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<mods:title id="0B2D0C169E003E6AD8954BB992CDCF0A">Continental fishes of Nicaragua: diversity, distribution and conservation status; with an annotated and illustrated checklist of species and an identification guide to families</mods:title>
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<mods:namePart id="59C6181B1A67F22AD24603BB2BA70FFB">Angulo, Arturo</mods:namePart>
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<mods:namePart id="BFE3D0A04D102E089587B86E2FA3E17C">Betts, Joel T.</mods:namePart>
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<mods:affiliation id="AE17604373478B8AC4AD243E82EB7385">Universidad Americana (UAM, Costado Noroeste Camino de Oriente, Managua, Nicaragua edgar. ecologica @ gmail. com; https: // orcid. org / 0000 - 0001 - 8148 - 3195</mods:affiliation>
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<mods:affiliation id="364B1DAEBCC5FCC54888B0FA7486C622">Department of Environment and Development, Zamorano Biodiversity Center. Zamorano University of Tegucigalpa, P. O. Box 93, Honduras evandenberghe @ zamorano. edu; https: // orcid. org / 0000 - 0001 - 7566 - 0415</mods:affiliation>
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<mods:namePart id="429394B9C3CD84203E37A9B795DA3B7C">Elías, Diego J.</mods:namePart>
<mods:affiliation id="1B8DE9C83877DC6BA2A6FA7D6EF37691">Field Museum of Natural History (FMNH). Chicago, Illinois, United States of America delias @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 4215 - 0384 cmcmahan @ fieldmuseum. org; https: // orcid. org / 0000 - 0003 - 2113 - 8554</mods:affiliation>
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<mods:date id="096B9C25A041CF985CCD84BF10CBBF4B">2023</mods:date>
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<taxonomicName id="4C254D1DE861FF97FF7929F7FDD0327E" ID-CoL="K3G7" authority="(Gunther 1861)" baseAuthorityName="Gunther" baseAuthorityYear="1861" box="[151,625,1512,1538]" family="Gobiidae" genus="Awaous" isUncertain="true" kingdom="Animalia" order="Perciformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="transandeanus">
<emphasis id="B951EA8CE861FF97FF7929F7FF4F327E" box="[151,238,1512,1538]" italics="true" pageId="34" pageNumber="35">Awaous</emphasis>
cf.
<emphasis id="B951EA8CE861FF97FEF929F7FE1D327E" box="[279,444,1512,1538]" italics="true" pageId="34" pageNumber="35">transandeanus</emphasis>
(Günther 1861)
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. Pacific River Goby; Chupapiedras, Guavina, Lamearenas, Gobio, Gobio de Río del Pacífico, Dormilón
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<emphasis id="B951EA8CE861FF97FF792A2FFEB13236" bold="true" box="[151,272,1584,1610]" pageId="34" pageNumber="35">Vouchers:</emphasis>
UCR 0052-009 (1), 0183-004 (2), 0184-003 (30) and 0270-005 (23).
<emphasis id="B951EA8CE861FF97FBA12A2FFB4A3236" bold="true" box="[1103,1259,1584,1610]" pageId="34" pageNumber="35">Distribution:</emphasis>
North, Central and South America; from central Baja California Sur and Sonora, Mexico, to Tumbes, Peru, Pacific drainages;
<emphasis id="B951EA8CE861FF97FF792A67FE8632EE" bold="true" box="[151,295,1656,1682]" pageId="34" pageNumber="35">
(
<collectingCountry id="F332760EE861FF97FF712A67FEBE32EE" box="[159,287,1656,1682]" name="Nicaragua" pageId="34" pageNumber="35">Nicaragua</collectingCountry>
)
</emphasis>
Ch, Ne, ER, PN, Ni (Pacific);
<quantity id="4CDD9B7BE861FF97FD6F2A67FCA732EE" box="[641,774,1656,1682]" metricMagnitude="2" metricUnit="m" metricValue="1.595" metricValueMax="3.19" metricValueMin="0.0" pageId="34" pageNumber="35" unit="m" value="159.5" valueMax="319.0" valueMin="0.0">0319 masl</quantity>
; Per, Dia, Cat.
<emphasis id="B951EA8CE861FF97FC412A67FAC432EE" bold="true" box="[943,1381,1656,1682]" pageId="34" pageNumber="35">Occurrence and conservation status:</emphasis>
Nat; LC (2020), population trend unknown.
<emphasis id="B951EA8CE861FF97FDB52A83FD4032CA" bold="true" box="[603,737,1692,1718]" pageId="34" pageNumber="35">Literature:</emphasis>
<bibRefCitation id="EFB44B6FE861FF97FD072A83FC0932CA" author="Villa" box="[745,936,1692,1718]" firstAuthor="Villa" page="227" pageId="34" pageNumber="35" refId="ref86666" refString="Villa, J. (1982) Peces nicaraguenses de agua dulce. Editorial Union Cardoza y Cia. Ltda, Managua, 253 pp." type="book" year="1982">Villa (1982: 227</bibRefCitation>
; detailed description, including illustrations and an identification key, and information on distribution and ecology),
<bibRefCitation id="EFB44B6FE861FF97FC052ADFFAF032A6" author="Carr &amp; Giovannoli" box="[1003,1361,1728,1754]" firstAuthor="Carr" page="111" pageId="34" pageNumber="35" pagination="95 - 115" refId="ref81867" refString="Carr, A. F. Jr. &amp; Giovannoli, L. (1994) The fishes of the Choluteca drainage of southern Honduras. Ceiba, 35 (2), 95 - 115." type="journal article" year="1994">Carr &amp; Giovannoli (1994: 111</bibRefCitation>
, as
<taxonomicName id="4C254D1DE861FF97FA6A2ADFFF513282" baseAuthorityName="Lichtenstein" baseAuthorityYear="1822" family="Gobiidae" genus="Awaous" kingdom="Animalia" order="Perciformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="tajasica">
<emphasis id="B951EA8CE861FF97FA6A2ADFFF513282" italics="true" pageId="34" pageNumber="35">A. tajasica</emphasis>
</taxonomicName>
—misidentification; brief description, including information on distribution and ecology),
<bibRefCitation id="EFB44B6FE861FF97FB1F2AFBFF65335E" author="Bussing" firstAuthor="Bussing" page="398" pageId="34" pageNumber="35" refId="ref81779" refString="Bussing, W. A. (1998). Freshwater fishes of Costa Rica. Editorial Universidad de Costa Rica, San Jose´, Costa Rica, 468 pp." type="book" year="1998">Bussing (1998: 398</bibRefCitation>
; brief description, including illustrations and an identification key, and information on distribution, with a map, and ecology),
<bibRefCitation id="EFB44B6FE861FF97FE952B33FDEB333A" author="Angulo, A." box="[379,586,1836,1862]" pageId="34" pageNumber="35" pagination="1 - 72" refId="ref80295" refString="Angulo, A. (2021) New records and range extensions to the Costa Rican freshwater fish fauna, with an updated checklist. Zootaxa, 5083 (1), 1 - 72, https: // doi. org / 10.11646 / zootaxa. 5083.1.1" type="journal article" year="2021">Angulo (2021: 36</bibRefCitation>
; listed, including information on distribution) and
<bibRefCitation id="EFB44B6FE861FF97FB7C2B33FF653316" author="McMahan" etAl="et al." firstAuthor="McMahan" page="970" pageId="34" pageNumber="35" pagination="970 - 979" refId="ref84371" refString="McMahan, C. D., Elias, D. J., Li, Y., Dominguez-Dominguez, O., Rodriguez-Machado, S., Morales-Cabrera, A., Velasquez- Ramirez, D., Piller, K. R., Chakrabarty, P. &amp; Matamoros, W. A. (2021) Molecular systematics of the Awaous banana complex (River gobies; Teleostei: Oxudercidae). Journal of Fish Biology, 99 (3), 970 - 979. https: // doi. org / 10.1111 / jfb. 14783" type="journal article" year="2021">
McMahan
<emphasis id="B951EA8CE861FF97FAE12B32FAEB333A" box="[1295,1354,1836,1862]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(2021: 970
</bibRefCitation>
; phylogenetic relationships, and information on distribution, with a map, and ecology).
<emphasis id="B951EA8CE861FF97FB382B4FFAED3316" bold="true" box="[1238,1356,1872,1898]" pageId="34" pageNumber="35">Remarks:</emphasis>
Based on molecular data,
<bibRefCitation id="EFB44B6FE861FF97FE962B6BFD2433F2" author="McMahan" box="[376,645,1908,1934]" etAl="et al." firstAuthor="McMahan" pageId="34" pageNumber="35" pagination="970 - 979" refId="ref84371" refString="McMahan, C. D., Elias, D. J., Li, Y., Dominguez-Dominguez, O., Rodriguez-Machado, S., Morales-Cabrera, A., Velasquez- Ramirez, D., Piller, K. R., Chakrabarty, P. &amp; Matamoros, W. A. (2021) Molecular systematics of the Awaous banana complex (River gobies; Teleostei: Oxudercidae). Journal of Fish Biology, 99 (3), 970 - 979. https: // doi. org / 10.1111 / jfb. 14783" type="journal article" year="2021">
McMahan
<emphasis id="B951EA8CE861FF97FE1B2B6AFD9133F2" box="[501,560,1908,1934]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(2021)
</bibRefCitation>
suggested the presence of two different lineages under the name
<taxonomicName id="4C254D1DE861FF97FA6A2B6BFE9D33CE" baseAuthorityName="Gunther" baseAuthorityYear="1861" family="Gobiidae" genus="Awaous" kingdom="Animalia" order="Perciformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="transandeanus">
<emphasis id="B951EA8CE861FF97FA6A2B6BFE9D33CE" italics="true" pageId="34" pageNumber="35">A. transandeanus</emphasis>
</taxonomicName>
in the Pacific coast of Mexico and Central America. The boundary between the two lineages is the northern portion of the Costa Rican Pacific (Guanacaste) coast. Morphological studies are necessary to corroborate this and to properly diagnose both species.
</paragraph>
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