470 lines
54 KiB
XML
470 lines
54 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1193.114957" ID-Pensoft-Pub="1313-2970-1193-181" ID-Pensoft-UUID="E1868F688C4451E787BCFCC7E618177E" ID-ZooBank="311FA5D6CBA54C738811861D5A27F8FF" ModsDocID="1313-2970-1193-181" checkinTime="1709803639292" checkinUser="pensoft" docAuthor="Constant, Jerome, Pham, Thai-Hong, Le, Cuong Viet Canh, Vu, Trung Thanh, Nguyen, Hoai Thu Thi & Tran, Hai Nam" docDate="2024" docId="81D7FFA74E0B5432A46C66C3F695FA8F" docLanguage="en" docName="ZooKeys 1193: 181-194" docOrigin="ZooKeys 1193" docPubDate="2024-03-06" docSource="http://dx.doi.org/10.3897/zookeys.1193.114957" docTitle="Goniopsarites mientrunganus Constant & Pham 2024, sp. nov." docType="treatment" docUuid="689317C8-D1E4-4526-81CC-E89B9C11E42B" docUuidSource="ZooBank" docVersion="1" id="E1868F688C4451E787BCFCC7E618177E" lastPageNumber="181" masterDocId="E1868F688C4451E787BCFCC7E618177E" masterDocTitle="Pisachini planthoppers of Vietnam: new records of Pisacha and a new Goniopsarites species from Central Vietnam (Hemiptera, Fulgoromorpha, Nogodinidae)" masterLastPageNumber="194" masterPageNumber="181" pageNumber="181" updateTime="1709803639292" updateUser="pensoft">
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<mods:titleInfo id="66E1821B5CD42A8E7A32E8D6B1079326">
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<mods:title id="16352B5711D532AD2A81832AF2F20399">Pisachini planthoppers of Vietnam: new records of Pisacha and a new Goniopsarites species from Central Vietnam (Hemiptera, Fulgoromorpha, Nogodinidae)</mods:title>
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<mods:name id="D1FD2C5A5213437E889DF64A1FC23559" type="personal">
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<mods:namePart id="5316561CAC321E8E9A1F60ED86BA4FA9">Constant, Jerome</mods:namePart>
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<mods:nameIdentifier id="8D778D6DD2426F509B59EBD12710E982" type="ORCID">https://orcid.org/0000-0003-0254-0863</mods:nameIdentifier>
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<mods:affiliation id="722468A6AACE37055A9CADB0232CEC80">Royal Belgian Institute of Natural Sciences, O. D. Taxonomy & Phylogeny - Entomology, Vautier street 29, B- 1000 Brussels, Belgium</mods:affiliation>
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<mods:nameIdentifier id="770BA8DF8A9077D4D42798EB658ACA4F" type="email">jconstant@naturalsciences.be</mods:nameIdentifier>
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<mods:roleTerm id="AEBED45969D1FF6125CCF0167C5AD98E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="455E8F2114811730B1CE0E81D06F7BBE">Pham, Thai-Hong</mods:namePart>
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<mods:nameIdentifier id="CC5C67AF4AF4BB91B6F928930C631063" type="ORCID">https://orcid.org/0000-0002-4763-3679</mods:nameIdentifier>
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<mods:affiliation id="1EF2508FDB155DC8A970621C0E10039E">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam & Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier id="934218DDAC4DF06232B89682009875D2" type="email">phamthai1976@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:namePart id="53ED29684C66F1D24BF4F47CD811E471">Le, Cuong Viet Canh</mods:namePart>
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<mods:nameIdentifier id="670DB3CA3E2EB36826991FF11125BDD1" type="ORCID">https://orcid.org/0000-0002-1430-6305</mods:nameIdentifier>
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<mods:affiliation id="8180309B00A01A4265BCB2C8514CC934">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name id="5768D34CD1C50D67E298F3FD1D096A3E" type="personal">
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<mods:namePart id="731FB11265BCE6D35F842096DEDB81FC">Vu, Trung Thanh</mods:namePart>
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<mods:nameIdentifier id="0AB93EBBCA68566C8F8506379F991B4B" type="ORCID">https://orcid.org/0009-0002-8987-1672</mods:nameIdentifier>
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<mods:affiliation id="2BFA2625CBD3D1F0BC451121603CDFBC">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<mods:name id="A5FC8F55F620DC297285345A2B9C36BF" type="personal">
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<mods:roleTerm id="660677D56D866E5ED9129EB5DC87AA09">Author</mods:roleTerm>
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<mods:namePart id="926FF64E72ED8F7EDAC0546EDB33B96F">Nguyen, Hoai Thu Thi</mods:namePart>
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<mods:nameIdentifier id="F87FB0704F5FC8C136E54F258666A44E" type="ORCID">https://orcid.org/0009-0004-9130-2940</mods:nameIdentifier>
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<mods:affiliation id="ABBD6E7A3719DB95B4C7AA999CAB091E">Vietnam National Museum of Nature, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="E40D0F62E457EA4B401D613F805DBBDA">Tran, Hai Nam</mods:namePart>
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<mods:nameIdentifier id="7C511C96B96FA0DE4896DE0F9EBABC12" type="ORCID">https://orcid.org/0009-0003-1557-1666</mods:nameIdentifier>
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<mods:affiliation id="3C2534807B198C3252741B54A4E20376">Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam & Department of Biology, Hanoi National University of Education, 136 Xuan Thuy, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:typeOfResource id="36589F48527C92DADD6764F9761E0C30">text</mods:typeOfResource>
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<mods:title id="72DEB2B706F4ED401975E9B4D8DA8214">ZooKeys</mods:title>
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<mods:part id="19BF76FACF067CF9C03AD3020DE571F5">
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<mods:date id="D10EF03C512EF9C1D92A7FD1D26A2A42">2024</mods:date>
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<mods:number id="6568DF20DCE57A751E3A078E089C0685">2024-03-06</mods:number>
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<mods:detail id="FC844ACA9163511717802419EA862406" type="volume">
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<mods:number id="F0086E8698A8374808D26F200F33586D">1193</mods:number>
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<mods:start id="1487A93901BE6BA16532FC9D91A116F3">181</mods:start>
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<mods:end id="713626EDB13814EE332C54EEFDD90252">194</mods:end>
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<mods:url id="132A71094EEE7A986A865E1BD9B860DF">http://dx.doi.org/10.3897/zookeys.1193.114957</mods:url>
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<mods:classification id="F9F735BD5BB9BBD7E6FD5328842D35F6">journal article</mods:classification>
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<mods:identifier id="CB346703145E1F8C74466351899835F1" type="DOI">http://dx.doi.org/10.3897/zookeys.1193.114957</mods:identifier>
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<mods:identifier id="4708432D82C1026089AF3F7527B61A12" type="Pensoft-Pub">1313-2970-1193-181</mods:identifier>
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<treatment id="81D7FFA74E0B5432A46C66C3F695FA8F" LSID="urn:lsid:zoobank.org:act:689317C8-D1E4-4526-81CC-E89B9C11E42B" httpUri="http://treatment.plazi.org/id/81D7FFA74E0B5432A46C66C3F695FA8F" lastPageNumber="181" pageId="0" pageNumber="181">
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<subSubSection id="A218713E15898EEAA7390A075C69FBA3" pageId="0" pageNumber="181" type="nomenclature">
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<paragraph id="3E830C62F0925D81E1EEABC53BB4A1A7" pageId="0" pageNumber="181">
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<taxonomicName id="4635CE51A683EA1CFAFBF3E511E230B8" LSID="https://zoobank.org/689317C8-D1E4-4526-81CC-E89B9C11E42B" authority="Constant & Pham" authorityName="Constant & Pham" authorityYear="2024" class="Insecta" family="Nogodinidae" genus="Goniopsarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniopsarites mientrunganus" order="Hemiptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="mientrunganus" status="sp. nov.">Goniopsarites mientrunganus Constant & Pham</taxonomicName>
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<taxonomicNameLabel id="2368D723D0EBCBE81C073D230ABF21F3" pageId="0" pageNumber="181">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="425EBDDC23CD9E00C3CC28DFEB278D1A" pageId="0" pageNumber="181" type="description">
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<paragraph id="2E9861C9CD3D3A46A3E5715C4532AB62" pageId="0" pageNumber="181">
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<figureCitation id="48E7F2E8A8938BFE64B8F26E44ED3EE7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">Figs 1</figureCitation>
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<figureCitation id="BDD7751729270BA8FDF7A7C315D1C098" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">, 2</figureCitation>
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<figureCitation id="4F7156586DEAC28A3E3704C5DD06CD93" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Goniopsarites mientrunganus Constant & Pham, sp. nov., in nature and habitat A-C Bach Ma National Park, low altitude, 15 May 2023 A adult on stem B habitat C nymph D-F Bach Ma National Park, Pheasant Trail, 20 May 2023 D habitat E adult on tree trunk F idem, general view G, H Phong Dien District, CCRR, 23 May 2023 G adult on stem H habitat." figureDoi="10.3897/zookeys.1193.114957.figure3" httpUri="https://binary.pensoft.net/fig/997499" pageId="0" pageNumber="181">, 3</figureCitation>
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<figureCitation id="DEF826B808ED50F05BE7D2F96C661355" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution maps A Goniopsarites species in Vietnam B Pisacha yinggensis Meng, Wang & Wang, 2014." figureDoi="10.3897/zookeys.1193.114957.figure4" httpUri="https://binary.pensoft.net/fig/997500" pageId="0" pageNumber="181">, 4A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="67203921396FCB708F338FE1879E35AE" pageId="0" pageNumber="181" type="type material">
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<paragraph id="47294E9855AA85DC6095A9825D7ECDD1" pageId="0" pageNumber="181">Type material.</paragraph>
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<paragraph id="40874415271993C1C70463BC89C7AFF0" pageId="0" pageNumber="181">
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<emphasis id="8153162F2BD70BB8A1196451F81FC05A" bold="true" italics="true" pageId="0" pageNumber="181">Holotype</emphasis>
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♂, Vietnam • Thừa
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<normalizedToken id="4FD4479659448440FE60B39B1DDDDE5E" originalValue="Thiên-Hu">Thien-Hu</normalizedToken>
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ế Province, Bach Ma National Park, near ranger station;
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<geoCoordinate id="B372FE2B2A0352CF6C8B5E8939E91F1E" degrees="16" direction="north" minutes="08" orientation="latitude" precision="15" seconds="37" value="16.143612">16°08'37"N</geoCoordinate>
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,
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<geoCoordinate id="0030EA6688A158BDD835E1A520D707C5" degrees="107" direction="east" minutes="49" orientation="longitude" precision="15" seconds="36" value="107.82667">107°49'36"E</geoCoordinate>
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; 18 May 2023; alt. 300-600 m; J. Constant & L. Semeraro leg.; VNMN_E000.017.000.
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</paragraph>
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<paragraph id="A97D2665453B4FCAF7B5A2590C3C317E" pageId="0" pageNumber="181">
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<emphasis id="47F58C6252C3A5081E18D8534755608F" bold="true" italics="true" pageId="0" pageNumber="181">Paratypes</emphasis>
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, Vietnam • 1 ♀; Thừa
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<normalizedToken id="375A95FA269D9D188A91B7E20E2FBB18" originalValue="Thiên-Hu">Thien-Hu</normalizedToken>
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ế Province, Bach Ma National Park;
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<geoCoordinate id="ED98A63CDA1FEC9C906A2FEF55F6E4A1" degrees="16" direction="north" minutes="13" orientation="latitude" precision="15" seconds="14" value="16.220556">16°13'14"N</geoCoordinate>
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,
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<geoCoordinate id="2F4ECCA9D0B47AE573EF885209F18028" degrees="107" direction="east" minutes="53" orientation="longitude" precision="15" seconds="10" value="107.88611">107°53'10"E</geoCoordinate>
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; 9 Mar. 2023; by net; V.T. Trung leg.; VNMN_ E000.017.001 • 1 ♂; Thừa
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<normalizedToken id="73E051D9554E1BD327243925C7FA5C26" originalValue="Thiên-Hu">Thien-Hu</normalizedToken>
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ế Province, Bach Ma National Park, Pheasant trail;
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<geoCoordinate id="CA7F60E3A8CB63E846BABAF146AAC594" degrees="16" direction="north" minutes="13" orientation="latitude" precision="15" seconds="38" value="16.227222">16°13'38"N</geoCoordinate>
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,
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<geoCoordinate id="0DB9875AED8FEA7FB9468F76AF505FAF" degrees="107" direction="east" minutes="51" orientation="longitude" precision="15" seconds="20" value="107.85555">107°51'20"E</geoCoordinate>
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; 10-20 May 2023; alt. 500-600 m; J. Constant & L. Semeraro leg.; I.G.: 34.640; RBINS • 2 ♂♂, 1 ♀; Thừa
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<normalizedToken id="FA9789696B76C965CED2A5FAFBFC3D72" originalValue="Thiên-Hu">Thien-Hu</normalizedToken>
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ế Province, Bach Ma National Park; low altitude;
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<geoCoordinate id="1355394EC7F4904B97814C1EE0E4D630" degrees="16" direction="north" minutes="13" orientation="latitude" precision="15" seconds="05" value="16.218056">16°13'05"N</geoCoordinate>
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,
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<geoCoordinate id="28A5463ECAD2ECCCAAB25B89CBF57298" degrees="107" direction="east" minutes="42" orientation="longitude" precision="15" seconds="27" value="107.7075">107°42'27"E</geoCoordinate>
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; 17 May 2023; alt. 100-200 m; J. Constant & L. Semeraro leg.; I.G.: 34.640; RBINS • 1 ♀; Thừa
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<normalizedToken id="DA1C6BDBDE2689FED5A8654284E9AFFE" originalValue="Thiên-Hu">Thien-Hu</normalizedToken>
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ế Province; Phong Dien District, CCRR;
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<geoCoordinate id="C85855B83CCB9DF231EB226512569AD1" degrees="16" direction="north" minutes="30" orientation="latitude" precision="15" seconds="27" value="16.5075">16°30'27"N</geoCoordinate>
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,
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<geoCoordinate id="72C5A36077006A06F4CB516C2DFC1D00" degrees="107" direction="east" minutes="16" orientation="longitude" precision="15" seconds="05" value="107.26806">107°16'05"E</geoCoordinate>
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; 23 May 2023; alt. 350-400 m; J. Constant & L. Semeraro leg.; I.G.: 34.640; RBINS • 1 ♀; same collection data as for preceding; VNMN_E000.017.002.
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</paragraph>
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</subSubSection>
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<subSubSection id="CCF7DAE12376C4835B34450DD611F180" pageId="0" pageNumber="181" type="diagnosis">
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<paragraph id="E0EBF754213A631296D541B02C0147DB" pageId="0" pageNumber="181">Diagnosis.</paragraph>
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<paragraph id="0FBA0AA6DFF2BB5CC62DF6779E0364AC" pageId="0" pageNumber="181">
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The species is very close externally to
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<taxonomicName id="22B6379F37BB6046757C024492EE6954" lsidName="G. fronticonvexus" pageId="0" pageNumber="181" rank="species" species="fronticonvexus">
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<emphasis id="E7BCD493A171FA4FAB7155247BB9CC21" italics="true" pageId="0" pageNumber="181">G. fronticonvexus</emphasis>
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</taxonomicName>
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Meng, Wang & Wang, 2014 and
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<taxonomicName id="80A236C2FFE09375B5104253151062CC" lsidName="G. tonkinensis" pageId="0" pageNumber="181" rank="species" species="tonkinensis">
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<emphasis id="E0DBDC76FF7D7E0C45708AD89CE6C6F7" italics="true" pageId="0" pageNumber="181">G. tonkinensis</emphasis>
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</taxonomicName>
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Constant & Pham, 2016 but has slightly more elongate tegmina, 2.2 times as long as wide (1.9 times in
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<taxonomicName id="3C1580EDDB533B4B45A7E920FE728F42" lsidName="G. fronticonvexus" pageId="0" pageNumber="181" rank="species" species="fronticonvexus">
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<emphasis id="048D398869E9ADEE210774AA23728DCE" italics="true" pageId="0" pageNumber="181">G. fronticonvexus</emphasis>
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</taxonomicName>
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; 2.0 times in
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<taxonomicName id="2E202AE6D8514DA6C461D43EFFAB3752" lsidName="G. tonkinensis" pageId="0" pageNumber="181" rank="species" species="tonkinensis">
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<emphasis id="865A28AE401C60244B45246C99176FD2" italics="true" pageId="0" pageNumber="181">G. tonkinensis</emphasis>
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</taxonomicName>
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). These species are better separated by the male genitalia characters as follows.
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</paragraph>
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<paragraph id="3A1B02FD73E654D79B931723A9DB2F01" pageId="0" pageNumber="181">
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<taxonomicName id="2A0D87C6F3360BE3743CB9029BBA51B9" authorityName="Constant & Pham" authorityYear="2024" class="Insecta" family="Nogodinidae" genus="Goniopsarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniopsarites mientrunganus" order="Hemiptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="mientrunganus">
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<emphasis id="D47D289BAB2E6C213DD63495C151A591" italics="true" pageId="0" pageNumber="181">Goniopsarites mientrunganus</emphasis>
|
|
</taxonomicName>
|
|
Constant & Pham, sp. nov. can be separated from the other two species by
|
|
<emphasis id="D3830D316E950C309D2A050B265D764A" bold="true" pageId="0" pageNumber="181">(1)</emphasis>
|
|
its large laminate posterior processes of the periandrium (
|
|
<emphasis id="2666CD879A030D4A4A8E91B8E2C467DD" italics="true" pageId="0" pageNumber="181">ppp</emphasis>
|
|
- Fig.
|
|
<figureCitation id="029A2FC4A84593BAAF7F7B45554606D9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">2D</figureCitation>
|
|
), which are absent in
|
|
<taxonomicName id="59139807ACC8B4350C1D1E3267884C33" lsidName="G. fronticonvexus" pageId="0" pageNumber="181" rank="species" species="fronticonvexus">
|
|
<emphasis id="DBBEBC184CB4767DB9C902F87237AEE3" italics="true" pageId="0" pageNumber="181">G. fronticonvexus</emphasis>
|
|
</taxonomicName>
|
|
and much more slender, curved the other way round, in
|
|
<taxonomicName id="E0918EC731E7CD9EC42166EB87B36A2F" lsidName="G. tonkinensis" pageId="0" pageNumber="181" rank="species" species="tonkinensis">
|
|
<emphasis id="0341F215508D9C691CB74DD86B9A9900" italics="true" pageId="0" pageNumber="181">G. tonkinensis</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="105E0435C0AA0D7C3269D70060F40117" bold="true" pageId="0" pageNumber="181">(2)</emphasis>
|
|
the posterodorsal process of the gonostyli bulging apically (
|
|
<emphasis id="D7CF646A272B8C01D069A218A146B1FF" italics="true" pageId="0" pageNumber="181">G</emphasis>
|
|
- Fig.
|
|
<figureCitation id="BD4C2B925B640D9CFF81A07F3222D40A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">2A</figureCitation>
|
|
), not bulging in both other species; and
|
|
<emphasis id="459D7B4F9DB1EEDF5F30A17EC1D4BEDF" bold="true" pageId="0" pageNumber="181">(3)</emphasis>
|
|
the strongly developed basiventral lobe of the proximal half of the anal tube (
|
|
<emphasis id="2A06B84FFD1946A0E0687AAA7FD6386E" italics="true" pageId="0" pageNumber="181">An</emphasis>
|
|
- Fig.
|
|
<figureCitation id="A5052D49F72F692E40F9593F8C8F8804" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">2A</figureCitation>
|
|
), clearly less developed in both other species.
|
|
</paragraph>
|
|
<paragraph id="C74B1908BC6350D75EC4255DD9F3342E" pageId="0" pageNumber="181">
|
|
From
|
|
<taxonomicName id="D8ACB829EB61F2DAFE8787BBAFE9ECFF" lsidName="G. tonkinensis" pageId="0" pageNumber="181" rank="species" species="tonkinensis">
|
|
<emphasis id="3C2103E6603E2A0FCB0F0DF87AB6F571" italics="true" pageId="0" pageNumber="181">G. tonkinensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="49F4503CAE831024E3B36134891381EC" lsidName="G. mientrunganus" pageId="0" pageNumber="181" rank="species" species="mientrunganus">
|
|
<emphasis id="97E0D89A3C7AFF24909B826467595DE0" italics="true" pageId="0" pageNumber="181">G. mientrunganus</emphasis>
|
|
</taxonomicName>
|
|
Constant & Pham, sp. nov. also differs in
|
|
<emphasis id="4D9C4525411D2F4FFBE748E35DD11D30" bold="true" pageId="0" pageNumber="181">(1)</emphasis>
|
|
having the maximum width of the anal tube in the basal third (
|
|
<emphasis id="55685B8187FC8946C3C61C9502982CFB" italics="true" pageId="0" pageNumber="181">An</emphasis>
|
|
- Fig.
|
|
<figureCitation id="B540EBDF3BBA138A617AD5A42E8DAC6A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">2C</figureCitation>
|
|
), which is in the distal half of the anal tube in
|
|
<taxonomicName id="F328431CC40601BD827CE3F22D527B6B" lsidName="G. tonkinensis" pageId="0" pageNumber="181" rank="species" species="tonkinensis">
|
|
<emphasis id="2F030938C855E979DE8F1AA5423CE837" italics="true" pageId="0" pageNumber="181">G. tonkinensis</emphasis>
|
|
</taxonomicName>
|
|
(the state of this character is not known for
|
|
<taxonomicName id="A9DCB880FEF94E2EA2E86B8F0512BDDE" lsidName="G. fronticonvexus" pageId="0" pageNumber="181" rank="species" species="fronticonvexus">
|
|
<emphasis id="F2E2FF8678E49AD2EC85C94F06B470C6" italics="true" pageId="0" pageNumber="181">G. fronticonvexus</emphasis>
|
|
</taxonomicName>
|
|
);
|
|
<emphasis id="E6E34ECEA8F575E484A84EAAF1184C02" bold="true" pageId="0" pageNumber="181">(2)</emphasis>
|
|
in having a strongly developed dorsal laminate process along basal half of periandrium (
|
|
<emphasis id="2A08C3453567DC91F54CC769F42BA650" italics="true" pageId="0" pageNumber="181">dlp</emphasis>
|
|
- Fig.
|
|
<figureCitation id="F32A5B41E3C9928CCC0E919F0A511B4D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">2D, F, G</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="869DE46AB66667A7337701187516E53A" pageId="0" pageNumber="181" type="description">
|
|
<paragraph id="1F9B964F4B9BD60FAF581C328391843F" pageId="0" pageNumber="181">Description.</paragraph>
|
|
<paragraph id="6F6191DAE32B999BECF44607027E173F" pageId="0" pageNumber="181">
|
|
<emphasis id="F4E1F79DC280C7D36C809E6E8AFECD1C" bold="true" italics="true" pageId="0" pageNumber="181">Measurements and ratios</emphasis>
|
|
: LT: ♂ (
|
|
<emphasis id="CD023DB20A4A07B7BA6F21A62D24E8FB" italics="true" pageId="0" pageNumber="181">n</emphasis>
|
|
= 4): 11.3 mm (11. 1-11.7); ♀ (
|
|
<emphasis id="867948B7744E2067CAE610F062945D17" italics="true" pageId="0" pageNumber="181">n</emphasis>
|
|
= 4): 13.5 (13. 0-14.5). LTg/BTg = 2.2; LW/BW = 1.4; LV/BV = 8.5; LF/BF = 1.4.
|
|
</paragraph>
|
|
<paragraph id="2531321EC190D5C3F6B644D69E90DF33" pageId="0" pageNumber="181">
|
|
<emphasis id="D7C754A8F70403B50EDD6EDFEBF72D02" bold="true" italics="true" pageId="0" pageNumber="181">Head</emphasis>
|
|
: (Fig.
|
|
<figureCitation id="FB2D74E405CB19074175FDD93C7E9C6E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">1A-E</figureCitation>
|
|
) vertex dark brown, with yellowish median and lateral carinae; concave with lateral margins carinate, and with anterior and posterior margins slightly carinate and rather strongly concave in dorsal view, resulting in a very narrow central portion. Frons varying from variegated brown and yellowish, darker under V-shaped carina, carina yellowish to reddish, to nearly completely black-brown, sometimes with yellowish markings around mid-height of lateral margin; narrow, well-defined, yellowish line above frontoclypeal suture; frons elongate, broader dorsally, concave with median carina on middle of disc and with a strong projection on ventral half marked by strong, V-shaped carina extending ventrad on clypeus; projection rounded in lateral view; lateral margins of frons carinate; frontoclypeal suture grooved and rounded ventrally. Genae largely variegated yellow-brown, dark brown under antennae, with pale yellowish spot along anterior margin. Clypeus yellowish, variegated with black-brown on sides towards apex, and with brown oblique lines on each side; clypeus elongate, narrower and shorter than frons, and showing a strong median carina; roundly convex in lateral view. Labium elongate and narrow, yellow-brown, slightly surpassing posterior coxae. Antennae black with scape ring-shaped; pedicel subglobose.
|
|
</paragraph>
|
|
<paragraph id="F19A6C4D468B6ADF3653201D4E4CFA32" pageId="0" pageNumber="181">
|
|
<emphasis id="E036B872B1A130004A2BC22E4A8EEDFC" bold="true" italics="true" pageId="0" pageNumber="181">Thorax</emphasis>
|
|
: (Fig.
|
|
<figureCitation id="68198B84AF4BEC2848D23CCD91BDE53D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">1A, C-E</figureCitation>
|
|
) pronotum variegated yellow-brown, darker in middle portion, rather densely pitted, and with median carina weakly marked; lateral fields of prothorax coloured as pronotum, darker in ventral portion, widening ventrally and with ventral margin rounded. Mesonotum dark brown, with yellowish markings on sides and yellowish spot on scutellum; lateral fields moderately pitted; median carina obsolete; lateral carinae marked with yellowish, fusing anteriorly in a rounded carina parallel to posterior margin of pronotum. Tegulae brown.
|
|
</paragraph>
|
|
<caption id="210B4D33081D223611E547CA4A4F8350" doi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181" start="Figure 1" startId="F1">
|
|
<paragraph id="AE5FF4B7788C236D40A47EEE08C7F3B1" pageId="0" pageNumber="181">
|
|
<emphasis id="B92C4E4040C4EC175FD203E43F8329EB" bold="true" pageId="0" pageNumber="181">Figure 1.</emphasis>
|
|
<taxonomicName id="A3365AB8693DCB525A73E9CB190213BC" authorityName="Constant & Pham" authorityYear="2024" class="Insecta" family="Nogodinidae" genus="Goniopsarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniopsarites mientrunganus" order="Hemiptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="mientrunganus">
|
|
<emphasis id="A707F65FD5EE4BA3A49E6358BA1C6B1A" italics="true" pageId="0" pageNumber="181">Goniopsarites mientrunganus</emphasis>
|
|
</taxonomicName>
|
|
Constant & Pham, sp. nov.
|
|
<emphasis id="22672D6C0809101E9A7D54AEF9986272" bold="true" pageId="0" pageNumber="181">A-F</emphasis>
|
|
holotype ♂
|
|
<emphasis id="59006F48E6D394EB2EDB266E568F9F50" bold="true" pageId="0" pageNumber="181">A</emphasis>
|
|
habitus, dorsal
|
|
<emphasis id="FD1D60EA3DEFA2CFD54F68AF282D5484" bold="true" pageId="0" pageNumber="181">B</emphasis>
|
|
habitus, ventral
|
|
<emphasis id="4BEE5D4AC313153263CD4D4E9BA6D258" bold="true" pageId="0" pageNumber="181">C</emphasis>
|
|
habitus, lateral
|
|
<emphasis id="C7BA5D5AE1E9EBA159A886D9AF4202C6" bold="true" pageId="0" pageNumber="181">D</emphasis>
|
|
habitus, perpendicular view of frons
|
|
<emphasis id="A87372BF7716C4E9D716383C8EB427CD" bold="true" pageId="0" pageNumber="181">E</emphasis>
|
|
habitus, anterolateral
|
|
<emphasis id="663BBC228F501D2A2395AE9754618F31" bold="true" pageId="0" pageNumber="181">F</emphasis>
|
|
right hind wing
|
|
<emphasis id="05D2C11E80CE5506B84469FA67FF4947" bold="true" pageId="0" pageNumber="181">G</emphasis>
|
|
paratypes ♀♀, posterior margin of sternite VII.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="1B0A1B1C7B957133937E24E09D1D2DDE" pageId="0" pageNumber="181">
|
|
<emphasis id="09D0EFA4FB6053273351DDA7268A1F79" bold="true" italics="true" pageId="0" pageNumber="181">Tegmina</emphasis>
|
|
: (Fig.
|
|
<figureCitation id="99C240A2308950FCBCD57283DF46919E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">1A-C</figureCitation>
|
|
) elongate, with costal margin broadly rounded on proximal half and rather strongly sinuate on posterior half; brown with large, irregular, black-brown marking on basal half, not extending on clavus; large, pale yellowish marking along costal margin in distal half; clavus with irregular, slightly darker markings on posterior half; hypocostal plate narrow, visible in proximal third; posterior margin obliquely rounded; clavus closed, extending to posterior angle of tegmen. Veins reddish brown, sometimes black in black areas.
|
|
</paragraph>
|
|
<paragraph id="EB56A473DF8EF175C5178DF4C9013C0C" pageId="0" pageNumber="181">
|
|
<emphasis id="A597C2777D9E0089147F0AA0762232F3" bold="true" italics="true" pageId="0" pageNumber="181">Hind wings</emphasis>
|
|
: (Fig.
|
|
<figureCitation id="15F0014DBD2F223C16D3018D02058B4A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">1F</figureCitation>
|
|
) brown, darker towards posterocostal angle and with large basicostal yellow-brown area; well developed, with posterior margin trilobed; costal margin sinuate, with coupling apparatus at 2/3 of length.
|
|
</paragraph>
|
|
<paragraph id="280A2D750A97AC36B1D5B19776816D5D" pageId="0" pageNumber="181">
|
|
<emphasis id="A635CE78A65561D3B729DFF75CA5E7B1" bold="true" italics="true" pageId="0" pageNumber="181">Legs</emphasis>
|
|
: (Fig.
|
|
<figureCitation id="B3B3F29AD29E7F1CBAF371369F4CE02A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">1A-C</figureCitation>
|
|
) profemora black-brown, with narrow, pale yellow rings; mesofemora pale yellow, with brown rings; pro- and mesotibiae elongate and slender, with 6 rings alternatively pale yellow and brown, basal one brown, apical one yellow; metafemora yellow, with brown markings; metatibiae rather short, broadening towards apex, pale yellow, infuscate basally, with 2 strong lateral spines near apex. Tarsi brown, darker apically. Metatibiotarsal formula: (2) 10 / 2 (+7 on underside) / 2.
|
|
</paragraph>
|
|
<paragraph id="576DBB1AF46FD7148EF6C8C6B9C97FDB" pageId="0" pageNumber="181">
|
|
<emphasis id="4AB9DF715BA15F840F439FD18D76FF79" bold="true" italics="true" pageId="0" pageNumber="181">Abdomen</emphasis>
|
|
: (Fig.
|
|
<figureCitation id="FABCB6584131E57AD19ABDFD2E5B3A09" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">1B</figureCitation>
|
|
) pale yellowish brown.
|
|
</paragraph>
|
|
<paragraph id="2E150AC42199F9126028EC262380EAC1" pageId="0" pageNumber="181">
|
|
<emphasis id="3A823884093AAAAD64F91F06165620DB" bold="true" italics="true" pageId="0" pageNumber="181">Terminalia</emphasis>
|
|
♂: (Fig.
|
|
<figureCitation id="654975EE4A3DFA142C3226B2213BFD98" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">2</figureCitation>
|
|
) pygofer (
|
|
<emphasis id="C75806CC3E1A1B340518072D05AD3EB7" italics="true" pageId="0" pageNumber="181">Py</emphasis>
|
|
) higher than long in lateral view, with anterior and posterior margins moderately sinuate; posterodorsal angles with dorsally developed, subtriangular laminate process, with posterodorsal angle right and rounded. Anal tube (
|
|
<emphasis id="99F84FF21173A6D1C28633959E8444AA" italics="true" pageId="0" pageNumber="181">An</emphasis>
|
|
) massive, elongate, with lateral margins in dorsal view, widely rounded in proximal half, then sinuately tapering towards posterior, leaving widely rounded ventral margin visible on sides in distal half; 1.78 times as long in midline, as wide with maximum width at proximal 1/3 in dorsal view; basal half in lateral view with large ventral lobe; strongly curved ventrally at midlength with lateral margins strongly produced ventrally into broad lobe in distal half. Gonostyli (
|
|
<emphasis id="EC1CC705EB99FD02755B9B2AC1F88B59" italics="true" pageId="0" pageNumber="181">G</emphasis>
|
|
) (Fig.
|
|
<figureCitation id="055424A655949CE5CB7C095C30690A6C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution maps A Goniopsarites species in Vietnam B Pisacha yinggensis Meng, Wang & Wang, 2014." figureDoi="10.3897/zookeys.1193.114957.figure4" httpUri="https://binary.pensoft.net/fig/997500" pageId="0" pageNumber="181">4A</figureCitation>
|
|
) elongate in lateral view, rounded apically and with strong process projecting dorsomesad at posterodorsal angle; ventral margin nearly straight and posterior margin weakly sinuate in lateral view; dorsal process slightly twisted internally, bulging in distal portion and with apex with blunt point directed anteromesad. Aedeagus (Fig.
|
|
<figureCitation id="FEBE480641404E187807A848D1952B0F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goniopsarites mientrunganus Constant & Pham, sp. nov., holotype ♂, terminalia A-C pygofer, gonostyli and anal tube A lateral B caudal C dorsal D-I aedeagus, phallobase and connective D left lateral E dorsal F lateroventral G laterodorsal H caudal I ventral. cv = connective; dlp = dorsal laminate process of periandrium; lpa = lateral process of aedeagus; ppp = posterior process of periandrium; spp = sclerotized process of phallus; td = tectiductus; vlp = ventral lobe of periandrium." figureDoi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181">2D-I</figureCitation>
|
|
) strongly curved in lateral view, with a pair of lateral, elongate, strongly sinuate processes (
|
|
<emphasis id="603637E3E73E48B9B304A008DEDE6876" italics="true" pageId="0" pageNumber="181">lpa</emphasis>
|
|
) directed anteriorly, attached posteriorly at 2/3 of length; processes well visible and sinuate in ventral view. Periandrium posteriorly with a pair of large laminate processes (
|
|
<emphasis id="2C6B9AB132B113B5F479BF55DFB0B69C" italics="true" pageId="0" pageNumber="181">ppp</emphasis>
|
|
), lobe-shaped and developed dorsally into sinuate, elongate, narrowly pointed processes directed mesodorsad; well-developed laminate process (
|
|
<emphasis id="AAC8AEFD9008327DF6DE7D221F71FFF2" italics="true" pageId="0" pageNumber="181">dlp</emphasis>
|
|
) running dorsally in basal half, abruptly terminated at right angle at midlength. Ventral lobe of periandrium (
|
|
<emphasis id="6A7324549A5D048EBD582E0B0EA2A7FA" italics="true" pageId="0" pageNumber="181">vlp</emphasis>
|
|
) elongate, lanceolate in distal portion in ventral view, with sides rounded. Phallus membranous, large, with pair of curved, sclerotized processes (
|
|
<emphasis id="6DC23436512D6A53D23EFBDED308060A" italics="true" pageId="0" pageNumber="181">spp</emphasis>
|
|
) directed cephalodorsad before apex; strongly, angularly projecting posterodorsad; apex narrowing, directed anteriorly, and curved ventrally. Connective (
|
|
<emphasis id="29E501DB3660DC4432BF088B250938B0" italics="true" pageId="0" pageNumber="181">cv</emphasis>
|
|
) with well-developed, elongate tectiductus (
|
|
<emphasis id="0AB196E3F2EBA25B64BB29387A92876D" italics="true" pageId="0" pageNumber="181">td</emphasis>
|
|
) showing complete, broadly rounded dorsal crista.
|
|
</paragraph>
|
|
<caption id="E43CFD4555761387AD8118FA2389CFE0" doi="10.3897/zookeys.1193.114957.figure2" httpUri="https://binary.pensoft.net/fig/997498" pageId="0" pageNumber="181" start="Figure 2" startId="F2">
|
|
<paragraph id="76BACA28E690C3360428D1576A3FC5C4" pageId="0" pageNumber="181">
|
|
<emphasis id="6B76E4A5BCA387A28B2844B4D3374635" bold="true" pageId="0" pageNumber="181">Figure 2.</emphasis>
|
|
<taxonomicName id="225EE910AF83A51977DB0C8660BFA111" authorityName="Constant & Pham" authorityYear="2024" class="Insecta" family="Nogodinidae" genus="Goniopsarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniopsarites mientrunganus" order="Hemiptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="mientrunganus">
|
|
<emphasis id="67707D1034968A4AF3C2D8C7CD6422EE" italics="true" pageId="0" pageNumber="181">Goniopsarites mientrunganus</emphasis>
|
|
</taxonomicName>
|
|
Constant & Pham, sp. nov., holotype ♂, terminalia
|
|
<emphasis id="39EA174421E326B5753C4BD9D274014F" bold="true" pageId="0" pageNumber="181">A-C</emphasis>
|
|
pygofer, gonostyli and anal tube
|
|
<emphasis id="DC246744B32C354042ADDF94FF3EFC11" bold="true" pageId="0" pageNumber="181">A</emphasis>
|
|
lateral
|
|
<emphasis id="AD2B6932AB1B6A6F00D4CDE8E7EB1B93" bold="true" pageId="0" pageNumber="181">B</emphasis>
|
|
caudal
|
|
<emphasis id="5E11C4E87991118AEEA60FC8A4D67F68" bold="true" pageId="0" pageNumber="181">C</emphasis>
|
|
dorsal
|
|
<emphasis id="5BBE1398E0B770070868C5421BE8572F" bold="true" pageId="0" pageNumber="181">D-I</emphasis>
|
|
aedeagus, phallobase and connective
|
|
<emphasis id="013EA42A34951D50414B71473E80FCC1" bold="true" pageId="0" pageNumber="181">D</emphasis>
|
|
left lateral
|
|
<emphasis id="2BA89253D120038EA071941AE6848428" bold="true" pageId="0" pageNumber="181">E</emphasis>
|
|
dorsal
|
|
<emphasis id="DF5BEB13CABD8A48FA3401CE54C935DA" bold="true" pageId="0" pageNumber="181">F</emphasis>
|
|
lateroventral
|
|
<emphasis id="327268C49B7DED0386C2D586B524BCE6" bold="true" pageId="0" pageNumber="181">G</emphasis>
|
|
laterodorsal
|
|
<emphasis id="7CBBC7E6D5962927E75D8D5F76D5901D" bold="true" pageId="0" pageNumber="181">H</emphasis>
|
|
caudal
|
|
<emphasis id="E188D549B1D03FD143D4AE26BA254266" bold="true" pageId="0" pageNumber="181">I</emphasis>
|
|
ventral.
|
|
<emphasis id="A41D1D53A6FFC16B09FF494F8E5AF3BA" italics="true" pageId="0" pageNumber="181">cv</emphasis>
|
|
= connective;
|
|
<emphasis id="5BAA936B15AFBA1B815DE2CB3D681186" italics="true" pageId="0" pageNumber="181">dlp</emphasis>
|
|
= dorsal laminate process of periandrium;
|
|
<emphasis id="07FB9FDD799C1F6FD51E7FBA6CFEF262" italics="true" pageId="0" pageNumber="181">lpa</emphasis>
|
|
= lateral process of aedeagus;
|
|
<emphasis id="3C5CC1BA80ABD0D86F281A086ECA9E65" italics="true" pageId="0" pageNumber="181">ppp</emphasis>
|
|
= posterior process of periandrium;
|
|
<emphasis id="125B5976DD6815D37EEE5079B4234288" italics="true" pageId="0" pageNumber="181">spp</emphasis>
|
|
= sclerotized process of phallus;
|
|
<emphasis id="5A574065A95145CF4F30EDBDFB7562D1" italics="true" pageId="0" pageNumber="181">td</emphasis>
|
|
= tectiductus;
|
|
<emphasis id="8C218847B16BEF3F9DE9CC80780ADC42" italics="true" pageId="0" pageNumber="181">vlp</emphasis>
|
|
= ventral lobe of periandrium.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="4E021D8115DC64598206B196E7202CCE" pageId="0" pageNumber="181" type="notes">
|
|
<paragraph id="04F10DBC3B9D0D738AB7F0B93F6244B8" pageId="0" pageNumber="181">Note.</paragraph>
|
|
<paragraph id="C4EAE4072D7A4204AC6D13796A2C5A32" pageId="0" pageNumber="181">
|
|
The female genital structures were checked and found to be similar to those of
|
|
<taxonomicName id="EB4AE28179429FFECF49FAEFD5A66E50" lsidName="G. fronticonvexus" pageId="0" pageNumber="181" rank="species" species="fronticonvexus">
|
|
<emphasis id="2484E2B32FDEB2B13FD0454E403AD523" italics="true" pageId="0" pageNumber="181">G. fronticonvexus</emphasis>
|
|
</taxonomicName>
|
|
, as well as to these of
|
|
<taxonomicName id="0BB5E968166D123B05E075E2EED1B41D" lsidName="G. tonkinensis" pageId="0" pageNumber="181" rank="species" species="tonkinensis">
|
|
<emphasis id="421F9992C7AA067ECC29FB5E96E42A53" italics="true" pageId="0" pageNumber="181">G. tonkinensis</emphasis>
|
|
</taxonomicName>
|
|
, and, as it is the case in most nogodinids that we have observed so far, female genitalia do not help with species identification. The indentation in middle of the hind margin of sternum VII is always present but was found to vary in depth between the specimens of
|
|
<taxonomicName id="EA1DC17B199F982E6A9CE659D42AC962" lsidName="G. mientrunganus" pageId="0" pageNumber="181" rank="species" species="mientrunganus">
|
|
<emphasis id="700830C472203695C1258EB5E7530B47" italics="true" pageId="0" pageNumber="181">G. mientrunganus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. that we have in hand (Fig.
|
|
<figureCitation id="38DFE31EDEEFBFB9945E5A47C0AC4AD4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Goniopsarites mientrunganus Constant & Pham, sp. nov. A-F holotype ♂ A habitus, dorsal B habitus, ventral C habitus, lateral D habitus, perpendicular view of frons E habitus, anterolateral F right hind wing G paratypes ♀♀, posterior margin of sternite VII." figureDoi="10.3897/zookeys.1193.114957.figure1" httpUri="https://binary.pensoft.net/fig/997497" pageId="0" pageNumber="181">1G</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="666B356275D5B87363091BC23876E1C3" pageId="0" pageNumber="181" type="etymology">
|
|
<paragraph id="4F0ED3692494C5286C3117FAAD1A7E78" pageId="0" pageNumber="181">Etymology.</paragraph>
|
|
<paragraph id="80684E5A6A80E44FB3CD4F603D59B422" pageId="0" pageNumber="181">
|
|
The species epithet
|
|
<emphasis id="AD3F4CF80CE5A619EE6BFD25F8CC4568" italics="true" pageId="0" pageNumber="181">mientrunganus</emphasis>
|
|
refers to the region where the new species was discovered: Central Vietnam, "Miền Trung" in Vietnamese.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="284AFCEF13C27302D137B57BBCD14CD2" pageId="0" pageNumber="181" type="biology_ecology">
|
|
<paragraph id="AA8DDED065412B829C8821331AFD32A7" pageId="0" pageNumber="181">Biology.</paragraph>
|
|
<paragraph id="A137DED70A25CAEDA9FA1F5675826208" pageId="0" pageNumber="181">
|
|
The specimens were found sitting on stems of bushes (Fig.
|
|
<figureCitation id="0B3123B46BB220E91771C0A8E82E6C42" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Goniopsarites mientrunganus Constant & Pham, sp. nov., in nature and habitat A-C Bach Ma National Park, low altitude, 15 May 2023 A adult on stem B habitat C nymph D-F Bach Ma National Park, Pheasant Trail, 20 May 2023 D habitat E adult on tree trunk F idem, general view G, H Phong Dien District, CCRR, 23 May 2023 G adult on stem H habitat." figureDoi="10.3897/zookeys.1193.114957.figure3" httpUri="https://binary.pensoft.net/fig/997499" pageId="0" pageNumber="181">3A, C, G</figureCitation>
|
|
) or on tree trunks (Fig.
|
|
<figureCitation id="02D71BEBA9D10608C097643906D76E8C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Goniopsarites mientrunganus Constant & Pham, sp. nov., in nature and habitat A-C Bach Ma National Park, low altitude, 15 May 2023 A adult on stem B habitat C nymph D-F Bach Ma National Park, Pheasant Trail, 20 May 2023 D habitat E adult on tree trunk F idem, general view G, H Phong Dien District, CCRR, 23 May 2023 G adult on stem H habitat." figureDoi="10.3897/zookeys.1193.114957.figure3" httpUri="https://binary.pensoft.net/fig/997499" pageId="0" pageNumber="181">3E, F</figureCitation>
|
|
), in subtropical evergreen forest (Fig.
|
|
<figureCitation id="6C8719987E75A96798943771F4651BAC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Goniopsarites mientrunganus Constant & Pham, sp. nov., in nature and habitat A-C Bach Ma National Park, low altitude, 15 May 2023 A adult on stem B habitat C nymph D-F Bach Ma National Park, Pheasant Trail, 20 May 2023 D habitat E adult on tree trunk F idem, general view G, H Phong Dien District, CCRR, 23 May 2023 G adult on stem H habitat." figureDoi="10.3897/zookeys.1193.114957.figure3" httpUri="https://binary.pensoft.net/fig/997499" pageId="0" pageNumber="181">3B, D, H</figureCitation>
|
|
) at the junction of the Northern Vietnam lowland rain forests, Southern Vietnam lowland rain forests, and Southern Annamites montane rain forests ecoregions, at rather low altitude (150-600 m). Some nymphs (Fig.
|
|
<figureCitation id="E6FC3B910181B66E1E377F4CD7D3C044" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Goniopsarites mientrunganus Constant & Pham, sp. nov., in nature and habitat A-C Bach Ma National Park, low altitude, 15 May 2023 A adult on stem B habitat C nymph D-F Bach Ma National Park, Pheasant Trail, 20 May 2023 D habitat E adult on tree trunk F idem, general view G, H Phong Dien District, CCRR, 23 May 2023 G adult on stem H habitat." figureDoi="10.3897/zookeys.1193.114957.figure3" httpUri="https://binary.pensoft.net/fig/997499" pageId="0" pageNumber="181">3C</figureCitation>
|
|
) were observed in May 2023 together with adult specimens.
|
|
</paragraph>
|
|
<caption id="19556DDEBD2E27BFE3C128748FC17CD2" doi="10.3897/zookeys.1193.114957.figure3" httpUri="https://binary.pensoft.net/fig/997499" pageId="0" pageNumber="181" start="Figure 3" startId="F3">
|
|
<paragraph id="06A17A4806AC1AFE14B75B983ECBF5B2" pageId="0" pageNumber="181">
|
|
<emphasis id="2C51CA44AF319D4601933D7647B0E703" bold="true" pageId="0" pageNumber="181">Figure 3.</emphasis>
|
|
<taxonomicName id="32C912CA388D0867F7349256F1F9324D" authorityName="Constant & Pham" authorityYear="2024" class="Insecta" family="Nogodinidae" genus="Goniopsarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniopsarites mientrunganus" order="Hemiptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="mientrunganus">
|
|
<emphasis id="D21E9D6A3D4DFA5492368D1561D04945" italics="true" pageId="0" pageNumber="181">Goniopsarites mientrunganus</emphasis>
|
|
</taxonomicName>
|
|
Constant & Pham, sp. nov., in nature and habitat
|
|
<emphasis id="8C1313131F5F2C594B6E2643AE302EF7" bold="true" pageId="0" pageNumber="181">A-C</emphasis>
|
|
Bach Ma National Park, low altitude, 15 May 2023
|
|
<emphasis id="0CAC3CF968369106DB263409C355072A" bold="true" pageId="0" pageNumber="181">A</emphasis>
|
|
adult on stem
|
|
<emphasis id="3B0DD13E9F2FD68E7DB1E3904F0A23D2" bold="true" pageId="0" pageNumber="181">B</emphasis>
|
|
habitat
|
|
<emphasis id="57B3943C1BBCB1468A355A44FD34CD20" bold="true" pageId="0" pageNumber="181">C</emphasis>
|
|
nymph
|
|
<emphasis id="1E95ED57163372C78B728F15427EF4BA" bold="true" pageId="0" pageNumber="181">D-F</emphasis>
|
|
Bach Ma National Park, Pheasant Trail, 20 May 2023
|
|
<emphasis id="2680C1C74E75902D18A0F4C6E09D0D1B" bold="true" pageId="0" pageNumber="181">D</emphasis>
|
|
habitat
|
|
<emphasis id="F5C53092C8E8971077EF71E90027B10B" bold="true" pageId="0" pageNumber="181">E</emphasis>
|
|
adult on tree trunk
|
|
<emphasis id="48D5338EBAD503306065DD024724909A" bold="true" pageId="0" pageNumber="181">F</emphasis>
|
|
idem, general view
|
|
<emphasis id="2BCF18631D6407D0593FD66155BB8E5C" bold="true" pageId="0" pageNumber="181">G, H</emphasis>
|
|
Phong Dien District, CCRR, 23 May 2023
|
|
<emphasis id="F4245BE244C1E933251EA4B81C96D4E7" bold="true" pageId="0" pageNumber="181">G</emphasis>
|
|
adult on stem
|
|
<emphasis id="DBCDB13E4719F3C75F8EAC8592E6F1C0" bold="true" pageId="0" pageNumber="181">H</emphasis>
|
|
habitat.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="FBBFE5E5F58A77D7F3C135C16C25B03F" pageId="0" pageNumber="181" type="distribution">
|
|
<paragraph id="A794E20EA73671E95B93767B8FFD8416" pageId="0" pageNumber="181">Distribution.</paragraph>
|
|
<paragraph id="B8913D401D00B115A617445040EDD087" pageId="0" pageNumber="181">
|
|
Vietnam, Thua Tinh-Hue Province, Bach Ma National Park, and Phong Dien district, CCRR (Fig.
|
|
<figureCitation id="2A8DBD5C52B75DFE1D5761E010993E6C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution maps A Goniopsarites species in Vietnam B Pisacha yinggensis Meng, Wang & Wang, 2014." figureDoi="10.3897/zookeys.1193.114957.figure4" httpUri="https://binary.pensoft.net/fig/997500" pageId="0" pageNumber="181">4A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="118EAD2A6F11F1274438B32043179B81" doi="10.3897/zookeys.1193.114957.figure4" httpUri="https://binary.pensoft.net/fig/997500" pageId="0" pageNumber="181" start="Figure 4" startId="F4">
|
|
<paragraph id="89549DFE28B1BE33030518B0FADFA0A3" pageId="0" pageNumber="181">
|
|
<emphasis id="144E21A702472D0A348E7568F9196159" bold="true" pageId="0" pageNumber="181">Figure 4.</emphasis>
|
|
Distribution maps
|
|
<emphasis id="20BD8E845681B328592FE7B55DFA5B48" bold="true" pageId="0" pageNumber="181">A</emphasis>
|
|
<taxonomicName id="B07111C5B508A1944C5E34649C358595" authorityName="Meng, Wang & Wang" authorityYear="2014" class="Insecta" family="Nogodinidae" genus="Goniopsarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniopsarites" order="Hemiptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B3987E8B06104CA10CA09EE1BA9282C3" italics="true" pageId="0" pageNumber="181">Goniopsarites</emphasis>
|
|
</taxonomicName>
|
|
species in Vietnam
|
|
<emphasis id="6A19F48F0F00A9BFAEE34467195198C8" bold="true" pageId="0" pageNumber="181">B</emphasis>
|
|
<taxonomicName id="4E2CEA79625BD28E41E75E944E82119F" authorityName="Meng, Wang & Wang" authorityYear="2014" class="Insecta" family="Nogodinidae" genus="Pisacha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pisacha yinggensis" order="Hemiptera" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="yinggensis">
|
|
<emphasis id="55ABF03FF2A1CF048BCB72EF6B7D19B7" italics="true" pageId="0" pageNumber="181">Pisacha yinggensis</emphasis>
|
|
</taxonomicName>
|
|
Meng, Wang & Wang, 2014.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |