banner & Banner, 1981
|
||
</taxonomicName>
|
||
, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D—
|
||
<taxonomicName id="4C324D50FF98DF06FF55E45429453DA7" box="[194,275,1654,1676]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="58" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF98DF06FF55E45429453DA7" box="[194,275,1654,1676]" italics="true" pageId="10" pageNumber="58">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557BAFF98DF06FE8CE454293D3DA0" box="[283,363,1654,1675]" pageId="10" pageNumber="58" rank="species">
|
||
<emphasis id="B946EAC1FF98DF06FE8CE454293D3DA0" bold="true" box="[283,363,1654,1675]" pageId="10" pageNumber="58">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU-2010-4101), sitting on its host,
|
||
<taxonomicName id="4C324D50FF98DF06FF00E4B62A8A3D81" authority="Leuckart & Ruppel, 1829" authorityName="Leuckart & Ruppel" authorityYear="1829" box="[151,732,1684,1706]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="10" pageNumber="58" phylum="Echiura" rank="species" species="erythrogrammon">
|
||
<emphasis id="B946EAC1FF98DF06FF00E4B629983D81" box="[151,462,1684,1706]" italics="true" pageId="10" pageNumber="58">Ochetostoma erythrogrammon</emphasis>
|
||
Leuckart & Rüppel, 1829
|
||
</taxonomicName>
|
||
, and dorsolateral view (both
|
||
<emphasis id="B946EAC1FF98DF06FB92E4B72C193D81" box="[1029,1103,1685,1706]" italics="true" pageId="10" pageNumber="58">in vitro</emphasis>
|
||
); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (
|
||
<emphasis id="B946EAC1FF98DF06FD22E4962AA13DE2" box="[693,759,1716,1737]" italics="true" pageId="10" pageNumber="58">in situ</emphasis>
|
||
); E—
|
||
<taxonomicName id="4C324D50FF98DF06FCAFE4962C723DE2" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[824,1060,1715,1737]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="58" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="B946EAC1FF98DF06FCAFE4962BCC3DE2" box="[824,922,1716,1737]" italics="true" pageId="10" pageNumber="58">A. saurus</emphasis>
|
||
Anker, 2011
|
||
</taxonomicName>
|
||
, holotype, male from Lizard Island, Australia (QM W29048), lateral view; F, G—
|
||
<taxonomicName id="4C324D50FF98DF06FDE3E4F02B953DCC" authority="Berggren, 1991" authorityName="Berggren" authorityYear="1991" box="[628,963,1746,1768]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="58" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="B946EAC1FF98DF06FDE3E4F02B4F3DC3" box="[628,793,1746,1768]" italics="true" pageId="10" pageNumber="58">A. rubricinctuta</emphasis>
|
||
Berggren, 1991
|
||
</taxonomicName>
|
||
, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D39853), lateral (F) and dorsal (G) view in aquarium; H—
|
||
<taxonomicName id="4C324D50FF98DF06FB89E4D32DCE3C2D" authority="Banner & Banner, 1982" authorityName="Banner & Banner" authorityYear="1982" box="[1054,1432,1776,1798]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="58" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B946EAC1FF98DF06FB89E4D32CC03C2D" box="[1054,1174,1776,1798]" italics="true" pageId="10" pageNumber="58">A. australis</emphasis>
|
||
Banner & Banner, 1982
|
||
</taxonomicName>
|
||
; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8D36D3FF98DF06FF5EE5492C293CC4" blockId="10.[151,1436,1899,2031]" pageId="10" pageNumber="58">
|
||
<collectingCountry id="F3257643FF98DF06FF5EE54929553CA8" box="[201,259,1899,1923]" name="Malta" pageId="10" pageNumber="58">Male</collectingCountry>
|
||
second pleopod unknown; female second pleopod with appendix interna. Uropod with lateral lobe of protopod bearing large, acute lateral tooth and smaller but distinct acute mesial tooth adjacent to lateral tooth; exopod with diaeresis straight, deeply curving mesially; distolateral tooth small, subacute, adjacent distolateral spiniform seta moderately elongate; distomesial margin with row of spiniform setae (
|
||
<figureCitation id="13092A56FF98DF06FB8CE5F52C0B3CC4" box="[1051,1117,2007,2031]" captionStart="FIGURE 5" captionStartId="8.[151,250,1852,1875]" captionTargetBox="[171,1398,198,1824]" captionTargetId="figure@8.[151,1436,193,1849]" captionTargetPageId="8" captionText="FIGURE 5. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, carapace, lateral view; D, tooth on ventromesial carina of first article of antennular peduncle, lateral view; E, antennal scaphocerite, dorsal view; F, third maxilliped, lateral view; G, right second pereiopod, lateral view (chela missing); H, chela of left second pereiopod, mesial view; I, third pereiopod, lateral view; J, fourth pereiopod, lateral view; K, fifth pereiopod, lateral view; L, uropod, dorsal view (setae omitted); M, telson, dorsal view." httpUri="https://zenodo.org/record/215151/files/figure.png" pageId="10" pageNumber="58">Fig. 5</figureCitation>
|
||
L).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF99DF07FF5EE2B529553BFF" blockId="11.[151,1436,151,716]" pageId="11" pageNumber="59">
|
||
Gill-exopod formula typical for genus (see above); reduced exopod-like lobe present at basis of minor cheliped (
|
||
<figureCitation id="13092A56FF99DF07FF08E29E28B53BFF" box="[159,227,188,212]" captionStart="FIGURE 6" captionStartId="9.[151,250,949,972]" captionTargetBox="[188,1367,190,876]" captionTargetId="figure@9.[151,1436,161,928]" captionTargetPageId="9" captionText="FIGURE 6. Athanopsis gotoi sp. nov.: holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101); A, major cheliped, mesial view; B, same, lateral view; C, distal major chela, dorsomesial view; D, coxa and ischium of major cheliped, mesial view; E, minor cheliped, lateral view." httpUri="https://zenodo.org/record/215152/files/figure.png" pageId="11" pageNumber="59">Fig. 6</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF99DF07FF5EE2FD29713A4F" blockId="11.[151,1436,151,716]" pageId="11" pageNumber="59">
|
||
<emphasis id="B946EAC1FF99DF07FF5EE2FD29DC3BD3" bold="true" box="[201,394,223,248]" pageId="11" pageNumber="59">Colour pattern.</emphasis>
|
||
Carapace almost completely covered with white chromatophores dorsally; first to fifth abdominal somites completely covered with red chromatophores; sixth abdominal somite, telson and uropods white; major cheliped hyaline-white; antennules, antennae, minor cheliped, and second to fifth pereiopods whitish (
|
||
<figureCitation id="13092A56FF99DF07FF08E36E28B73A4F" box="[159,225,332,356]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="11" pageNumber="59">Fig. 7</figureCitation>
|
||
C, D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3286558FF99DF07FF5EE3522B7A3974" pageId="11" pageNumber="59" type="etymology">
|
||
<paragraph id="8B8D36D3FF99DF07FF5EE3522DDA3A87" blockId="11.[151,1436,151,716]" pageId="11" pageNumber="59">
|
||
<emphasis id="B946EAC1FF99DF07FF5EE35229063AA3" bold="true" box="[201,336,368,392]" pageId="11" pageNumber="59">Etymology.</emphasis>
|
||
This new species is named after Ryutaro Goto (Kyoto University,
|
||
<collectingCountry id="F3257643FF99DF07FBC3E34D2CCE3AAC" box="[1108,1176,367,391]" name="Japan" pageId="11" pageNumber="59">Japan</collectingCountry>
|
||
), whose interest in the macrofauna associated with echiurans (
|
||
<bibRefCitation id="EFA34B22FF99DF07FDD8E3B62B5E3A87" author="Goto" box="[591,776,404,428]" pageId="11" pageNumber="59" refString="Goto, R., Hamamura, Y., & Kato, M. (2011) Morphological and ecological adaptation of Basterotia bivalves (Galeommatoidea: Sportellidae) to symbiotic association with burrowing echiuran worms. Zoological Science, 28, 225 - 234." type="journal article" year="2011">
|
||
Goto
|
||
<emphasis id="B946EAC1FF99DF07FD18E3B72A9E3A87" box="[655,712,404,428]" italics="true" pageId="11" pageNumber="59">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA34B22FF99DF07FC83E3B62BBE3A87" author="Goto" box="[788,1000,404,428]" pageId="11" pageNumber="59" refString="Goto, R. & Kato, M. (2012) Geographic mosaic of mutually exclusive dominance of obligate commensals in symbiotic communities associated with a burrowing echiuran worm. Marine Biology, 159, 319 - 330." type="journal article" year="2012">Goto & Kato 2012</bibRefCitation>
|
||
) led to the collection of the
|
||
<typeStatus id="54898871FF99DF07FAB5E3B62DDE3A87" box="[1314,1416,404,428]" pageId="11" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF99DF07FF5EE39A2B7A3974" blockId="11.[151,1436,151,716]" pageId="11" pageNumber="59">
|
||
<emphasis id="B946EAC1FF99DF07FF5EE39A297B3AFB" bold="true" box="[201,301,440,464]" pageId="11" pageNumber="59">Ecology.</emphasis>
|
||
The
|
||
<typeStatus id="54898871FF99DF07FEFDE395299A3AE4" box="[362,460,439,463]" pageId="11" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
was collected on a sand flat composed of gravel and coral sand exposed at low tide, under large rocks, in the gallery of the echiuran
|
||
<taxonomicName id="4C324D50FF99DF07FD20E3FE2D613ADF" authority="Leuckart & Ruppel, 1828" authorityName="Leuckart & Ruppel" authorityYear="1828" box="[695,1335,476,500]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="11" pageNumber="59" phylum="Echiura" rank="species" species="erythrogrammon">
|
||
<emphasis id="B946EAC1FF99DF07FD20E3FE2C583ADF" box="[695,1038,476,500]" italics="true" pageId="11" pageNumber="59">Ochetostoma erythrogrammon</emphasis>
|
||
<bibRefCitation id="EFA34B22FF99DF07FB81E3FE2D613ADF" author="Leuckart" box="[1046,1335,476,500]" pageId="11" pageNumber="59" refString="Leuckart, F. S. & Ruppel, W. P. S. (1828) Neue wirbellose Thiere des Rothen Meeres. Atlas zu der Reise im nordlichen Afrika, 1, Zoologie, 6 - 9." type="journal article" year="1828">Leuckart & Rüppel, 1828</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C324D50FF99DF07FAD1E3FE29B6393C" authority="Ochetostomatinae" authorityName="Ochetostomatinae" class="Echiuroidea" family="Thalassematidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiurida" pageId="11" pageNumber="59" phylum="Echiura" rank="family">Thalassematidae, Ochetostomatinae</taxonomicName>
|
||
) (
|
||
<figureCitation id="13092A56FF99DF07FE60E3DD2A6F393C" box="[503,569,511,535]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="11" pageNumber="59">Fig. 7</figureCitation>
|
||
C). The
|
||
<typeStatus id="54898871FF99DF07FD03E0222AA23933" box="[660,756,512,536]" pageId="11" pageNumber="59" type="paratype">paratype</typeStatus>
|
||
was found under rocks in
|
||
<quantity id="4CCA9B36FF99DF07FB8EE3DD2C12393C" box="[1049,1092,511,535]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="59" unit="m" value="6.0">6 m</quantity>
|
||
deep water, where it was associated with a generally similar-looking echiuran, possibly a species of
|
||
<taxonomicName id="4C324D50FF99DF07FC79E0062CD43917" box="[1006,1154,548,572]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="11" pageNumber="59" phylum="Echiura" rank="genus">
|
||
<emphasis id="B946EAC1FF99DF07FC79E0062CD43917" box="[1006,1154,548,572]" italics="true" pageId="11" pageNumber="59">Ochetostoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13092A56FF99DF07FB0FE0062CB43917" box="[1176,1250,548,572]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="11" pageNumber="59">Fig. 7</figureCitation>
|
||
D). No detailed ecological notes are available for the Indonesian specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3286558FF99DF07FF5EE04E2C7039AF" box="[201,1062,619,644]" pageId="11" pageNumber="59" type="materials_examined">
|
||
<paragraph id="8B8D36D3FF99DF07FF5EE04E2C7039AF" blockId="11.[151,1436,151,716]" box="[201,1062,619,644]" pageId="11" pageNumber="59">
|
||
<emphasis id="B946EAC1FF99DF07FF5EE04E293339AF" bold="true" box="[201,357,619,644]" pageId="11" pageNumber="59">
|
||
<typeStatus id="54898871FF99DF07FF5EE04E295439AF" box="[201,258,620,644]" pageId="11" pageNumber="59">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Southern
|
||
<collectingCountry id="F3257643FF99DF07FE4EE04F2A4A39AF" box="[473,540,621,644]" name="Japan" pageId="11" pageNumber="59">Japan</collectingCountry>
|
||
, Ryukyu Archipelago, Amami-Oshima
|
||
<collectingCountry id="F3257643FF99DF07FC4DE04E2C7439AF" box="[986,1058,620,644]" name="Iceland" pageId="11" pageNumber="59">Island</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3286558FF99DF07FF5EE0AD296A39E7" pageId="11" pageNumber="59" type="distribution">
|
||
<paragraph id="8B8D36D3FF99DF07FF5EE0AD296A39E7" blockId="11.[151,1436,151,716]" pageId="11" pageNumber="59">
|
||
<emphasis id="B946EAC1FF99DF07FF5EE0AD29343983" bold="true" box="[201,354,655,680]" pageId="11" pageNumber="59">Distribution.</emphasis>
|
||
Presently known only from the Ryukyu Archipelago,
|
||
<collectingCountry id="F3257643FF99DF07FC5EE0B22C5A398C" box="[969,1036,656,679]" name="Japan" pageId="11" pageNumber="59">Japan</collectingCountry>
|
||
, and Talaud Islands,
|
||
<collectingCountry id="F3257643FF99DF07FB6DE0AD2D31398C" box="[1274,1383,655,679]" name="Indonesia" pageId="11" pageNumber="59">Indonesia</collectingCountry>
|
||
(see map in
|
||
<figureCitation id="13092A56FF99DF07FF7CE096297839E7" box="[235,302,692,716]" captionStart="FIGURE 8" captionStartId="11.[151,250,1722,1745]" captionTargetBox="[257,1316,769,1680]" captionTargetId="figure@11.[257,1316,769,1690]" captionTargetPageId="11" captionText="FIGURE 8. Map of the Indo-West Pacific showing all collection localities for seven species of Athanopsis Coutière, 1897: A. platyrhynchus Coutière, 1897 (Djibouti; Pohnpei record questionable); A. dentipes Miya, 1980 (Japan: Nomo Bay, Sea of Ariake); A. brevirostris Banner & Banner, 1981 (Eritrea: Dahlak Archipelago; Madagascar: Nosy-Bé; Japan: Okinawa); A. australis Banner & Banner, 1982 (Australia: Port Phillip Bay); A. rubricinctuta Berggren, 1991 (Mozambique: Inhaca); A. saurus Anker, 2011 (Australia: Lizard Island); A. gotoi sp. nov. (Japan: Amami-shima, Okinawa; Indonesia: Talaud Islands)." httpUri="https://zenodo.org/record/215154/files/figure.png" pageId="11" pageNumber="59">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4D665BFF99DF07FF00E4982D153C60" httpUri="https://zenodo.org/record/215154/files/figure.png" pageId="11" pageNumber="59" targetBox="[257,1316,769,1680]" targetPageId="11">
|
||
<paragraph id="8B8D36D3FF99DF07FF00E4982D153C60" blockId="11.[151,1436,1722,1867]" pageId="11" pageNumber="59">
|
||
<emphasis id="B946EAC1FF99DF07FF00E49829453DFB" bold="true" box="[151,275,1722,1745]" pageId="11" pageNumber="59">FIGURE 8.</emphasis>
|
||
Map of the Indo-West Pacific showing all collection localities for seven species of
|
||
<taxonomicName id="4C324D50FF99DF07FBFDE4992D2A3DFB" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[1130,1404,1722,1745]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B946EAC1FF99DF07FBFDE4992C8D3DFA" box="[1130,1243,1723,1745]" italics="true" pageId="11" pageNumber="59">Athanopsis</emphasis>
|
||
Coutière, 1897
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C324D50FF99DF07FA11E49929923DC4" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="B946EAC1FF99DF07FA11E49929713DC4" italics="true" pageId="11" pageNumber="59">A. platyrhynchus</emphasis>
|
||
Coutière, 1897
|
||
</taxonomicName>
|
||
(Djibouti; Pohnpei record questionable);
|
||
<taxonomicName id="4C324D50FF99DF07FCFBE4F82C0F3DC4" authority="Miya, 1980" authorityName="Miya" authorityYear="1980" box="[876,1113,1753,1775]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="dentipes">
|
||
<emphasis id="B946EAC1FF99DF07FCFBE4F82B8B3DC4" box="[876,989,1753,1775]" italics="true" pageId="11" pageNumber="59">A. dentipes</emphasis>
|
||
Miya, 1980
|
||
</taxonomicName>
|
||
(Japan: Nomo Bay, Sea of Ariake);
|
||
<taxonomicName id="4C324D50FF99DF07FF58E4DB2A0E3C25" authority="Banner & Banner, 1981" authorityName="Banner & Banner" authorityYear="1981" box="[207,600,1784,1806]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="B946EAC1FF99DF07FF58E4DB29373C25" box="[207,353,1784,1806]" italics="true" pageId="11" pageNumber="59">A. brevirostris</emphasis>
|
||
Banner & Banner, 1981
|
||
</taxonomicName>
|
||
(Eritrea: Dahlak Archipelago; Madagascar: Nosy-Bé; Japan: Okinawa);
|
||
<taxonomicName id="4C324D50FF99DF07FAA0E4DB29E63C07" authority="Banner & Banner, 1982" authorityName="Banner & Banner" authorityYear="1982" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B946EAC1FF99DF07FAA0E4DB28F93C06" italics="true" pageId="11" pageNumber="59">A. australis</emphasis>
|
||
Banner & Banner, 1982
|
||
</taxonomicName>
|
||
(Australia: Port Phillip Bay);
|
||
<taxonomicName id="4C324D50FF99DF07FD7BE53A2C6F3C07" authority="Berggren, 1991" authorityName="Berggren" authorityYear="1991" box="[748,1081,1815,1837]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="B946EAC1FF99DF07FD7BE53A2BC73C06" box="[748,913,1815,1837]" italics="true" pageId="11" pageNumber="59">A. rubricinctuta</emphasis>
|
||
Berggren, 1991
|
||
</taxonomicName>
|
||
(Mozambique: Inhaca);
|
||
<taxonomicName id="4C324D50FF99DF07FAADE53A29403C60" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="B946EAC1FF99DF07FAADE53A2DCA3C06" box="[1338,1436,1816,1837]" italics="true" pageId="11" pageNumber="59">A. saurus</emphasis>
|
||
Anker, 2011
|
||
</taxonomicName>
|
||
(Australia: Lizard Island);
|
||
<taxonomicName id="4C324D50FF99DF07FDBDE5142A2C3C60" box="[554,634,1846,1867]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF99DF07FDBDE5142A2C3C60" box="[554,634,1846,1867]" italics="true" pageId="11" pageNumber="59">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557BAFF99DF07FD16E5142A823C60" box="[641,724,1846,1867]" pageId="11" pageNumber="59" rank="species">
|
||
<emphasis id="B946EAC1FF99DF07FD16E5142A823C60" bold="true" box="[641,724,1846,1867]" pageId="11" pageNumber="59">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Japan: Amami-shima, Okinawa; Indonesia: Talaud Islands).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3286558FF99DF00FF5EE5512BC53964" lastPageId="12" lastPageNumber="60" pageId="11" pageNumber="59" type="discussion">
|
||
<paragraph id="8B8D36D3FF99DF00FF5EE5512B963A76" blockId="11.[151,1436,1907,2035]" lastBlockId="12.[151,1437,151,592]" lastPageId="12" lastPageNumber="60" pageId="11" pageNumber="59">
|
||
<emphasis id="B946EAC1FF99DF07FF5EE551296B3CA0" bold="true" box="[201,317,1907,1931]" pageId="11" pageNumber="59">Remarks.</emphasis>
|
||
<taxonomicName id="4C324D50FF99DF07FED2E5512A533CA0" box="[325,517,1907,1931]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF99DF07FED2E5512A533CA0" box="[325,517,1907,1931]" italics="true" pageId="11" pageNumber="59">Athanopsis gotoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557BAFF99DF07FD9AE5512A3F3CA0" box="[525,617,1907,1931]" pageId="11" pageNumber="59" rank="species">
|
||
<emphasis id="B946EAC1FF99DF07FD9AE5512A3F3CA0" bold="true" box="[525,617,1907,1931]" pageId="11" pageNumber="59">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is most closely related to
|
||
<taxonomicName id="4C324D50FF99DF07FC0EE5512C0E3CA0" box="[921,1112,1907,1931]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="B946EAC1FF99DF07FC0EE5512C0E3CA0" box="[921,1112,1907,1931]" italics="true" pageId="11" pageNumber="59">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C324D50FF99DF07FBFFE5512D5F3CA0" box="[1128,1289,1907,1931]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="B946EAC1FF99DF07FBFFE5512D5F3CA0" box="[1128,1289,1907,1931]" italics="true" pageId="11" pageNumber="59">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
(see above), and
|
||
<taxonomicName id="4C324D50FF99DF07FF5EE5B429633C85" box="[201,309,1942,1966]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="B946EAC1FF99DF07FF5EE5B429633C85" box="[201,309,1942,1966]" italics="true" pageId="11" pageNumber="59">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFA34B22FF99DF07FEE2E5B72A5B3C86" author="Anker" box="[373,525,1941,1965]" pageId="11" pageNumber="59" refString="Anker, A. (2011 a) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734, 1 - 22." type="journal article" year="2011" yearSuffix="a">Anker 2011a</bibRefCitation>
|
||
). These four species are characterised by the unarmed meri of the third to fifth pereiopods, as opposed to the other three species of the genus, viz.
|
||
<taxonomicName id="4C324D50FF99DF07FC17E59A2BAA3CFB" box="[896,1020,1976,2000]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="dentipes">
|
||
<emphasis id="B946EAC1FF99DF07FC17E59A2BAA3CFB" box="[896,1020,1976,2000]" italics="true" pageId="11" pageNumber="59">A. dentipes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C324D50FF99DF07FB9EE59A2CDC3CFB" box="[1033,1162,1976,2000]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B946EAC1FF99DF07FB9EE59A2CDC3CFB" box="[1033,1162,1976,2000]" italics="true" pageId="11" pageNumber="59">A. australis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C324D50FF99DF07FB51E59A2D2F3CFB" box="[1222,1401,1976,2000]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="59" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="B946EAC1FF99DF07FB51E59A2D2F3CFB" box="[1222,1401,1976,2000]" italics="true" pageId="11" pageNumber="59">A. rubricinctuta</emphasis>
|
||
</taxonomicName>
|
||
, in which the meri of the third to fifth pereiopods are armed with one or more stout spiniform setae. Based on morphological grounds,
|
||
<taxonomicName id="4C324D50FF9EDF00FEC6E2BA29FE3B9B" box="[337,424,152,176]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF9EDF00FEC6E2BA29FE3B9B" box="[337,424,152,176]" italics="true" pageId="12" pageNumber="60">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557BAFF9EDF00FE38E2BA2A5E3B9B" box="[431,520,152,176]" pageId="12" pageNumber="60" rank="species">
|
||
<emphasis id="B946EAC1FF9EDF00FE38E2BA2A5E3B9B" bold="true" box="[431,520,152,176]" pageId="12" pageNumber="60">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be separated from all of them by the dorsally markedly domed carapace; from
|
||
<taxonomicName id="4C324D50FF9EDF00FF00E29929023BF8" box="[151,340,187,211]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="B946EAC1FF9EDF00FF00E29929023BF8" box="[151,340,187,211]" italics="true" pageId="12" pageNumber="60">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
by the non-gaping, serrated finger cutting edges of the major chela (gaping and armed with a few large teeth in
|
||
<taxonomicName id="4C324D50FF9EDF00FEA4E2FF29A73BDE" box="[307,497,221,245]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="B946EAC1FF9EDF00FEA4E2FF29A73BDE" box="[307,497,221,245]" italics="true" pageId="12" pageNumber="60">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
, cf.
|
||
<figureCitation id="13092A56FF9EDF00FDB1E2FF2A303BDE" box="[550,614,221,245]" captionStart="FIGURE 1" captionStartId="2.[151,250,702,725]" captionTargetBox="[181,1356,205,636]" captionTargetId="figure@2.[151,1436,193,681]" captionTargetPageId="2" captionText="FIGURE 1. Athanopsis platyrhynchus Coutière, 1897: type specimen from Djibouti: A, anterior carapace, dorsal view; B, same, lateral view; C, rostrum and right frontal appendages, dorsal view; D, major cheliped, mesial view; E, minor cheliped, mesial view (all figures adapted from Coutière 1899)." httpUri="https://zenodo.org/record/215147/files/figure.png" pageId="12" pageNumber="60">Fig. 1</figureCitation>
|
||
D); and from both
|
||
<taxonomicName id="4C324D50FF9EDF00FCD7E2FF2BB73BDE" box="[832,993,221,245]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="B946EAC1FF9EDF00FCD7E2FF2BB73BDE" box="[832,993,221,245]" italics="true" pageId="12" pageNumber="60">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C324D50FF9EDF00FB8DE2FF2CD33BDE" box="[1050,1157,221,245]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="B946EAC1FF9EDF00FB8DE2FF2CD33BDE" box="[1050,1157,221,245]" italics="true" pageId="12" pageNumber="60">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of dense setae on the fingers of the major chela, the distally blunter and much shorter stylocerite (not reaching third article of the antennular peduncle in
|
||
<taxonomicName id="4C324D50FF9EDF00FE7EE3012A153A10" box="[489,579,291,315]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF9EDF00FE7EE3012A153A10" box="[489,579,291,315]" italics="true" pageId="12" pageNumber="60">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B946EAC1FF9EDF00FDDDE3012AF33A10" bold="true" box="[586,677,291,315]" pageId="12" pageNumber="60">
|
||
<taxonomicNameLabel id="A27557BAFF9EDF00FDDDE3012AF33A10" box="[586,677,291,315]" pageId="12" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
vs. reaching mid-length of this article in the other two species; see
|
||
<figureCitation id="13092A56FF9EDF00FF00E367288F3A76" box="[151,217,325,349]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1822]" captionTargetBox="[177,1393,367,1748]" captionTargetId="figure@4.[174,1410,357,1774]" captionTargetPageId="4" captionText="FIGURE 2. Athanopsis brevirostris Banner & Banner, 1981: A – N, ovigerous female from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14487), O – Q, male from Nosy-Bé (FLMNH UF Arthropoda 14486); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, tooth on ventromesial carina of first article of antennular peduncle, lateral view; D, third maxilliped, lateral view; E, second pereiopod, lateral view; F, third pereiopod, lateral view; G, same, distal propodus and dactylus, lateral view; H, fourth pereiopod, lateral view; I, fifth pereiopod, lateral view; J, same, distal propodus and dactylus, lateral view; K, uropod, dorsal view; L, same, distal exopod, dorsal view; M, telson, dorsal view; N, same, posterior margin, dorsal view; O, frontal margin of carapace, lateral view; P, fifth pereiopod, ischium, lateral view; Q, second pleopod, appendix masculina and appendix interna, lateral view." httpUri="https://zenodo.org/record/215148/files/figure.png" pageId="12" pageNumber="60">Fig. 2</figureCitation>
|
||
A;
|
||
<bibRefCitation id="EFA34B22FF9EDF00FF6CE36729C63A76" author="Anker" box="[251,400,325,349]" pageId="12" pageNumber="60" refString="Anker, A. (2011 a) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734, 1 - 22." type="journal article" year="2011" yearSuffix="a">Anker 2011a</bibRefCitation>
|
||
), and the more tapering telson (cf.
|
||
<figureCitation id="13092A56FF9EDF00FC81E3672B343A76" box="[790,866,325,349]" captionStart="FIGURE 2" captionStartId="4.[151,250,1799,1822]" captionTargetBox="[177,1393,367,1748]" captionTargetId="figure@4.[174,1410,357,1774]" captionTargetPageId="4" captionText="FIGURE 2. Athanopsis brevirostris Banner & Banner, 1981: A – N, ovigerous female from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14487), O – Q, male from Nosy-Bé (FLMNH UF Arthropoda 14486); A, frontal margin of carapace and frontal appendages, dorsal view; B, same, lateral view; C, tooth on ventromesial carina of first article of antennular peduncle, lateral view; D, third maxilliped, lateral view; E, second pereiopod, lateral view; F, third pereiopod, lateral view; G, same, distal propodus and dactylus, lateral view; H, fourth pereiopod, lateral view; I, fifth pereiopod, lateral view; J, same, distal propodus and dactylus, lateral view; K, uropod, dorsal view; L, same, distal exopod, dorsal view; M, telson, dorsal view; N, same, posterior margin, dorsal view; O, frontal margin of carapace, lateral view; P, fifth pereiopod, ischium, lateral view; Q, second pleopod, appendix masculina and appendix interna, lateral view." httpUri="https://zenodo.org/record/215148/files/figure.png" pageId="12" pageNumber="60">Figs. 2</figureCitation>
|
||
M, 5M).
|
||
</paragraph>
|
||
<paragraph id="8B8D36D3FF9EDF00FF5EE3452BC53964" blockId="12.[151,1437,151,592]" pageId="12" pageNumber="60">
|
||
The colour pattern of
|
||
<taxonomicName id="4C324D50FF9EDF00FE28E34A2A4E3AAB" box="[447,536,360,384]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="gotoi" status="sp. nov.">
|
||
<emphasis id="B946EAC1FF9EDF00FE28E34A2A4E3AAB" box="[447,536,360,384]" italics="true" pageId="12" pageNumber="60">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27557BAFF9EDF00FDB7E34A2A2C3AAB" box="[544,634,360,384]" pageId="12" pageNumber="60" rank="species">
|
||
<emphasis id="B946EAC1FF9EDF00FDB7E34A2A2C3AAB" bold="true" box="[544,634,360,384]" pageId="12" pageNumber="60">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
appears to be unique within the genus by combining the uniformly red first to fifth abdominal somites with the entirely white carapace, sixth abdominal somite and tail fan (
|
||
<figureCitation id="13092A56FF9EDF00FA84E3A82D003A89" box="[1299,1366,394,418]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="12" pageNumber="60">Fig. 7</figureCitation>
|
||
C, D). This colour pattern clearly differs from those of
|
||
<taxonomicName id="4C324D50FF9EDF00FD26E38F2B063AEE" box="[689,848,429,453]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="B946EAC1FF9EDF00FD26E38F2B063AEE" box="[689,848,429,453]" italics="true" pageId="12" pageNumber="60">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13092A56FF9EDF00FCC8E38F2BF73AED" box="[863,929,429,454]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="12" pageNumber="60">Fig. 7</figureCitation>
|
||
A, B) and
|
||
<taxonomicName id="4C324D50FF9EDF00FB82E38F2C293AEE" box="[1045,1151,429,453]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="B946EAC1FF9EDF00FB82E38F2C293AEE" box="[1045,1151,429,453]" italics="true" pageId="12" pageNumber="60">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13092A56FF9EDF00FB1AE38F2C993AEE" box="[1165,1231,429,453]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="12" pageNumber="60">Fig. 7</figureCitation>
|
||
E), in which a few broad white bands are intercalated into the mostly red abdomen, and from that of
|
||
<taxonomicName id="4C324D50FF9EDF00FBBBE3F22CBF3AC3" box="[1068,1257,464,488]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="B946EAC1FF9EDF00FBBBE3F22CBF3AC3" box="[1068,1257,464,488]" italics="true" pageId="12" pageNumber="60">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
, with red transverse bands on the carapace (
|
||
<bibRefCitation id="EFA34B22FF9EDF00FE70E3D02B583921" author="Coutiere" box="[487,782,498,522]" pageId="12" pageNumber="60" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, serie 8, 9, 1 - 559, pls. 1 - 6. Masson, Paris." type="journal article" year="1899">Coutière 1899; see above</bibRefCitation>
|
||
). The three remaining and more distantly related species, viz.
|
||
<taxonomicName id="4C324D50FF9EDF00FF5FE037292B3906" box="[200,381,533,557]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="B946EAC1FF9EDF00FF5FE037292B3906" box="[200,381,533,557]" italics="true" pageId="12" pageNumber="60">A. rubricinctuta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C324D50FF9EDF00FE1BE0372A593906" box="[396,527,533,557]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B946EAC1FF9EDF00FE1BE0372A593906" box="[396,527,533,557]" italics="true" pageId="12" pageNumber="60">A. australis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C324D50FF9EDF00FDD8E0372A9B3906" box="[591,717,533,557]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="dentipes">
|
||
<emphasis id="B946EAC1FF9EDF00FDD8E0372A9B3906" box="[591,717,533,557]" italics="true" pageId="12" pageNumber="60">A. dentipes</emphasis>
|
||
</taxonomicName>
|
||
exhibit even more different colourations (
|
||
<figureCitation id="13092A56FF9EDF00FB38E0372CA43906" box="[1199,1266,533,557]" captionStart="FIGURE 7" captionStartId="10.[151,250,1592,1615]" captionTargetBox="[151,1436,193,1569]" captionTargetId="figure@10.[151,1436,193,1571]" captionTargetPageId="10" captionText="FIGURE 7. Colour patterns of five species of Athanopsis Coutière, 1897 A, B— A. brevirostris Banner & Banner, 1981, male (A) and ovigerous female (B) from Nosy-Bé, Madagascar (FLMNH UF Arthropoda 14486, 14487), both in lateral view; C, D— A. gotoi sp. nov., C, holotype, ovigerous female from Amami-Oshima, Japan (MNHN-IU- 2010 - 4101), sitting on its host, Ochetostoma erythrogrammon Leuckart & Rüppel, 1829, and dorsolateral view (both in vitro); D, non-type, ovigerous female from Okinawa (CBM-ZC 10620) on echiuran host (in situ); E— A. saurus Anker, 2011, holotype, male from Lizard Island, Australia (QM W 29048), lateral view; F, G— A. rubricinctuta Berggren, 1991, ovigerous female from Inhaca, Moçambique (probably allotype, RMNH D 39853), lateral (F) and dorsal (G) view in aquarium; H— A. australis Banner & Banner, 1982; male from Port Phillip Bay, Victoria, Australia (NMV), lateral view. Photographs: A, B, E, by the author; C, by R. Goto; D, by Y. Yamada; F, G, by M. Berggren; H, by M. Marmach / Museum Victoria." httpUri="https://zenodo.org/record/215153/files/figure.png" pageId="12" pageNumber="60">Fig. 7</figureCitation>
|
||
F, G, H; colour pattern of
|
||
<taxonomicName id="4C324D50FF9EDF00FE9DE01A29D0397B" box="[266,390,568,592]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="dentipes">
|
||
<emphasis id="B946EAC1FF9EDF00FE9DE01A29D0397B" box="[266,390,568,592]" italics="true" pageId="12" pageNumber="60">A. dentipes</emphasis>
|
||
</taxonomicName>
|
||
similar to that of
|
||
<taxonomicName id="4C324D50FF9EDF00FDD9E01A2A99397B" box="[590,719,568,592]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="60" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B946EAC1FF9EDF00FDD9E01A2A99397B" box="[590,719,568,592]" italics="true" pageId="12" pageNumber="60">A. australis</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="EFA34B22FF9EDF00FC9FE0152BD03964" author="Miya" box="[776,902,567,591]" pageId="12" pageNumber="60" refString="Miya, Y. (1980) Two new records of the genera, Athanopsis and Prionalpheus, from Japan, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biology Laboratory, 5, 117 - 131." type="journal article" year="1980">Miya 1980</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |