114 lines
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114 lines
18 KiB
XML
<document id="37C2577C1E0E6BE7080D6CA2075CF232" ID-DOI="10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="10110744" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630265040447" checkinUser="carolina" docAuthor="SMETANA, ALEŠ & DAVIES, ANTHONY" docDate="2000" docId="0384878FD83DFB30FC10691208D9FE01" docLanguage="en" docName="N3287.pdf" docOrigin="American Museum Novitates 3287" docSource="http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2" docStyle="DocumentStyle:059C9B7006157ECE636E30DEF6793B3A.3:AmMusNovit.2000-2010.journal_article.type2" docStyleId="059C9B7006157ECE636E30DEF6793B3A" docStyleName="AmMusNovit.2000-2010.journal_article.type2" docStyleVersion="3" docTitle="Creophilus Leach 1819" docType="treatment" docVersion="3" masterDocId="FFBDFFF7D829FB25FFD86B530951FFD4" masterDocTitle="Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1699682540511" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="45421BFE0B138A005DEDCDA16FC777FC">
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<mods:title id="1225B70299FE4757841127EE260F0C8A">Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="63F0E48523D6B709E6BBF8E27868E209">SMETANA, ALEŠ</mods:namePart>
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<mods:namePart id="9B47031CB20F544E7B4E7731A73D175C">DAVIES, ANTHONY</mods:namePart>
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<mods:title id="F34C1E63C3F96E3D3186D1B07AFD1EC0">American Museum Novitates</mods:title>
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<mods:part id="277D13FD96EC0A0F90CCCE5C5BDEA4FB">
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<mods:date id="C806057268F80BFAF2E58FD3720FD5A4">2000</mods:date>
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<mods:number id="6C04790C06754DF9FD66B0A1BF7DEDF6">2000-02-02</mods:number>
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</mods:detail>
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<mods:detail id="C01BC0EEA859F591CBFB59CEEABFECB7" type="volume">
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<mods:number id="42172EDA55800179F0319E099414BE93">3287</mods:number>
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<mods:url id="6D4C6A99BFE3658A31D3F5A0EC215164">http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2</mods:url>
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<mods:classification id="F52670DA56885F165D4F27D22F94E19D">journal article</mods:classification>
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<mods:identifier id="898E228120C0999B1B147BBCDF5AB190" type="DOI">10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2</mods:identifier>
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<mods:identifier id="823886D955805EC7A32DEB9C2B9F7ECE" type="ISSN">0003-0082</mods:identifier>
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<mods:identifier id="AE327B87D5653F09A38C5DD89F8C6A14" type="Zenodo-Dep">10110744</mods:identifier>
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<treatment id="0384878FD83DFB30FC10691208D9FE01" LSID="urn:lsid:plazi:treatment:0384878FD83DFB30FC10691208D9FE01" httpUri="http://treatment.plazi.org/id/0384878FD83DFB30FC10691208D9FE01" lastPageId="21" pageId="20">
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<subSubSection id="C3376512D83DFB31FC1069120DB9FD8C" box="[968,1256,577,601]" pageId="20" type="nomenclature">
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<paragraph id="8B923699D83DFB31FC1069120DB9FD8C" blockId="20.[968,1256,577,601]" box="[968,1256,577,601]" pageId="20">
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<heading id="D0DA81F5D83DFB31FC1069120DB9FD8C" box="[968,1256,577,601]" centered="true" fontSize="36" level="2" pageId="20" reason="8">
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<taxonomicName id="4C2D4D1AD83DFB31FC1069120DB9FD8C" ID-CoL="8VRLQ" authority="Leach, 1819" authorityName="Leach" authorityYear="1819" box="[968,1256,577,601]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD83DFB31FC1069120D1DFD8D" bold="true" box="[968,1100,577,601]" italics="true" pageId="20">Creophilus</emphasis>
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Leach, 1819
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376512D83DFB31FCB5693D0AEFFD77" pageId="20" type="materials_examined">
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<paragraph id="8B923699D83DFB31FCB5693D0AEFFD77" blockId="20.[848,1377,622,1731]" pageId="20">
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<materialsCitation id="3B453CC4D83DFB31FCB5693D0AEAFD77" collectionCode="SPECIES" pageId="20" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5496883BD83DFB31FCB5693D0AF6FD51" box="[877,935,622,645]" pageId="20">TYPE</typeStatus>
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<collectionCode id="ED3CAE5CD83DFB31FC68693D0D59FD51" box="[944,1032,622,645]" pageId="20">SPECIES</collectionCode>
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:
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<taxonomicName id="4C2D4D1AD83DFB31FBC1693C0C74FD52" authorityName="Linnaeus" authorityYear="1758" box="[1049,1317,623,646]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="species" species="maxillosus">
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<emphasis id="B959EA8BD83DFB31FBC1693C0C74FD52" box="[1049,1317,623,646]" italics="true" pageId="20">Staphylinus maxillosus</emphasis>
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</taxonomicName>
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Linne´, 1758
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3376512D83DFB30FCB569FA08D9FE01" lastPageId="21" pageId="20" type="discussion">
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<paragraph id="8B923699D83DFB31FCB569FA0A93FAFC" blockId="20.[848,1377,622,1731]" pageId="20">
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COMMENTS: Members of this genus may be easily recognised by the following shared character states: (
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<bibRefCitation id="EFBC4B68D83DFB31FBF169B70D64FD2F" author="Natural History Museum, London" box="[1065,1077,740,763]" pageId="20" pagination="204" refId="ref25242" refString="1. Ocypus almorensis (Cameron, 1932). Cameron (1932: 204) described the species (in Staphylinus) from specimens from '' West Almora: Kali Valley, alt. 9000 feet' ' and '' Punjab: Triun, alt. 9300 feet. '' We were able to study one of the male syntypes in the Natural History Museum, London, England, labelled as follows: '' Triun, 9,300 ft Dharm- sala. Punjab. H. G. C. / M. Cameron. Bequest. B. M. 1955 - 147. / Staphylinus (Ocypus) almorensis. '' The specimen is in very good shape. It was dissected, and the aedeagus and the genital segment were glued to the plate with the beetle. It is hereby designated as the lectotype of Staphylinus almorensis; the label '' LECTOTYPE Staphylinus almorensis Cameron A. Smetana des. 1999 '' was attached to it. The specimen cannot be distinguished from those of Ocypus helleni J. Muller that occur in northern Pakistan and in Kashmir. The name O. almorensis becomes a junior synonym of O. helleni (syn. nov.). The corresponding determination label was attached to the specimen." type="journal article" year="1955">1</bibRefCitation>
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) palpifer with one long subapical seta and three to four shorter apical setae; (
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<bibRefCitation id="EFBC4B68D83DFB31FC7D684D0AE4FCE1" author="Ocypus" box="[933,949,798,821]" pageId="20" pagination="170" refId="ref25465" refString="2. Ocypus auricomus (Lindberg, 1953: 4), described in Staphylinus, is preoccupied by Staphylinus auricomus Brulle', 1842: pl. 5, fig. 6, now a synonym of Glenus chrysis (Gravenhorst, 1806), and by Staphylinus auricomus Cameron, 1929: 65. The name Ocypus mateui (Coiffait, 1954: 170), listed at present as a junior synonym of O. auricomus, becomes the valid name of this species (stat. nov.)." type="journal article" year="1842">2</bibRefCitation>
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) disc of pronotum and most of dorsal face of pronotum virtually impunctate and asetose; (
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<bibRefCitation id="EFBC4B68D83DFB31FC20680A0D56FCA4" author="Lordithon" box="[1016,1031,857,880]" pageId="20" refId="ref25558" refString="3. Ocypus bimaculatus (Cameron, 1932: 207), described in Staphylinus, is preoccupied by several names in Staphylinus, the oldest one being Staphylinus bimaculatus Schrank, 1798: 644, now a synonym of Lordithon trinotatus (Erichson, 1839 b: 409). The replacement name Ocypus cameroni, nom. nov., is hereby proposed for Ocypus bimaculatus (Cameron, 1932)." type="journal volume" year="1839">3</bibRefCitation>
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) middle coxae widely separated by broadly rounded apical margin of mesosternum; (
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<bibRefCitation id="EFBC4B68D83DFB31FBD168C70D48FC7F" author="Naturhistorisches Museum in Wien" box="[1033,1049,916,939]" pageId="20" pagination="172" refId="ref25632" refString="4. Ocypus fulvotomentosus Eppelsheim, 1889: 172. The collection at the Naturhistorisches Museum in Wien, Austria, contains one male specimen under this name. It is labelled as follows: '' Kan-ssa 1885 G. Patanin / c. Eppelsh. Steind. d. / fulvotomentosus det. Bernhauer / fulvotomentosus det. J. Muller / TYPUS. '' The specimen is in good condition; however, the entire left middle leg, out- er three segments on left hind tarsus, and the entire right middle tarsus are missing. It was dissected and the aedeagus was glued to a plate attached to the pin, with the beetle. The specimen is hereby designated as the lectotype of O. fulvotomentosus; the label '' LEC- TOTYPE Ocypus fulvotomentosus Eppelsheim A. Smetana des. 1997 '' was attached to it." type="journal article" year="1997">4</bibRefCitation>
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) pronotal hypomeron with superior line becoming obsolete near anterior angles of pronotum and not joining the inferior line (
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<figureCitation id="13162A1CD83DFB31FC3A68BF0D68FBD7" box="[994,1081,1004,1027]" captionStart="Figs" captionStartId="68.[298,346,1509,1530]" captionTargetBox="[309,1338,176,1488]" captionTargetId="figure-35@68.[309,1338,176,1488]" captionTargetPageId="68" captionText="Figs. 45–52. 45. Philonthus caeruleipennis, superior and inferior marginal lines of pronotal hypomeron not joined, ventral view. 46. Agelosus carinatus, superior and inferior marginal lines of pronotal hypomeron distinctly not joined, ventral view. 47. Ocypus weisei, superior and inferior marginal lines of pronotal hypomeron not quite joined, ventral view. 48. Ocypus tenebricosus, superior and inferior marginal lines of pronotal hypomeron not quite joined, ventral view. 49. Creophilus maxillosus, superior marginal line of pronotal hypomeron becoming obsolete near anterior angles, ventral view. 50. Staphylinus dimidiaticornis, joining of superior and inferior marginal lines of pronotal hypomeron, ventral view. 51. Miobdelus sp., joining of superior and inferior marginal lines of pronotal hypomeron, ventral view. 52. Platydracus stercorarius, superior and inferior marginal lines of pronotal hypomeron not quite joined, appearing as a grooved carina anteriorly, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10110766" httpUri="https://zenodo.org/record/10110766/files/figure.png" pageId="20">fig. 49</figureCitation>
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); (
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<bibRefCitation id="EFBC4B68D83DFB31FB8768BF0D3FFBD7" box="[1119,1134,1004,1027]" pageId="20" refId="ref25784" refString="5. Ocypus hauseri (Bernhauer, 1933: 32). The collection at the Field Museum of Natural History, Chicago, Illinois, contains one male specimen (holotype) under this name. It is labelled as follows: '' Kalgan Jnn-shan 980 - 2100 m / 285 / Hauseri Brh. Typus unic. / Hauseri Bernh. Typus unic. / Chicago NHMus M. Bernhauer Collection. '' The specimen is in very good condition. It was dissected, compared to the lectotype of O. fulvotomentosus (see above), and was found to be conspecific with it. The name O. hauseri becomes a junior synonym of O. fulvotomentosus (syn. nov.); the holotype was labelled accordingly." type="book">5</bibRefCitation>
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) presence, on both dorsal and ventral faces of body, of long, moreorless variegate pubescence varying in color from black to grayishsilver, grayishgolden, yellowishred, or brownishred (such pubescence not present in a few species occurring elsewhere); (
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<bibRefCitation id="EFBC4B68D83DFB31FB9D6FCF0D04FB67" author="Natural History Museum, London" box="[1093,1109,1180,1203]" pageId="20" pagination="288" refId="ref25919" refString="6. Ocypus himalayicus (Cameron, 1935: 288) (Staphylinus). Cameron (loc. cit.) described the species from a single female from Karakorum range. The holotype, deposited at the Natural History Museum, London, England, is labelled as follows: '' Type [round disc with red margin] / Sind valley 2000 - 2800 m 2 - 5. V. 1929 / Nederlandsche Karakorum- Expeditie J. A. Sillem leg. / M. Cameron. Bequest. B. M. 1955 - 147. / Staphylinus himalayicus Cam. TYPE. '' The specimen is in fair shape (last segment of left front tarsus and claws of middle right tarsus are missing, and most of the pubescence of the head and pronotum is missing as well). The specimen cannot be distinguished from those of Ocypus (Pseudocypus) helleni J. Muller that occurs in northern Pakistan and in Kashmir. The name O. himalayicus becomes a junior synonym of O. helleni (syn. nov.). The corresponding determination label was attached to the specimen." type="journal article" year="2000">6</bibRefCitation>
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) anterior projection of metasternum very broad, with margins obsolete; and (
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<bibRefCitation id="EFBC4B68D83DFB31FC376F850AAFFB39" author="Erichson" box="[1007,1022,1238,1261]" pageId="20" pagination="406" refId="ref26121" refString="7. Ocypus italicus (Aragona, 1830: 10). By coincidence, Erichson (1840: 406) later redescribed this species as new, using the same name as Aragona. Subsequent authors considered Erichson's description as a subsequent citation of Aragona's species. However, based on Erichson's treatment of his Ocypus italicus, including his comments un- der '' Obs., '' there is little doubt that Erichson considered it a new species. Therefore, the name Ocypus italicus Erichson, 1840 becomes a junior synonym of Ocypus italicus (Aragona, 1830) (syn. nov.)." type="journal article" year="1840">7</bibRefCitation>
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) dorsal apicolateral lobe of hind coxa without distinct spines, or spines very fine.
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</paragraph>
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<paragraph id="8B923699D83DFB31FCA36E7D0D0EF9BF" blockId="20.[848,1377,622,1731]" pageId="20">Further character states include: mandibles each with microsculpture on mediobasal portion; mandibular prostheca copiously ciliate along entire length, with long, dense basal ciliae gradually shortening apicad; postmandibular ridge absent; deflected portion of pronotal disc visible in ventral view from about anterior edge of pronotal epimeron, which is well developed and membranous; metasternum oblique, highly convex between coxae, then horizontal.</paragraph>
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<paragraph id="8B923699D83DFB30FCB56D2208D9FE01" blockId="20.[848,1377,622,1731]" lastBlockId="21.[238,767,182,469]" lastPageId="21" pageId="20">
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This genus is assigned by some authors (e.g., Coiffait, 1974: 557; Moore and Legner, 1979: 32) to the subtribe
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<taxonomicName id="4C2D4D1AD83DFB31FB646DFF0C30F917" baseAuthorityName="Naomi" baseAuthorityYear="1983" box="[1212,1377,1708,1731]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
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based on the configuration of the superior and inferior lines of the pronotal hypomeron (see above). Although
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<taxonomicName id="4C2D4D1AD83CFB30FE2E6BA20B26FEDC" authorityName="Leach" authorityYear="1819" box="[502,631,241,264]" class="Insecta" family="Staphylinidae" genus="Creophilus" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD83CFB30FE2E6BA20B26FEDC" box="[502,631,241,264]" italics="true" pageId="21">Creophilus</emphasis>
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</taxonomicName>
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(with
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<taxonomicName id="4C2D4D1AD83CFB30FD1A6BA20879FEF2" authorityName="Maklin" authorityYear="1852" class="Insecta" family="Staphylinidae" genus="Hadrotes" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD83CFB30FD1A6BA20879FEF2" italics="true" pageId="21">Hadrotes</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2D4D1AD83CFB30FEB16A5C08BDFEF2" authorityName="LeConte" authorityYear="1852" box="[361,492,271,294]" class="Insecta" family="Staphylinidae" genus="Thinopinus" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="genus">
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<emphasis id="B959EA8BD83CFB30FEB16A5C08BDFEF2" box="[361,492,271,294]" italics="true" pageId="21">Thinopinus</emphasis>
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</taxonomicName>
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) seems to be intermediate between the two subtribes, characters 1, 6, and 7 above are shared exclusively with members of the
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<taxonomicName id="4C2D4D1AD83CFB30FE606A350B1EFEA9" authorityName="Latreille" authorityYear="1802" box="[440,591,358,381]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
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; together with the absence of the postmandibular ridge, these confirm their closer affinity with the
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<taxonomicName id="4C2D4D1AD83CFB30FF366AED08D4FE01" authorityName="Latreille" authorityYear="1802" box="[238,389,446,469]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |