Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae) Author SMETANA, ALEŠ Author DAVIES, ANTHONY text American Museum Novitates 2000 2000-02-02 3287 1 1 http://www.bioone.org/doi/abs/10.1206/0003-0082%282000%29287%3C0001%3AROTNTT%3E2.0.CO%3B2 journal article 10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2 0003-0082 10110744 Creophilus Leach, 1819 TYPE SPECIES : Staphylinus maxillosus Linne´, 1758 . COMMENTS: Members of this genus may be easily recognised by the following shared character states: ( 1 ) palpifer with one long subapical seta and three to four shorter apical setae; ( 2 ) disc of pronotum and most of dorsal face of pronotum virtually impunctate and asetose; ( 3 ) middle coxae widely separated by broadly rounded apical margin of mesosternum; ( 4 ) pronotal hypomeron with superior line becoming obsolete near anterior angles of pronotum and not joining the inferior line ( fig. 49 ); ( 5 ) presence, on both dorsal and ventral faces of body, of long, more­or­less variegate pubescence varying in color from black to grayish­silver, grayishgolden, yellowish­red, or brownish­red (such pubescence not present in a few species occurring elsewhere); ( 6 ) anterior projection of metasternum very broad, with margins obsolete; and ( 7 ) dorsal apicolateral lobe of hind coxa without distinct spines, or spines very fine. Further character states include: mandibles each with microsculpture on mediobasal portion; mandibular prostheca copiously ciliate along entire length, with long, dense basal ciliae gradually shortening apicad; postmandibular ridge absent; deflected portion of pronotal disc visible in ventral view from about anterior edge of pronotal epimeron, which is well developed and membranous; metasternum oblique, highly convex between coxae, then horizontal. This genus is assigned by some authors (e.g., Coiffait, 1974: 557; Moore and Legner, 1979: 32) to the subtribe Xanthopygina based on the configuration of the superior and inferior lines of the pronotal hypomeron (see above). Although Creophilus (with Hadrotes and Thinopinus ) seems to be intermediate between the two subtribes, characters 1, 6, and 7 above are shared exclusively with members of the Staphylinina ; together with the absence of the postmandibular ridge, these confirm their closer affinity with the Staphylinina .