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<mods:title id="D54201A783624D488E6A3A3693C0F348">A taxonomic review of Sueus Murayama, 1951 ambrosia beetles (Coleoptera: Curculionidae: Scolytinae: Hyorrhynchini) aided by molecular phylogenetic analyses</mods:title>
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<mods:affiliation id="039D719A2B8E956EBEFE6A9BDFAB93B5">Department of Entomology, Michigan State University, 288 Farm Lane, 243 Natural Science Bldg, East Lansing, MI 48824, USA.</mods:affiliation>
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<taxonomicName id="4C3A4D52F273FFF8878BFCF9FE5EFCCB" ID-CoL="8HRLR" authority="Murayama, 1951" authorityName="Murayama" authorityYear="1951" box="[151,446,826,853]" class="Insecta" family="Curculionidae" genus="Sueus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC3F273FFF8878BFCF9FE5EFCCB" bold="true" box="[151,446,826,853]" pageId="7" pageNumber="482">
<emphasis id="B94EEAC3F273FFF8878BFCF9FF00FCCB" bold="true" box="[151,224,826,852]" italics="true" pageId="7" pageNumber="482">Sueus</emphasis>
<bibRefCitation id="EFAB4B20F273FFF887F4FCF8FE5EFCCB" author="Murayama, J. J." box="[232,446,826,853]" pageId="7" pageNumber="482" pagination="1 - 7" refId="ref9413" refString="Murayama, J. J. (1951) New genus and species of Scolytidae (Coleoptera) from Ohshima and Shionomisaki, Wakayama prefecture. Bulletin of the Faculty of Agriculture, Yamaguti University, 2, 1 - 7." type="journal article" year="1951">Murayama, 1951</bibRefCitation>
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<taxonomicName id="4C3A4D52F273FFF8878BFCBCFE41FC08" ID-CoL="8HRLR" authority="Murayama, 1951: 1" authorityName="Murayama" authorityPageNumber="1" authorityYear="1951" box="[151,417,895,919]" class="Insecta" family="Curculionidae" genus="Sueus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC3F273FFF8878BFCBCFF32FC09" box="[151,210,895,918]" italics="true" pageId="7" pageNumber="482">Sueus</emphasis>
<bibRefCitation id="EFAB4B20F273FFF887C5FCBCFE41FC08" author="Murayama, J. J." box="[217,417,895,919]" pageId="7" pageNumber="482" pagination="1 - 7" refId="ref9413" refString="Murayama, J. J. (1951) New genus and species of Scolytidae (Coleoptera) from Ohshima and Shionomisaki, Wakayama prefecture. Bulletin of the Faculty of Agriculture, Yamaguti University, 2, 1 - 7." type="journal article" year="1951">Murayama, 1951: 1</bibRefCitation>
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.
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<typeStatus id="54818873F273FFF886B0FCBCFE3EFC08" box="[428,478,895,919]" pageId="7" pageNumber="482">Type</typeStatus>
species
<taxonomicName id="4C3A4D52F273FFF88528FCBCFC38FC08" ID-CoL="7B587" authority="Murayama, 1951" authorityName="Murayama" authorityYear="1951" box="[564,984,895,919]" class="Insecta" family="Curculionidae" genus="Sueus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="sphaerotrypoides">
<emphasis id="B94EEAC3F273FFF88528FCBCFCC4FC08" box="[564,804,895,919]" italics="true" pageId="7" pageNumber="482">Sueus sphaerotrypoides</emphasis>
<bibRefCitation id="EFAB4B20F273FFF88437FCBCFC38FC08" author="Murayama, J. J." box="[811,984,895,919]" pageId="7" pageNumber="482" pagination="1 - 7" refId="ref9413" refString="Murayama, J. J. (1951) New genus and species of Scolytidae (Coleoptera) from Ohshima and Shionomisaki, Wakayama prefecture. Bulletin of the Faculty of Agriculture, Yamaguti University, 2, 1 - 7." type="journal article" year="1951">Murayama, 1951</bibRefCitation>
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=
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<taxonomicName id="4C3A4D52F273FFF8878BFC5DFDEEFC29" ID-CoL="3NM25" authority="Eggers, 1926" authorityName="Eggers" authorityYear="1926" box="[151,526,926,950]" class="Insecta" family="Curculionidae" genus="Hyorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="species" species="niisimai">
<emphasis id="B94EEAC3F273FFF8878BFC5DFE60FC2A" box="[151,384,926,950]" italics="true" pageId="7" pageNumber="482">Hyorrhynchus niisimai</emphasis>
<bibRefCitation id="EFAB4B20F273FFF8869AFC5DFDEEFC29" author="Eggers, H." box="[390,526,926,950]" pageId="7" pageNumber="482" pagination="133 - 138" refId="ref9139" refString="Eggers, H. (1926) Japanische Borkenkafer, I. Entomologische Blatter, 22, 133 - 138 + 145 - 148." type="journal article" year="1926">Eggers, 1926</bibRefCitation>
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, original designation.
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<emphasis id="B94EEAC3F273FFF8878BFC27FEF2FC61" bold="true" box="[151,274,996,1022]" pageId="7" pageNumber="482">Diagnosis.</emphasis>
Length 1.63.0 mm, 1.822.5
<emphasis id="B94EEAC3F273FFF8856CFC26FD63FC61" box="[624,643,997,1022]" italics="true" pageId="7" pageNumber="482">×</emphasis>
longer than wide, body black.
<taxonomicName id="4C3A4D52F273FFF884FDFC27FBC2FC61" authorityName="Murayama" authorityYear="1951" box="[993,1058,996,1022]" class="Insecta" family="Curculionidae" genus="Sueus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC3F273FFF884FDFC27FBC2FC61" box="[993,1058,996,1022]" italics="true" pageId="7" pageNumber="482">Sueus</emphasis>
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is distinguished by the following combination of characters: eyes completely divided without a separating carina in either sex; antennal club less than 1.5
<emphasis id="B94EEAC3F273FFF887A6FBEEFF2DFBD9" box="[186,205,1069,1094]" italics="true" pageId="7" pageNumber="482">×</emphasis>
longer than wide, forming a slightly flattened cone; female antennal funicle five or six segmented; protibia outer apical angle produced into one conspicuous spine; procoxae widely separated; and dwarf males.
<taxonomicName id="4C3A4D52F273FFF88237FB93FA8CFBF5" authorityName="Murayama" authorityYear="1951" box="[1323,1388,1104,1130]" class="Insecta" family="Curculionidae" genus="Sueus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC3F273FFF88237FB93FA8CFBF5" box="[1323,1388,1104,1130]" italics="true" pageId="7" pageNumber="482">Sueus</emphasis>
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can be further distinguished from the other
<taxonomicName id="4C3A4D52F273FFF8854EFBB7FD17FB11" authorityName="Hopkins" authorityYear="1915" box="[594,759,1140,1166]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="492" phylum="Arthropoda" rank="tribe" tribe="Hyorrhynchini">Hyorrhynchini</taxonomicName>
genera
<taxonomicName id="4C3A4D52F273FFF8844DFBB7FC11FB11" box="[849,1009,1140,1166]" class="Insecta" family="Curculionidae" genus="Hyorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC3F273FFF8844DFBB7FC11FB11" box="[849,1009,1140,1166]" italics="true" pageId="7" pageNumber="482">Hyorrhynchus</emphasis>
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and
<taxonomicName id="4C3A4D52F273FFF88334FBB7FAF4FB11" box="[1064,1300,1140,1166]" class="Insecta" family="Curculionidae" genus="Pseudohyorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC3F273FFF88334FBB7FAF4FB11" box="[1064,1300,1140,1166]" italics="true" pageId="7" pageNumber="482">Pseudohyorrhynchus</emphasis>
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which have eyes completely divided and separated, in part, by a carina (males); antennal club that is 1.52.0
<emphasis id="B94EEAC3F273FFF883DBFB5AFB3AFB2D" box="[1223,1242,1177,1202]" italics="true" pageId="7" pageNumber="482">×</emphasis>
longer then wide and flattened; antennal funicle six segmented; and males that are comparable to females in size.
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