377 lines
41 KiB
XML
377 lines
41 KiB
XML
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<mods:title id="EDABFA18E7A962C5EA177741375D7FE1">To the knowledge of Oriental species of subgenus Pseudochlaeniellus Jeannel, 1949 (Coleoptera, Carabidae, Chlaenius)</mods:title>
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<treatment id="038487BBFFE4DA2356CD333E822E6BEB" ID-DOI="http://doi.org/10.5281/zenodo.6040738" ID-GBIF-Taxon="127694620" ID-Zenodo-Dep="6040738" LSID="urn:lsid:plazi:treatment:038487BBFFE4DA2356CD333E822E6BEB" httpUri="http://treatment.plazi.org/id/038487BBFFE4DA2356CD333E822E6BEB" lastPageId="3" lastPageNumber="190" pageId="1" pageNumber="188">
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<subSubSection id="C3376526FFE4DA2156CD333E802569BC" pageId="1" pageNumber="188" type="nomenclature">
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<paragraph id="8B9236ADFFE4DA2156CD333E821D699E" blockId="1.[151,862,870,930]" box="[151,862,870,896]" pageId="1" pageNumber="188">
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<heading id="D0DA81C1FFE4DA2156CD333E821D699E" bold="true" box="[151,862,870,896]" fontSize="11" level="1" pageId="1" pageNumber="188" reason="1">
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<taxonomicName id="4C2D4D2EFFE4DA2156CD333E821D699E" ID-CoL="5XVN6" authority="Chaudoir, 1876" authorityName="Chaudoir" authorityYear="1876" box="[151,862,870,896]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="germanus" subGenus="Pseudochlaeniellus">
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<emphasis id="B959EABFFFE4DA2156CD333E821D699E" bold="true" box="[151,862,870,896]" pageId="1" pageNumber="188">
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<emphasis id="B959EABFFFE4DA2156CD333E8050699E" bold="true" box="[151,275,870,896]" italics="true" pageId="1" pageNumber="188">Chlaenius</emphasis>
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(
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<emphasis id="B959EABFFFE4DA21577E333E834D699E" bold="true" box="[292,526,870,896]" italics="true" pageId="1" pageNumber="188">Pseudochlaeniellus</emphasis>
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)
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<emphasis id="B959EABFFFE4DA215445333E83D4699E" bold="true" box="[543,663,870,896]" italics="true" pageId="1" pageNumber="188">germanus</emphasis>
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<bibRefCitation id="EFBC4B5CFFE4DA2154C5333E821D699E" author="Chaudoir" box="[671,862,870,896]" pageId="1" pageNumber="188" refString="Chaudoir, M. de (1876) Monographie des Chlaeniens. Annali del Museo Civico di Storia Naturale di Genova, 8, 5 - 315." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9236ADFFE4DA2156CD33D1802569BC" blockId="1.[151,862,870,930]" box="[151,358,905,930]" pageId="1" pageNumber="188">
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(
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<figureCitation id="13162A28FFE4DA2156C533D1804869BC" box="[159,267,905,930]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1930,1953]" captionTargetBox="[223,1318,205,1871]" captionTargetId="figure@2.[212,1335,193,1883]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Habitus of C. (Pseudochlaeniellus) germanus: 1 — Type of C. germanus. Female (MNHN); 2 — C. (P.) buriensis syn. n., Paratype, female (Thailand); 3 — Phitsanulok, Thailand. Female (NMW); 4 — Sylhet, Bangladesh. Male (BMNH); 5 – 6 — Variability of apical pattern. Karnataka, India. Males (cAA)." httpUri="https://zenodo.org/record/291263/files/figure.png" pageId="1" pageNumber="188">Figs. 1–6</figureCitation>
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,
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<figureCitation id="13162A28FFE4DA21574C33D2801D69BC" box="[278,350,906,930]" captionStart="FIGURES 28 – 36" captionStartId="13.[151,264,1805,1828]" captionTargetBox="[160,1422,730,1777]" captionTargetId="figure@13.[151,1436,724,1785]" captionTargetPageId="13" captionText="FIGURES 28 – 36. Aedeagus: 28 — Holotype of C. chloodes = C. neochloodes (BMNH); 29 — C. caesitius, Loc. Typ. (NMW); 30 — C. puncticollis. India (cAA); 31 — Paratype of C. mesopotamicus = lucasi (NMW); 32 — C. germanus, Karnataka, India (cAA); 33 — C. germanus, Sylhet, Bangladesh. (BMNH); 34 - 35 variability of aedeagus of C. sobrinus, Rajaji, India (cAA); 36 — C. sobrinus, Tonkin, Vietnam (NMW)." httpUri="https://zenodo.org/record/291268/files/figure.png" pageId="1" pageNumber="188">32–33</figureCitation>
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)
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</paragraph>
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<subSubSection id="C3376526FFE4DA2156CD3389827F6E3A" pageId="1" pageNumber="188" type="reference_group">
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<paragraph id="8B9236ADFFE4DA2156CD3389837E69F9" blockId="1.[151,828,977,1060]" box="[151,573,977,999]" pageId="1" pageNumber="188">
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<treatmentCitationGroup id="AB3D1183FFE4DA2156CD3389837E69F9" box="[151,573,977,999]" pageId="1" pageNumber="188">
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<taxonomicName id="4C2D4D2EFFE4DA2156CD3389837E69F9" ID-CoL="5XVN6" authority="Chaudoir, 1876: 199" authorityName="Chaudoir" authorityPageNumber="199" authorityYear="1876" box="[151,573,977,999]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="germanus">
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<emphasis id="B959EABFFFE4DA2156CD3389802B69F9" box="[151,360,977,999]" italics="true" pageId="1" pageNumber="188">Chlaenius germanus</emphasis>
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<treatmentCitation id="0A8C10BCFFE4DA2157343389837E69F9" author="Chaudoir" box="[366,573,977,999]" page="199" pageId="1" pageNumber="188" year="1876">
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<bibRefCitation id="EFBC4B5CFFE4DA2157343389834E69F9" author="Chaudoir" box="[366,525,977,999]" pageId="1" pageNumber="188" refString="Chaudoir, M. de (1876) Monographie des Chlaeniens. Annali del Museo Civico di Storia Naturale di Genova, 8, 5 - 315." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
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: 199
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9236ADFFE4DA2156CD33A883DA6E1B" blockId="1.[151,828,977,1060]" box="[151,665,1008,1030]" pageId="1" pageNumber="188">
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<treatmentCitationGroup id="AB3D1183FFE4DA2156CD33A883DA6E1B" box="[151,665,1008,1030]" pageId="1" pageNumber="188">
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<taxonomicName id="4C2D4D2EFFE4DA2156CD33A8830E6E18" ID-CoL="8MKM2" authority="Chaudoir, 1876: 196" authorityName="Chaudoir" authorityPageNumber="196" authorityYear="1876" box="[151,589,1008,1030]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="laevipennis">
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<emphasis id="B959EABFFFE4DA2156CD33A880346E18" box="[151,375,1008,1030]" italics="true" pageId="1" pageNumber="188">Chlaenius laevipennis</emphasis>
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<treatmentCitation id="0A8C10BCFFE4DA21572733A8830E6E18" author="Chaudoir" box="[381,589,1008,1030]" page="196" pageId="1" pageNumber="188" year="1876">
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<bibRefCitation id="EFBC4B5CFFE4DA21572733A8835E6E18" author="Chaudoir" box="[381,541,1008,1030]" pageId="1" pageNumber="188" refString="Chaudoir, M. de (1876) Monographie des Chlaeniens. Annali del Museo Civico di Storia Naturale di Genova, 8, 5 - 315." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
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: 196
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</treatmentCitation>
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</taxonomicName>
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<emphasis id="B959EABFFFE4DA21540E33A883DA6E1B" bold="true" box="[596,665,1008,1029]" pageId="1" pageNumber="188">
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<taxonomicNameLabel id="A26A57C4FFE4DA21540E33A883DA6E1B" box="[596,665,1008,1029]" pageId="1" pageNumber="188">syn. n.</taxonomicNameLabel>
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</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9236ADFFE4DA2156CD3457827F6E3A" blockId="1.[151,828,977,1060]" box="[151,828,1038,1060]" pageId="1" pageNumber="188">
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<treatmentCitationGroup id="AB3D1183FFE4DA2156CD3457827F6E3A" box="[151,828,1038,1060]" pageId="1" pageNumber="188">
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<taxonomicName id="4C2D4D2EFFE4DA2156CD345783B26E3A" ID-CoL="8MDCF" authority="Kirschenhofer 1998: 321" authorityName="Kirschenhofer" authorityPageNumber="321" authorityYear="1998" box="[151,753,1038,1060]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="buriensis" subGenus="Pseudochlaeniellus">
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||
<emphasis id="B959EABFFFE4DA2156CD345780AE6E3A" box="[151,493,1038,1060]" italics="true" pageId="1" pageNumber="188">C. (Pseudochlaeniellus) buriensis</emphasis>
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<treatmentCitation id="0A8C10BCFFE4DA2157AE345683B26E3A" author="Kirschenhofer" box="[500,753,1038,1060]" page="321" pageId="1" pageNumber="188" year="1998">
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<bibRefCitation id="EFBC4B5CFFE4DA2157AE345683826E3A" author="Kirschenhofer" box="[500,705,1038,1060]" pageId="1" pageNumber="188" refString="Kirschenhofer, E. (1998) Neue Chlaeniinae der palaearktischen und orientalischen Region (Coleoptera, Carabidae). Entomofauna, 19 (20), 317 - 332." type="journal article" year="1998">Kirschenhofer 1998</bibRefCitation>
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: 321
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</treatmentCitation>
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</taxonomicName>
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<emphasis id="B959EABFFFE4DA2154AD3457827F6E3A" bold="true" box="[759,828,1039,1060]" pageId="1" pageNumber="188">
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<taxonomicNameLabel id="A26A57C4FFE4DA2154AD3457827F6E3A" box="[759,828,1039,1060]" pageId="1" pageNumber="188">syn. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C3376526FFE4DA2156CD340980466F5C" pageId="1" pageNumber="188" type="materials_examined">
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<paragraph id="8B9236ADFFE4DA2156CD340980466F5C" blockId="1.[151,1437,1105,2030]" pageId="1" pageNumber="188">
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<materialsCitation id="3B453CF0FFE4DA2156CD340980FA6E77" ID-GBIF-Occurrence="1427603904" box="[151,441,1105,1130]" collectorName="Material. & Type" country="Laos" location="LAOS" pageId="1" pageNumber="188" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<emphasis id="B959EABFFFE4DA2156CD340980456E74" bold="true" box="[151,262,1105,1130]" pageId="1" pageNumber="188">
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<collectorName id="26D8537BFFE4DA2156CD340980456E74" box="[151,262,1105,1130]" pageId="1" pageNumber="188">Material.</collectorName>
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<collectingCountry id="F33A763DFFE4DA215757340A80156E74" box="[269,342,1106,1130]" name="Laos" pageId="1" pageNumber="188">LAOS</collectingCountry>
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:
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<collectorName id="26D8537BFFE4DA21573F340A80E36E74" box="[357,416,1106,1130]" pageId="1" pageNumber="188">Type</collectorName>
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,
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</materialsCitation>
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—
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<materialsCitation id="3B453CF0FFE4DA215782340A82C66E90" ID-GBIF-Occurrence="1427603910" collectingDate="2010-11-06" collectionCode="MNHN" collectorName="Ex Musaeo Chaudoir" country="Laos" county="Luangphabang prov." location="Namtap" municipality="Mt. Phou Phakhao" pageId="1" pageNumber="188" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<collectorName id="26D8537BFFE4DA215782340A838A6E74" box="[472,713,1105,1130]" pageId="1" pageNumber="188">Ex Musaeo Chaudoir</collectorName>
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(
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<collectionCode id="ED3CAE68FFE4DA215483340A82766E74" box="[729,821,1106,1130]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="188">MNHN</collectionCode>
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).
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—
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<collectingCountry id="F33A763DFFE4DA2155D0340A82856E77" box="[906,966,1106,1129]" name="Laos" pageId="1" pageNumber="188">Laos</collectingCountry>
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,
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<collectingCounty id="62F34E21FFE4DA21558B340985F96E77" box="[977,1210,1105,1130]" pageId="1" pageNumber="188">Luangphabang prov.</collectingCounty>
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,
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<collectingMunicipality id="6BF6ACD7FFE4DA21529F340A84DB6E74" box="[1221,1432,1105,1130]" pageId="1" pageNumber="188">Mt. Phou Phakhao</collectingMunicipality>
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,
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<location id="8EF26076FFE4DA2156CD342E81B36E93" LSID="urn:lsid:plazi:treatment:038487BBFFE4DA2356CD333E822E6BEB:8EF26076FFE4DA2156CD342E81B36E93" box="[151,240,1142,1165]" country="Laos" county="Luangphabang prov." municipality="Mt. Phou Phakhao" name="Namtap" pageId="1" pageNumber="188">Namtap</location>
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vill. 7–
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<date id="FF93106DFFE4DA21573A342D80A36E90" box="[352,480,1141,1166]" pageId="1" pageNumber="188" value="2010-11-06">
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<collectingDate id="EFD7E985FFE4DA21573A342D80A36E90" box="[352,480,1141,1166]" pageId="1" pageNumber="188" value="2010-11-06">11.06.2010</collectingDate>
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</date>
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, lg. Sasa Khamphilavong (cAD)
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</materialsCitation>
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.
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<materialsCitation id="3B453CF0FFE4DA2155C7342E853A6E90" ID-GBIF-Occurrence="1427603919" box="[925,1145,1142,1166]" country="Thailand" location="THAILAND" pageId="1" pageNumber="188" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<collectingCountry id="F33A763DFFE4DA2155C7342E85716E90" box="[925,1074,1142,1166]" name="Thailand" pageId="1" pageNumber="188">THAILAND</collectingCountry>
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:
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<specimenCount id="9D2BFD24FFE4DA215210342E853A6E90" box="[1098,1145,1142,1166]" pageId="1" pageNumber="188" type="female">1 ♀</specimenCount>
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—
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<materialsCitation id="3B453CF0FFE4DA2152C2342E83566EC8" ID-GBIF-Occurrence="1427603914" collectionCode="NMW" collectorName="Kirschenhofer & Han-Kha & Suphon-Buri & Thielen" country="Thailand" location="Thailand" pageId="1" pageNumber="188" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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“
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<typeStatus id="5496880FFFE4DA2152F8342E845B6E90" box="[1186,1304,1142,1166]" pageId="1" pageNumber="188" type="paratype">Paratypus</typeStatus>
|
||
<taxonomicName id="4C2D4D2EFFE4DA215376342D80B06EAC" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="buriensis" subGenus="Pseudochlaeniellus">Chlaenius (Pseudochlaeniellus) buriensis</taxonomicName>
|
||
mihi des.
|
||
<collectorName id="26D8537BFFE4DA21542434C1825D6EAC" box="[638,798,1177,1202]" pageId="1" pageNumber="188">Kirschenhofer</collectorName>
|
||
1996”, "
|
||
<collectingCountry id="F33A763DFFE4DA2155D434C182B46EAC" box="[910,1015,1177,1202]" name="Thailand" pageId="1" pageNumber="188">Thailand</collectingCountry>
|
||
,
|
||
<collectorName id="26D8537BFFE4DA21525334C185306EAC" box="[1033,1139,1177,1202]" pageId="1" pageNumber="188">Han-Kha</collectorName>
|
||
,
|
||
<quantity id="4CD59B48FFE4DA2152DF34C185906EAC" box="[1157,1235,1177,1202]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="188" unit="km" value="60.0">60 km</quantity>
|
||
n.
|
||
<collectorName id="26D8537BFFE4DA21535E34C184DB6EAC" box="[1284,1432,1177,1202]" pageId="1" pageNumber="188">Suphon-Buri</collectorName>
|
||
, 1.1990, leg.
|
||
<collectorName id="26D8537BFFE4DA21577834E580386EC8" box="[290,379,1213,1238]" pageId="1" pageNumber="188">Thielen</collectorName>
|
||
" (
|
||
<collectionCode id="ED3CAE68FFE4DA2157C834E680A36EC8" box="[402,480,1214,1238]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="188">NMW</collectionCode>
|
||
);
|
||
<specimenCount id="9D2BFD24FFE4DA2157A934E683566EC8" box="[499,533,1213,1238]" pageId="1" pageNumber="188" type="female">1 ♀</specimenCount>
|
||
</materialsCitation>
|
||
—
|
||
<materialsCitation id="3B453CF0FFE4DA21546E34E585AD6EC8" ID-GBIF-Occurrence="1427603913" box="[564,1262,1213,1238]" collectionCode="NMW" collectorName="H. Schmid" country="Thailand" location="Phitsanulok" pageId="1" pageNumber="188" specimenCount="1" stateProvince="Phitsanulok" typeStatus="paratype">
|
||
“
|
||
<collectingCountry id="F33A763DFFE4DA21546534E583E86EC8" box="[575,683,1213,1238]" name="Thailand" pageId="1" pageNumber="188">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E9F84FFFE4DA2154ED34E5827D6EC8" box="[695,830,1213,1238]" country="Thailand" name="Phitsanulok" pageId="1" pageNumber="188">Phitsanulok</collectingRegion>
|
||
, 1986.12.28, leg.
|
||
<collectorName id="26D8537BFFE4DA21525F34E6853C6EC8" box="[1029,1151,1213,1238]" pageId="1" pageNumber="188">H. Schmid</collectorName>
|
||
” (
|
||
<collectionCode id="ED3CAE68FFE4DA2152C234E685A56EC8" box="[1176,1254,1214,1238]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="188">NMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B453CF0FFE4DA2152A234E6843D6ECB" ID-GBIF-Occurrence="1427603917" box="[1272,1406,1214,1238]" country="India" location="INDIA" pageId="1" pageNumber="188" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
|
||
<collectingCountry id="F33A763DFFE4DA2152A234E684046EC8" box="[1272,1351,1214,1238]" name="India" pageId="1" pageNumber="188">INDIA</collectingCountry>
|
||
:
|
||
<specimenCount id="9D2BFD24FFE4DA21530C34E6843D6ECB" box="[1366,1406,1214,1238]" pageId="1" pageNumber="188" type="male">3 ♂</specimenCount>
|
||
</materialsCitation>
|
||
—
|
||
<materialsCitation id="3B453CF0FFE4DA2156CD34B984DA6EE4" ID-GBIF-Occurrence="1427603941" box="[151,1433,1249,1274]" collectingDate="2013-04-18" collectingDateMax="2013-04-19" collectingDateMin="2013-04-18" collectorName="Anichtchenko A." country="India" latitude="14.880417" location="Karnataka" longLatPrecision="1" longitude="74.26464" pageId="1" pageNumber="188" specimenCount="1" stateProvince="Karnataka" typeStatus="paratype">
|
||
“
|
||
<collectingCountry id="F33A763DFFE4DA2156FB34B9819C6EE4" box="[161,223,1249,1274]" name="India" pageId="1" pageNumber="188">India</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E9F84FFFE4DA2156B434B980276EE4" box="[238,356,1249,1274]" country="India" name="Karnataka" pageId="1" pageNumber="188">Karnataka</collectingRegion>
|
||
, KerwaDi
|
||
<geoCoordinate id="EE19506AFFE4DA2157B834BA83CD6EE4" box="[482,654,1249,1274]" direction="north" orientation="latitude" pageId="1" pageNumber="188" precision="1" value="14.880417">N 14°52’49.5‘‘</geoCoordinate>
|
||
<geoCoordinate id="EE19506AFFE4DA2154D434B9826E6EE4" box="[654,813,1249,1274]" direction="east" orientation="longitude" pageId="1" pageNumber="188" precision="1" value="74.26464">E74°15’52.7‘‘</geoCoordinate>
|
||
lake,
|
||
<date id="FF93106DFFE4DA21552D34BA857B6EE4" box="[887,1080,1250,1274]" pageId="1" pageNumber="188" value="2013-04-18" valueMax="2013-04-19" valueMin="2013-04-18">
|
||
<collectingDate id="EFD7E985FFE4DA21552D34BA857B6EE4" box="[887,1080,1250,1274]" pageId="1" pageNumber="188" value="2013-04-18" valueMax="2013-04-19" valueMin="2013-04-18">18.–19. IV. 2013</collectingDate>
|
||
</date>
|
||
<collectorName id="26D8537BFFE4DA21521834B984476EE4" box="[1090,1284,1249,1274]" pageId="1" pageNumber="188">Anichtchenko A.</collectorName>
|
||
leg.“ (cAA)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B453CF0FFE4DA2156CD355E80466F5C" ID-GBIF-Occurrence="1427603944" collectionCode="BMNH" collectorName="Bowring" country="Bangladesh" location="Sylhet" pageId="1" pageNumber="188" specimenCount="1" stateProvince="Sylhet" typeStatus="paratype">
|
||
<collectingCountry id="F33A763DFFE4DA2156CD355E80166F00" box="[151,341,1286,1310]" name="Bangladesh" pageId="1" pageNumber="188">BANGLADESH</collectingCountry>
|
||
: 1m—“
|
||
<collectingRegion id="49E9F84FFFE4DA2157ED355D83476F00" box="[439,516,1285,1310]" country="Bangladesh" name="Sylhet" pageId="1" pageNumber="188">Sylhet</collectingRegion>
|
||
”, “
|
||
<collectorName id="26D8537BFFE4DA215477355D83D26F00" box="[557,657,1285,1310]" pageId="1" pageNumber="188">Bowring</collectorName>
|
||
63–47”, “
|
||
<taxonomicName id="4C2D4D2EFFE4DA215549355D840B6F00" authority="Chaud. H. E. Andrewes" authorityName="Chaud. H. E. Andrewes" box="[787,1352,1285,1310]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="laevipennis">Chlaenius laevipennis Chaud. H. E. Andrewes</taxonomicName>
|
||
det. “ (
|
||
<collectionCode id="ED3CAE68FFE4DA2156C5357281BA6F5C" box="[159,249,1322,1346]" pageId="1" pageNumber="188">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3376526FFE4DA21569D351582476FCC" pageId="1" pageNumber="188" type="diagnosis">
|
||
<paragraph id="8B9236ADFFE4DA21569D351582476FCC" blockId="1.[151,1437,1105,2030]" pageId="1" pageNumber="188">
|
||
<emphasis id="B959EABFFFE4DA21569D351580026F78" bold="true" box="[199,321,1357,1382]" pageId="1" pageNumber="188">Diagnosis.</emphasis>
|
||
Smallest species of the subgenus. By its size can be confused only with small males of sympatric
|
||
<taxonomicName id="4C2D4D2EFFE4DA2153D8351681B46F97" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="sobrinus">
|
||
<emphasis id="B959EABFFFE4DA2153D8351681B46F97" italics="true" pageId="1" pageNumber="188">C. sobrinus</emphasis>
|
||
</taxonomicName>
|
||
, and by elytral pattern with
|
||
<taxonomicName id="4C2D4D2EFFE4DA215416352A83916F97" box="[588,722,1394,1417]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="caesitius">
|
||
<emphasis id="B959EABFFFE4DA215416352A83916F97" box="[588,722,1394,1417]" italics="true" pageId="1" pageNumber="188">C. caesitius</emphasis>
|
||
</taxonomicName>
|
||
. From
|
||
<taxonomicName id="4C2D4D2EFFE4DA215576352A82F16F97" box="[812,946,1393,1417]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="sobrinus">
|
||
<emphasis id="B959EABFFFE4DA215576352A82F16F97" box="[812,946,1393,1417]" italics="true" pageId="1" pageNumber="188">C. sobrinus</emphasis>
|
||
</taxonomicName>
|
||
it is differentiated, by the metallic lateral margin of pronotum, i.e. in
|
||
<taxonomicName id="4C2D4D2EFFE4DA21578F35CE831B6FB3" box="[469,600,1429,1453]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="sobrinus">
|
||
<emphasis id="B959EABFFFE4DA21578F35CE831B6FB3" box="[469,600,1429,1453]" italics="true" pageId="1" pageNumber="188">C. sobrinus</emphasis>
|
||
</taxonomicName>
|
||
is yellowish translucent. From
|
||
<taxonomicName id="4C2D4D2EFFE4DA21559835CE85066FB0" box="[962,1093,1429,1454]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="caesitius">C. caesitius</taxonomicName>
|
||
it is differentiated by its size. The aedeagus also differs absolutely from both species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3376526FFE4DA21569D358583A86D24" pageId="1" pageNumber="188" type="description">
|
||
<paragraph id="8B9236ADFFE4DA21569D3585855D6C98" blockId="1.[151,1437,1105,2030]" pageId="1" pageNumber="188">
|
||
<emphasis id="B959EABFFFE4DA21569D3585801A6FE8" bold="true" box="[199,345,1501,1526]" pageId="1" pageNumber="188">Description.</emphasis>
|
||
Body length
|
||
<quantity id="4CD59B48FFE4DA2157A83586833E6FEB" box="[498,637,1502,1526]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="7.8" metricValueMin="7.4" pageId="1" pageNumber="188" unit="mm" value="7.6" valueMax="7.8" valueMin="7.4">7.4–7.8 mm</quantity>
|
||
. Dorsal surface with dense yellow pubescence. Coloration of head and pronotum metallic green with bluish overtones on the disc. Mouthparts, antennomeres and legs yellow. Antennomeres IV–XI slightly darker. Elytra metallic green or blue, with narrow yellow lateral band on 2–2.5 external intervals. Apical part of pattern vary in width. Sometimes elytra with large reddish spot on the disc (
|
||
<figureCitation id="13162A28FFE4DA21533E361181806C98" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1930,1953]" captionTargetBox="[223,1318,205,1871]" captionTargetId="figure@2.[212,1335,193,1883]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Habitus of C. (Pseudochlaeniellus) germanus: 1 — Type of C. germanus. Female (MNHN); 2 — C. (P.) buriensis syn. n., Paratype, female (Thailand); 3 — Phitsanulok, Thailand. Female (NMW); 4 — Sylhet, Bangladesh. Male (BMNH); 5 – 6 — Variability of apical pattern. Karnataka, India. Males (cAA)." httpUri="https://zenodo.org/record/291263/files/figure.png" pageId="1" pageNumber="188">Figs. 1–6</figureCitation>
|
||
). Ventral surface black, except wide lateral band on all abdominal segments.
|
||
</paragraph>
|
||
<paragraph id="8B9236ADFFE4DA21569D36C985D66CEC" blockId="1.[151,1437,1105,2030]" pageId="1" pageNumber="188">Head dull, microreticulated throughout, punctate behind eyes and on vertex. Pronotum microreticulate, punctation on disc dense and regular, less or no more than one diameter distance from each other. Lateral margins narrow. Elytra with striae deep and narrow, finely uniformly punctate. Intervals almost flat.</paragraph>
|
||
<paragraph id="8B9236ADFFE4DA21569D36A583A86D24" blockId="1.[151,1437,1105,2030]" pageId="1" pageNumber="188">
|
||
Aedeagus (
|
||
<figureCitation id="13162A28FFE4DA21571F36A580936D08" box="[325,464,1789,1814]" captionStart="FIGURES 28 – 36" captionStartId="13.[151,264,1805,1828]" captionTargetBox="[160,1422,730,1777]" captionTargetId="figure@13.[151,1436,724,1785]" captionTargetPageId="13" captionText="FIGURES 28 – 36. Aedeagus: 28 — Holotype of C. chloodes = C. neochloodes (BMNH); 29 — C. caesitius, Loc. Typ. (NMW); 30 — C. puncticollis. India (cAA); 31 — Paratype of C. mesopotamicus = lucasi (NMW); 32 — C. germanus, Karnataka, India (cAA); 33 — C. germanus, Sylhet, Bangladesh. (BMNH); 34 - 35 variability of aedeagus of C. sobrinus, Rajaji, India (cAA); 36 — C. sobrinus, Tonkin, Vietnam (NMW)." httpUri="https://zenodo.org/record/291268/files/figure.png" pageId="1" pageNumber="188">Figs. 32–33</figureCitation>
|
||
). Specimens from
|
||
<collectingCountry id="F33A763DFFE4DA2154F836A5839F6D08" box="[674,732,1789,1814]" name="India" pageId="1" pageNumber="188">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33A763DFFE4DA21554E36A582D46D08" box="[788,919,1789,1814]" name="Bangladesh" pageId="1" pageNumber="188">Bangladesh</collectingCountry>
|
||
show slight differences in shape of apice, but it should be explained by the intraspecific variability.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3376526FFE4DA23569D371D82A36BCE" lastPageId="3" lastPageNumber="190" pageId="1" pageNumber="188" type="discussion">
|
||
<paragraph id="8B9236ADFFE4DA23569D371D82A36BCE" blockId="1.[151,1437,1105,2030]" lastBlockId="3.[151,1437,151,501]" lastPageId="3" lastPageNumber="190" pageId="1" pageNumber="188">
|
||
<emphasis id="B959EABFFFE4DA21569D371D800C6D40" bold="true" box="[199,335,1861,1886]" pageId="1" pageNumber="188">Comments.</emphasis>
|
||
Species was described in base of single female from Laos. Study of Type revealed is conspecific with species
|
||
<emphasis id="B959EABFFFE4DA215775373280826D9F" box="[303,449,1897,1921]" italics="true" pageId="1" pageNumber="188">
|
||
<taxonomicName id="4C2D4D2EFFE4DA215775373280FE6D9F" box="[303,445,1897,1921]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="buriensis">C. buriensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
also described by females. Last one in differential diagnosis was compared with
|
||
<taxonomicName id="4C2D4D2EFFE4DA2153D8373280596DBB" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="puncticollis">
|
||
<emphasis id="B959EABFFFE4DA2153D8373280596DBB" italics="true" pageId="1" pageNumber="188">C. puncticollis</emphasis>
|
||
</taxonomicName>
|
||
, and as principal differential characters where indicated completely black elytra with slightly wider lateral band. The study of the Type specimen revealed that the elytra looked black because they were greasy (
|
||
<figureCitation id="13162A28FFE4DA21533537E981EB6DF0" captionStart="FIGURES 1 – 6" captionStartId="2.[151,264,1930,1953]" captionTargetBox="[223,1318,205,1871]" captionTargetId="figure@2.[212,1335,193,1883]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Habitus of C. (Pseudochlaeniellus) germanus: 1 — Type of C. germanus. Female (MNHN); 2 — C. (P.) buriensis syn. n., Paratype, female (Thailand); 3 — Phitsanulok, Thailand. Female (NMW); 4 — Sylhet, Bangladesh. Male (BMNH); 5 – 6 — Variability of apical pattern. Karnataka, India. Males (cAA)." httpUri="https://zenodo.org/record/291263/files/figure.png" pageId="1" pageNumber="188">Fig. 2</figureCitation>
|
||
). New synonymy is proposed here. We did not have the opportunity to study males genitalia from Laos and Thailand, but by its external morphology this specimens conspecific with material from Bangladesh and India. Another enigmatic species,
|
||
<taxonomicName id="4C2D4D2EFFE6DA23579730E4832F6ACA" box="[461,620,188,212]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="laevipennis">
|
||
<emphasis id="B959EABFFFE6DA23579730E4832F6ACA" box="[461,620,188,212]" italics="true" pageId="3" pageNumber="190">C. laevipennis</emphasis>
|
||
</taxonomicName>
|
||
was described from Dakkan (India). According to personal comments of Kirschenhofer and Taghavian, the Type of
|
||
<taxonomicName id="4C2D4D2EFFE6DA2354D830B882676AE6" box="[642,804,224,248]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="laevipennis">
|
||
<emphasis id="B959EABFFFE6DA2354D830B882676AE6" box="[642,804,224,248]" italics="true" pageId="3" pageNumber="190">C. laevipennis</emphasis>
|
||
</taxonomicName>
|
||
is unavailable. In original description it was compared with
|
||
<taxonomicName id="4C2D4D2EFFE6DA235695315C80306B02" box="[207,371,260,284]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="puncticollis">
|
||
<emphasis id="B959EABFFFE6DA235695315C80306B02" box="[207,371,260,284]" italics="true" pageId="3" pageNumber="190">C. puncticollis</emphasis>
|
||
</taxonomicName>
|
||
, but differing by densely punctured pronotum, more elongate elytra and bluish head, pronotum and elytra. Under this description fit only two species,
|
||
<taxonomicName id="4C2D4D2EFFE6DA23555D317082C96B21" box="[775,906,296,319]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="caesitius">
|
||
<emphasis id="B959EABFFFE6DA23555D317082C96B21" box="[775,906,296,319]" italics="true" pageId="3" pageNumber="190">C. caesitius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2D4D2EFFE6DA235599317085106B5E" box="[963,1107,296,320]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="germanus">
|
||
<emphasis id="B959EABFFFE6DA235599317085106B5E" box="[963,1107,296,320]" italics="true" pageId="3" pageNumber="190">C. germanus</emphasis>
|
||
</taxonomicName>
|
||
. We saw one specimen from Bangladesh (BMNH), determined by Andrewes as
|
||
<emphasis id="B959EABFFFE6DA235489311482396B7A" box="[723,890,332,356]" italics="true" pageId="3" pageNumber="190">
|
||
<taxonomicName id="4C2D4D2EFFE6DA235489311482356B7A" box="[723,886,332,356]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="laevipennis">C. laevipennis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and is conspecific with
|
||
<taxonomicName id="4C2D4D2EFFE6DA2352D03114845B6B7A" box="[1162,1304,332,356]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="germanus">
|
||
<emphasis id="B959EABFFFE6DA2352D03114845B6B7A" box="[1162,1304,332,356]" italics="true" pageId="3" pageNumber="190">C. germanus</emphasis>
|
||
</taxonomicName>
|
||
. Given that the
|
||
<taxonomicName id="4C2D4D2EFFE6DA23569A312880016B99" box="[192,322,368,391]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="caesitius">
|
||
<emphasis id="B959EABFFFE6DA23569A312880016B99" box="[192,322,368,391]" italics="true" pageId="3" pageNumber="190">C. caesitius</emphasis>
|
||
</taxonomicName>
|
||
is known from Vietnam only, while
|
||
<taxonomicName id="4C2D4D2EFFE6DA235481312882296B96" box="[731,874,368,392]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="germanus">
|
||
<emphasis id="B959EABFFFE6DA235481312882296B96" box="[731,874,368,392]" italics="true" pageId="3" pageNumber="190">C. germanus</emphasis>
|
||
</taxonomicName>
|
||
is widespreaded, we tentatively propose here new synonymy
|
||
<taxonomicName id="4C2D4D2EFFE6DA23574F31CC83336BB3" authority="Chaudoir, 1876" authorityName="Chaudoir" authorityYear="1876" box="[277,624,404,429]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="laevipennis">
|
||
<emphasis id="B959EABFFFE6DA23574F31CC80FB6BB2" box="[277,440,404,428]" italics="true" pageId="3" pageNumber="190">C. laevipennis</emphasis>
|
||
<bibRefCitation id="EFBC4B5CFFE6DA23579B31CC83336BB3" author="Chaudoir" box="[449,624,404,429]" pageId="3" pageNumber="190" refString="Chaudoir, M. de (1876) Monographie des Chlaeniens. Annali del Museo Civico di Storia Naturale di Genova, 8, 5 - 315." type="journal article" year="1876">Chaudoir, 1876</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B959EABFFFE6DA23542331CC83896BB3" bold="true" box="[633,714,404,429]" pageId="3" pageNumber="190">
|
||
<taxonomicNameLabel id="A26A57C4FFE6DA23542331CC83896BB3" box="[633,714,404,429]" pageId="3" pageNumber="190">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
=
|
||
<taxonomicName id="4C2D4D2EFFE6DA2354B131CC823F6BB2" box="[747,892,404,428]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="190" phylum="Arthropoda" rank="species" species="germanus">
|
||
<emphasis id="B959EABFFFE6DA2354B131CC823F6BB2" box="[747,892,404,428]" italics="true" pageId="3" pageNumber="190">C. germanus</emphasis>
|
||
</taxonomicName>
|
||
. To make a final decision about the taxonomic status of this taxon it is necessary to study the type and additional material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF526625FFE7DA2256CD37D282E06DC3" httpUri="https://zenodo.org/record/291263/files/figure.png" pageId="2" pageNumber="189" targetBox="[223,1318,205,1871]" targetPageId="2">
|
||
<paragraph id="8B9236ADFFE7DA2256CD37D282E06DC3" blockId="2.[151,1436,1930,2013]" pageId="2" pageNumber="189">
|
||
<emphasis id="B959EABFFFE7DA2256CD37D280036D81" bold="true" box="[151,320,1930,1953]" pageId="2" pageNumber="189">FIGURES 1–6.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C2D4D2EFFE7DA22579337D382716DBE" box="[457,818,1930,1952]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="germanus" subGenus="Pseudochlaeniellus">
|
||
<emphasis id="B959EABFFFE7DA22579337D380996DBE" box="[457,474,1931,1952]" italics="true" pageId="2" pageNumber="189">C</emphasis>
|
||
. (
|
||
<emphasis id="B959EABFFFE7DA2257AF37D283F96DBE" box="[501,698,1930,1952]" italics="true" pageId="2" pageNumber="189">Pseudochlaeniellus</emphasis>
|
||
)
|
||
<emphasis id="B959EABFFFE7DA22549437D382716DBE" box="[718,818,1931,1952]" italics="true" pageId="2" pageNumber="189">germanus</emphasis>
|
||
</taxonomicName>
|
||
: 1—Type of
|
||
<taxonomicName id="4C2D4D2EFFE7DA22559637D385106DBE" box="[972,1107,1931,1952]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="germanus">
|
||
<emphasis id="B959EABFFFE7DA22559637D3829E6DBE" box="[972,989,1931,1952]" italics="true" pageId="2" pageNumber="189">C</emphasis>
|
||
.
|
||
<emphasis id="B959EABFFFE7DA2255B537D385106DBE" box="[1007,1107,1931,1952]" italics="true" pageId="2" pageNumber="189">germanus</emphasis>
|
||
</taxonomicName>
|
||
. Female (MNHN); 2—
|
||
<taxonomicName id="4C2D4D2EFFE7DA22530E37D381B76DA1" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="189" phylum="Arthropoda" rank="species" species="buriensis" subGenus="Pseudochlaeniellus">
|
||
<emphasis id="B959EABFFFE7DA22530E37D384266DBE" box="[1364,1381,1931,1952]" italics="true" pageId="2" pageNumber="189">C</emphasis>
|
||
. (
|
||
<emphasis id="B959EABFFFE7DA2253DA37D384CE6DBE" box="[1408,1421,1931,1952]" italics="true" pageId="2" pageNumber="189">P</emphasis>
|
||
.)
|
||
<emphasis id="B959EABFFFE7DA2256CD37F181B76DA1" box="[151,244,1961,1983]" italics="true" pageId="2" pageNumber="189">buriensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B959EABFFFE7DA2256A537F180096DA0" bold="true" box="[255,330,1961,1982]" pageId="2" pageNumber="189">
|
||
<taxonomicNameLabel id="A26A57C4FFE7DA2256A537F180096DA0" box="[255,330,1961,1982]" pageId="2" pageNumber="189">syn. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Paratype, female (Thailand); 3—Phitsanulok, Thailand. Female (NMW); 4—Sylhet, Bangladesh. Male (BMNH); 5–6—Variability of apical pattern. Karnataka, India. Males (cAA).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3376526FFE6DA23569D3184822E6BEB" box="[199,877,476,501]" pageId="3" pageNumber="190" type="distribution">
|
||
<paragraph id="8B9236ADFFE6DA23569D3184822E6BEB" blockId="3.[151,1437,151,501]" box="[199,877,476,501]" pageId="3" pageNumber="190">
|
||
<emphasis id="B959EABFFFE6DA23569D318480236BEB" bold="true" box="[199,352,476,501]" pageId="3" pageNumber="190">Distribution.</emphasis>
|
||
<collectingCountry id="F33A763DFFE6DA23573D318480AE6BEB" box="[359,493,476,501]" name="Bangladesh" pageId="3" pageNumber="190">Bangladesh</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33A763DFFE6DA2357A2318483776BEB" box="[504,564,476,501]" name="India" pageId="3" pageNumber="190">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33A763DFFE6DA235465318583366BEA" box="[575,629,477,500]" name="Laos" pageId="3" pageNumber="190">Laos</collectingCountry>
|
||
(Loc. typ.),
|
||
<collectingCountry id="F33A763DFFE6DA2355583184822A6BEB" box="[770,873,476,501]" name="Thailand" pageId="3" pageNumber="190">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |