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<document id="C8FEDB36CFFFA5D735375D0DF0F1858E" ID-CLB-Dataset="8504" ID-DOI="10.11646/zootaxa.4920.1.3" ID-GBIF-Dataset="8c7a90dc-a88c-4ae7-9981-9ba0e02b9dce" ID-ISSN="1175-5326" ID-Zenodo-Dep="4471639" ID-ZooBank="F26DD750-81BE-4D94-A4ED-62FED16A4581" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611747615504" checkinUser="plazi" docAuthor="Ermilov, Sergey G., Hugo-Coetzee, Elizabeth A. &amp; Khaustov, Alexander A." docDate="2021" docId="033087806439834F20B1FA90FE444F9B" docLanguage="en" docName="zootaxa.4920.1.3.pdf" docOrigin="Zootaxa 4920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stictozetes ihaguensis Ermilov &amp; Hugo-Coetzee &amp; Khaustov 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="99" masterDocId="FF09FFF8643F83472026FFBDFF9E4D55" masterDocTitle="Three new species of oribatid mites of the family Galumnidae (Acari, Oribatida) from South Africa" masterLastPageNumber="100" masterPageNumber="91" pageNumber="97" updateTime="1698925638261" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="32BD6081EB7F57A93FF9E86EA398E57D">Three new species of oribatid mites of the family Galumnidae (Acari, Oribatida) from South Africa</mods:title>
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<mods:namePart id="CAF477A2CEC97D61537192337ADF1FB0">Ermilov, Sergey G.</mods:namePart>
<mods:affiliation id="99B9F35C88FA7C154C41392BEB768EB6">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia</mods:affiliation>
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<mods:namePart id="55F19A4F31F4672C5A22DADF0F94EBEC">Hugo-Coetzee, Elizabeth A.</mods:namePart>
<mods:affiliation id="8E8B623C7E1388F386224A2804C2A4B1">National Museum, Bloemfontein, South Africa &amp; University of the Free State, Bloemfontein, South Africa</mods:affiliation>
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<mods:namePart id="A16665A2347F7841B115FD4461817E18">Khaustov, Alexander A.</mods:namePart>
<mods:nameIdentifier id="11240F1BB6E359DA63A33C0D2D9F74F7" type="ORCID">0000-0002-0306-112X</mods:nameIdentifier>
<mods:affiliation id="64FB62C2F55F9862CF0EE2DCB6A7A8AD">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia &amp; alex 1973 khaustov @ gmail. com; https: // orcid. org / 0000 - 0002 - 0306 - 112 X</mods:affiliation>
<mods:nameIdentifier id="D4F9B35945BC0660B414D50237C0217B" type="email">alex1973khaustov@gmail.com</mods:nameIdentifier>
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<treatment id="033087806439834F20B1FA90FE444F9B" ID-DOI="http://doi.org/10.5281/zenodo.4471635" ID-GBIF-Taxon="177207332" ID-Zenodo-Dep="4471635" LSID="urn:lsid:plazi:treatment:033087806439834F20B1FA90FE444F9B" httpUri="http://treatment.plazi.org/id/033087806439834F20B1FA90FE444F9B" lastPageId="8" lastPageNumber="99" pageId="6" pageNumber="97">
<subSubSection id="C383651D6439834120B1FA90FD9D4812" box="[151,515,1325,1352]" pageId="6" pageNumber="97" type="nomenclature">
<paragraph id="8B2636966439834120B1FA90FD9D4812" blockId="6.[151,515,1325,1387]" box="[151,515,1325,1352]" pageId="6" pageNumber="97">
<heading id="D06E81FA6439834120B1FA90FD9D4812" bold="true" box="[151,515,1325,1352]" fontSize="11" level="1" pageId="6" pageNumber="97" reason="1">
<emphasis id="B9EDEA846439834120B1FA90FD9D4812" bold="true" box="[151,515,1325,1352]" pageId="6" pageNumber="97">
<taxonomicName id="4C994D156439834120B1FA90FE00481D" ID-CoL="8TBJN" authority="Ermilov &amp; Hugo-Coetzee &amp; Khaustov, 2021" authorityName="Ermilov &amp; Hugo-Coetzee &amp; Khaustov" authorityYear="2021" box="[151,414,1325,1352]" class="Arachnida" family="Galumnidae" genus="Stictozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="97" phylum="Arthropoda" rank="species" species="ihaguensis" status="sp. nov.">
<emphasis id="B9EDEA846439834120B1FA90FE00481D" bold="true" box="[151,414,1325,1352]" italics="true" pageId="6" pageNumber="97">Stictozetes ihaguensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2DE57FF643983412180FA90FD9D4812" box="[422,515,1325,1351]" pageId="6" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C383651D6439834120B1FAECFF75483E" box="[151,235,1361,1387]" pageId="6" pageNumber="97" type="description">
<paragraph id="8B2636966439834120B1FAECFF75483E" blockId="6.[151,515,1325,1387]" box="[151,235,1361,1387]" pageId="6" pageNumber="97">
(
<figureCitation id="13A22A13643983412086FAECFF7D483E" box="[160,227,1361,1387]" captionStart="FIGURE 4" captionStartId="7.[151,250,1957,1981]" captionTargetBox="[170,1422,717,1925]" captionTargetId="figure-273@7.[151,1436,705,1934]" captionTargetPageId="7" captionText="FIGURE 4. Stictozetes ihaguensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs omitted); C—lateral view (pteromorph, gnathosoma and legs omitted); D—posterior view (pteromorphs omitted). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.4471648" httpUri="https://zenodo.org/record/4471648/files/figure.png" pageId="6" pageNumber="97">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C383651D6439834120B1FA24FCE54B16" pageId="6" pageNumber="97" type="diagnosis">
<paragraph id="8B2636966439834120B1FA24FCE54B16" blockId="6.[151,1437,1433,2036]" pageId="6" pageNumber="97">
<emphasis id="B9EDEA846439834120B1FA24FE8C48E6" bold="true" box="[151,274,1433,1459]" pageId="6" pageNumber="97">Diagnosis.</emphasis>
Body size: 298332 × 232249. Body surface granulate. Rostral seta long, setiform, slightly barbed. Lamellar seta short, setiform, slightly barbed. Interlamellar seta minute. Bothridial seta long, with elongate, narrowly unilaterally dilated, barbed head, pointed apically. Dorsosejugal suture complete. With three pairs of rounded notogastral porose areas. Median pore represented by 24 foveae. Epimeral and anogenital setae short, setiform, roughened. Circumpedal carina long. Postanal porose area absent.
</paragraph>
</subSubSection>
<subSubSection id="C383651D6439834F20E1F9F0FCF44DE4" lastPageId="8" lastPageNumber="99" pageId="6" pageNumber="97" type="description">
<paragraph id="8B2636966439834120E1F9F0FA8C4BD9" blockId="6.[151,1437,1433,2036]" pageId="6" pageNumber="97">
<emphasis id="B9EDEA846439834120E1F9F0FE254B32" bold="true" box="[199,443,1613,1639]" pageId="6" pageNumber="97">Description of adult</emphasis>
.
<emphasis id="B9EDEA846439834121EAF9F3FDF04B32" box="[460,622,1614,1639]" italics="true" pageId="6" pageNumber="97">Measurements</emphasis>
. Body length: 315 (
<typeStatus id="5422883464398341237CF9F0FC5E4B32" box="[858,960,1613,1639]" pageId="6" pageNumber="97" type="holotype">holotype</typeStatus>
, female), 315, 332, 298 (three
<typeStatus id="5422883464398341250CF9F3FA064B3D" box="[1322,1432,1614,1640]" pageId="6" pageNumber="97" type="paratype">paratypes</typeStatus>
, male and two female, respectively); notogaster width: 232 (
<typeStatus id="54228834643983412314F9CCFC044BD9" box="[818,922,1649,1676]" pageId="6" pageNumber="97" type="holotype">holotype</typeStatus>
), 232, 249, 240 (three
<typeStatus id="542288346439834124B0F9CFFA994BD9" box="[1174,1287,1650,1676]" pageId="6" pageNumber="97" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="8B2636966439834120E1F92BFDD64BA2" blockId="6.[151,1437,1433,2036]" pageId="6" pageNumber="97">
<emphasis id="B9EDEA846439834120E1F92BFEDB4BFA" box="[199,325,1686,1711]" italics="true" pageId="6" pageNumber="97">Integument</emphasis>
. Body color brown. Body surface heavily granulate (diameter of granule up to 1). Lateral side of body between bothridium and acetabula IIV with microgranulate cerotegument. Leg femora IIV and trochanters III, IV partially tuberculate and striate.
</paragraph>
<paragraph id="8B2636966439834120E1F8BCFC044AFE" blockId="6.[151,1437,1433,2036]" pageId="6" pageNumber="97">
<emphasis id="B9EDEA846439834120E1F8BCFEDA4A4E" box="[199,324,1793,1819]" italics="true" pageId="6" pageNumber="97">Prodorsum</emphasis>
(
<figureCitation id="13A22A13643983412173F8BCFE4C4A4E" box="[341,466,1793,1819]" captionStart="FIGURE 4" captionStartId="7.[151,250,1957,1981]" captionTargetBox="[170,1422,717,1925]" captionTargetId="figure-273@7.[151,1436,705,1934]" captionTargetPageId="7" captionText="FIGURE 4. Stictozetes ihaguensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs omitted); C—lateral view (pteromorph, gnathosoma and legs omitted); D—posterior view (pteromorphs omitted). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.4471648" httpUri="https://zenodo.org/record/4471648/files/figure.png" pageId="6" pageNumber="97">Fig. 4A, C</figureCitation>
). Rostrum rounded. Lamellar and sublamellar lines thin, parallel, curving backwards. Lateral structure
<emphasis id="B9EDEA84643983412172F89BFEF94A6A" box="[340,359,1830,1855]" italics="true" pageId="6" pageNumber="97">N</emphasis>
and ridges
<emphasis id="B9EDEA846439834121C2F89BFE6B4A6A" box="[484,501,1830,1855]" italics="true" pageId="6" pageNumber="97">E</emphasis>
,
<emphasis id="B9EDEA84643983412224F89BFD8C4A6A" box="[514,530,1830,1855]" italics="true" pageId="6" pageNumber="97">T</emphasis>
well visible. Rostral (2428) and lamellar (1216) setae setiform, slightly barbed. Interlamellar seta (46) setiform, thin, smooth. Bothridial seta (7377) with long stalk and elongate, narrowly unilaterally dilated, barbed head, pointed apically. Dorsosejugal porose area rounded (6), located posterolateral to interlamellar seta. Dorsophragma distinctly elongated longitudinally.
</paragraph>
<paragraph id="8B2636966439834020E1F80BFCEB4C48" blockId="6.[151,1437,1433,2036]" lastBlockId="7.[151,1437,151,687]" lastPageId="7" lastPageNumber="98" pageId="6" pageNumber="97">
<emphasis id="B9EDEA846439834120E1F80BFEDD4A9A" box="[199,323,1974,1999]" italics="true" pageId="6" pageNumber="97">Notogaster</emphasis>
(
<figureCitation id="13A22A13643983412173F808FE644A85" box="[341,506,1973,2000]" captionStart="FIGURE 4" captionStartId="7.[151,250,1957,1981]" captionTargetBox="[170,1422,717,1925]" captionTargetId="figure-273@7.[151,1436,705,1934]" captionTargetPageId="7" captionText="FIGURE 4. Stictozetes ihaguensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs omitted); C—lateral view (pteromorph, gnathosoma and legs omitted); D—posterior view (pteromorphs omitted). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.4471648" httpUri="https://zenodo.org/record/4471648/files/figure.png" pageId="6" pageNumber="97">Fig. 4A, C, D</figureCitation>
). Dorsosejugal suture complete. Ten pairs of short (812), setiform, thin, smooth setae developed. Three pairs of rounded porose areas poorly visible (
<emphasis id="B9EDEA84643983412393F867FC4A4AA1" box="[949,980,2010,2036]" italics="true" pageId="6" pageNumber="97">Aa</emphasis>
and
<emphasis id="B9EDEA84643983412436F864FBB14AA6" box="[1040,1071,2009,2035]" italics="true" pageId="6" pageNumber="97">A1</emphasis>
, 68;
<emphasis id="B9EDEA8464398341245DF864FB044AA6" box="[1147,1178,2009,2035]" italics="true" pageId="6" pageNumber="97">A3</emphasis>
, 24). Porose area
<emphasis id="B9EDEA8464398341255BF867FA024AA1" box="[1405,1436,2010,2036]" italics="true" pageId="6" pageNumber="97">Aa</emphasis>
removed from pteromorphal hinge, located between setae
<emphasis id="B9EDEA8464388340230CFF2AFCDE4DE4" box="[810,832,151,177]" italics="true" pageId="7" pageNumber="98">la</emphasis>
and
<emphasis id="B9EDEA8464388340235EFF2AFC0A4DE4" box="[888,916,151,177]" italics="true" pageId="7" pageNumber="98">lm</emphasis>
(clearly closer to
<emphasis id="B9EDEA84643883402444FF2AFBE04DE4" box="[1122,1150,151,177]" italics="true" pageId="7" pageNumber="98">lm</emphasis>
). Median pore present in both sexes, represented by two (four in one female) foveae, located between
<emphasis id="B9EDEA846438834023D6FF06FC604D80" box="[1008,1022,187,213]" italics="true" pageId="7" pageNumber="98">h</emphasis>
<subScript id="171D34D36438834023D8FF70FB984D8E" attach="left" box="[1022,1030,205,219]" fontSize="6" pageId="7" pageNumber="98">2</subScript>
. Opisthonotal gland opening and all lyrifissures (except
<emphasis id="B9EDEA84643883402150FF62FE124DAD" box="[374,396,223,248]" italics="true" pageId="7" pageNumber="98">ip</emphasis>
not observed) distinct:
<emphasis id="B9EDEA846438834022BEFF62FD224DAC" box="[664,700,223,249]" italics="true" pageId="7" pageNumber="98">gla</emphasis>
located lateral to
<emphasis id="B9EDEA846438834023AEFF62FC364DAC" box="[904,936,223,249]" italics="true" pageId="7" pageNumber="98">A1</emphasis>
and removed from it;
<emphasis id="B9EDEA84643883402481FF62FB5D4DAD" box="[1191,1219,223,248]" italics="true" pageId="7" pageNumber="98">im</emphasis>
anterolateral to
<emphasis id="B9EDEA8464388340255BFF62FA034DAC" box="[1405,1437,223,249]" italics="true" pageId="7" pageNumber="98">A1</emphasis>
and removed from it;
<emphasis id="B9EDEA846438834021ADFEBEFE3F4C48" box="[395,417,259,285]" italics="true" pageId="7" pageNumber="98">ih</emphasis>
and
<emphasis id="B9EDEA846438834021FEFEBEFE674C49" box="[472,505,259,284]" italics="true" pageId="7" pageNumber="98">ips</emphasis>
close to each other, anterior to
<emphasis id="B9EDEA8464388340237EFEBEFCF84C49" box="[856,870,259,284]" italics="true" pageId="7" pageNumber="98">p</emphasis>
<subScript id="171D34D3643883402340FEA8FCF04C76" attach="left" box="[870,878,277,291]" fontSize="6" pageId="7" pageNumber="98">3</subScript>
.
</paragraph>
<paragraph id="8B2636966438834020E1FE9AFF754CF8" blockId="7.[151,1437,151,687]" pageId="7" pageNumber="98">
<emphasis id="B9EDEA846438834020E1FE9AFEC84C14" box="[199,342,295,321]" italics="true" pageId="7" pageNumber="98">Gnathosoma</emphasis>
. Generally, similar to that of
<taxonomicName id="4C994D15643883402287FE9AFC454C14" authority="Ermilov &amp; Hugo-Coetzee &amp; Khaustov, 2021" authorityName="Ermilov &amp; Hugo-Coetzee &amp; Khaustov" authorityYear="2021" box="[673,987,295,321]" class="Arachnida" family="Galumnidae" genus="Pilogalumna" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="hogsbackensis" status="sp. nov.">
<emphasis id="B9EDEA84643883402287FE9AFC454C14" box="[673,987,295,321]" italics="true" pageId="7" pageNumber="98">Pilogalumna hogsbackensis</emphasis>
</taxonomicName>
<emphasis id="B9EDEA846438834023C4FE9AFBA24C14" bold="true" box="[994,1084,295,321]" pageId="7" pageNumber="98">
<taxonomicNameLabel id="A2DE57FF6438834023C4FE9AFBA24C14" box="[994,1084,295,321]" pageId="7" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Size of subcapitulum: 73 × 65. Subcapitular (
<emphasis id="B9EDEA84643883402110FEF6FEDA4C31" box="[310,324,331,356]" italics="true" pageId="7" pageNumber="98">a</emphasis>
, 16;
<emphasis id="B9EDEA8464388340215BFEF6FE0F4C31" box="[381,401,331,356]" italics="true" pageId="7" pageNumber="98">m</emphasis>
, 12;
<emphasis id="B9EDEA846438834021EDFEF6FE474C30" box="[459,473,331,357]" italics="true" pageId="7" pageNumber="98">h</emphasis>
, 68) and adoral (8) setae setiform, roughened;
<emphasis id="B9EDEA846438834023DFFEF6FB994C31" box="[1017,1031,331,356]" italics="true" pageId="7" pageNumber="98">a</emphasis>
thicker than others. Length of palp: 57. Postpalpal seta (4) thorn-like, smooth. Length of chelicera: 94. Cheliceral setae (
<emphasis id="B9EDEA8464388340244FFED2FB0F4CDC" box="[1129,1169,367,393]" italics="true" pageId="7" pageNumber="98">cha</emphasis>
, 36;
<emphasis id="B9EDEA846438834024F6FED2FB664CDC" box="[1232,1272,367,393]" italics="true" pageId="7" pageNumber="98">chb</emphasis>
, 24) setiform, barbed.
</paragraph>
<paragraph id="8B2636966438834020E1FE0AFD334F4C" blockId="7.[151,1437,151,687]" pageId="7" pageNumber="98">
<emphasis id="B9EDEA846438834020E1FE0AFD0E4C85" box="[199,656,439,465]" italics="true" pageId="7" pageNumber="98">Epimeral and lateral podosomal regions</emphasis>
(
<figureCitation id="13A22A13643883402286FE0AFC844C84" box="[672,794,439,465]" captionStart="FIGURE 4" captionStartId="7.[151,250,1957,1981]" captionTargetBox="[170,1422,717,1925]" captionTargetId="figure-273@7.[151,1436,705,1934]" captionTargetPageId="7" captionText="FIGURE 4. Stictozetes ihaguensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs omitted); C—lateral view (pteromorph, gnathosoma and legs omitted); D—posterior view (pteromorphs omitted). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.4471648" httpUri="https://zenodo.org/record/4471648/files/figure.png" pageId="7" pageNumber="98">Fig. 4B, C</figureCitation>
). Epimeral setal formula: 1-1-1-3. Setae (812) setiform, thin, roughened;
<emphasis id="B9EDEA8464388340217EFE66FEEA4CA0" box="[344,372,475,501]" italics="true" pageId="7" pageNumber="98">2a</emphasis>
removed from median region. Pedotectum II rounded in ventral aspect. Discidium triangular. Circumpedal carina long, thick, directed to
<emphasis id="B9EDEA84643883402258FE42FD384F4C" box="[638,678,511,537]" italics="true" pageId="7" pageNumber="98">PdI</emphasis>
.
</paragraph>
<paragraph id="8B2636966438834020E1FD9EFA114FFC" blockId="7.[151,1437,151,687]" pageId="7" pageNumber="98">
<emphasis id="B9EDEA846438834020E1FD9EFE0C4F69" box="[199,402,547,573]" italics="true" pageId="7" pageNumber="98">Anogenital region</emphasis>
(
<figureCitation id="13A22A13643883402184FD9EFE784F68" box="[418,486,547,573]" captionStart="FIGURE 4" captionStartId="7.[151,250,1957,1981]" captionTargetBox="[170,1422,717,1925]" captionTargetId="figure-273@7.[151,1436,705,1934]" captionTargetPageId="7" captionText="FIGURE 4. Stictozetes ihaguensis sp. nov., adult: A—dorsal view; B—ventral view (gnathosoma and legs omitted); C—lateral view (pteromorph, gnathosoma and legs omitted); D—posterior view (pteromorphs omitted). Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.4471648" httpUri="https://zenodo.org/record/4471648/files/figure.png" pageId="7" pageNumber="98">Fig. 4</figureCitation>
BD). Genital (68), aggenital (1012), anal (1012), and adanal (1012) setae setiform, thin, roughened. Anterior edge of genital plate with three setae. Aggenital seta located between genital and anal apertures, closer to the former. Adanal lyrifissure located close and parallel to anal plate. Adanal setae
<emphasis id="B9EDEA8464388340256FFDD6FAFB4FD0" box="[1353,1381,619,645]" italics="true" pageId="7" pageNumber="98">ad</emphasis>
<subScript id="171D34D3643883402543FDC0FAF34FDE" attach="left" box="[1381,1389,637,651]" fontSize="6" pageId="7" pageNumber="98">1</subScript>
and
<emphasis id="B9EDEA846438834020B1FD32FF2D4FFC" box="[151,179,655,681]" italics="true" pageId="7" pageNumber="98">ad</emphasis>
<subScript id="171D34D3643883402095FD1CFF254FFA" attach="left" box="[179,187,673,687]" fontSize="6" pageId="7" pageNumber="98">2</subScript>
posterior,
<emphasis id="B9EDEA84643883402115FD32FED14FFC" box="[307,335,655,681]" italics="true" pageId="7" pageNumber="98">ad</emphasis>
<subScript id="171D34D3643883402169FD1CFEC94FFA" attach="left" box="[335,343,673,687]" fontSize="6" pageId="7" pageNumber="98">3</subScript>
lateral to
<emphasis id="B9EDEA846438834021EEFD32FE724FFC" box="[456,492,655,681]" italics="true" pageId="7" pageNumber="98">iad</emphasis>
and anal plate. Distance
<emphasis id="B9EDEA84643883402323FD32FCBF4FFC" box="[773,801,655,681]" italics="true" pageId="7" pageNumber="98">ad</emphasis>
<subScript id="171D34D3643883402307FD1CFCB74FFA" attach="left" box="[801,809,673,687]" fontSize="6" pageId="7" pageNumber="98">1</subScript>
<emphasis id="B9EDEA84643883402311FD32FCCD4FFC" box="[823,851,655,681]" italics="true" pageId="7" pageNumber="98">ad</emphasis>
<subScript id="171D34D3643883402375FD1CFCC54FFA" attach="left" box="[851,859,673,687]" fontSize="6" pageId="7" pageNumber="98">2</subScript>
shorter than
<emphasis id="B9EDEA846438834023C8FD32FB944FFC" box="[1006,1034,655,681]" italics="true" pageId="7" pageNumber="98">ad</emphasis>
<subScript id="171D34D364388340242CFD1CFB8C4FFA" attach="left" box="[1034,1042,673,687]" fontSize="6" pageId="7" pageNumber="98">2</subScript>
<emphasis id="B9EDEA84643883402406FD32FBA24FFC" box="[1056,1084,655,681]" italics="true" pageId="7" pageNumber="98">ad</emphasis>
<subScript id="171D34D364388340241AFD1CFBDA4FFA" attach="left" box="[1084,1092,673,687]" fontSize="6" pageId="7" pageNumber="98">3</subScript>
. Postanal porose area absent.
</paragraph>
<caption id="DFE6661E6438834020B1F818FB674AB7" ID-DOI="http://doi.org/10.5281/zenodo.4471648" ID-Zenodo-Dep="4471648" httpUri="https://zenodo.org/record/4471648/files/figure.png" pageId="7" pageNumber="98" startId="7.[151,250,1957,1981]" targetBox="[170,1422,717,1925]" targetPageId="7">
<paragraph id="8B2636966438834020B1F818FB674AB7" blockId="7.[151,1437,1957,2018]" pageId="7" pageNumber="98">
<emphasis id="B9EDEA846438834020B1F818FE8C4AE8" bold="true" box="[151,274,1957,1981]" pageId="7" pageNumber="98">FIGURE 4.</emphasis>
<taxonomicName id="4C994D1564388340213EF81BFE6B4AEB" authority="Ermilov &amp; Hugo-Coetzee &amp; Khaustov, 2021" authorityName="Ermilov &amp; Hugo-Coetzee &amp; Khaustov" authorityYear="2021" box="[280,501,1958,1982]" class="Arachnida" family="Galumnidae" genus="Stictozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="98" phylum="Arthropoda" rank="species" species="ihaguensis" status="sp. nov.">
<emphasis id="B9EDEA8464388340213EF81BFE6B4AEB" box="[280,501,1958,1982]" italics="true" pageId="7" pageNumber="98">Stictozetes ihaguensis</emphasis>
</taxonomicName>
<emphasis id="B9EDEA846438834021DDF81BFDD44AEB" bold="true" box="[507,586,1958,1982]" pageId="7" pageNumber="98">
<taxonomicNameLabel id="A2DE57FF6438834021DDF81BFDD44AEB" box="[507,586,1958,1982]" pageId="7" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult: A—dorsal view; B—ventral view (gnathosoma and legs omitted); C—lateral view (pteromorph, gnathosoma and legs omitted); D—posterior view (pteromorphs omitted). Scale bar 50 μm.
</paragraph>
</caption>
<paragraph id="8B2636966437834F20E1FF2AFCF44DE4" blockId="8.[151,1437,151,718]" box="[199,874,151,177]" pageId="8" pageNumber="99">
<emphasis id="B9EDEA846437834F20E1FF2AFF624DE5" box="[199,252,151,176]" italics="true" pageId="8" pageNumber="99">Legs</emphasis>
. Similar to that of
<taxonomicName id="4C994D156437834F21F6FF2AFC974DE4" authority="Ermilov &amp; Hugo-Coetzee &amp; Khaustov, 2021" authorityName="Ermilov &amp; Hugo-Coetzee &amp; Khaustov" authorityYear="2021" box="[464,777,151,177]" class="Arachnida" family="Galumnidae" genus="Pilogalumna" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="hogsbackensis" status="sp. nov.">
<emphasis id="B9EDEA846437834F21F6FF2AFC974DE4" box="[464,777,151,177]" italics="true" pageId="8" pageNumber="99">Pilogalumna hogsbackensis</emphasis>
</taxonomicName>
<emphasis id="B9EDEA846437834F2336FF2AFCF44DE4" bold="true" box="[784,874,151,177]" pageId="8" pageNumber="99">
<taxonomicNameLabel id="A2DE57FF6437834F2336FF2AFCF44DE4" box="[784,874,151,177]" pageId="8" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C383651D6437834F20E1FF07FAD24CF8" pageId="8" pageNumber="99" type="materials_examined">
<paragraph id="8B2636966437834F20E1FF07FC244C17" blockId="8.[151,1437,151,718]" pageId="8" pageNumber="99">
<emphasis id="B9EDEA846437834F20E1FF07FE2C4D81" bold="true" box="[199,434,186,212]" pageId="8" pageNumber="99">Material examined.</emphasis>
<materialsCitation id="3BF13CCB6437834F219CFF06FC244C17" ID-GBIF-Occurrence="3028881302" collectingDate="2019-09-14" collectionCode="V" collectorName="A. Khaustov &amp; S. G. Ermilov &amp; J. A. Neethling &amp; E. A. Hugo-Coetzee &amp; A. A. Khaustov" country="South Africa" county="Amathole mountains" latitude="-32.589333" location="Hogsback" longLatPrecision="1" longitude="26.960695" municipality="Hogsback State Forest" pageId="8" pageNumber="99" specimenCount="2" stateProvince="Eastern Cape" typeStatus="holotype">
<typeStatus id="542288346437834F219CFF06FDBC4D80" box="[442,546,187,213]" pageId="8" pageNumber="99" type="holotype">Holotype</typeStatus>
(female) and three paratypes (one male and two females):
<collectingCountry id="F38E76066437834F24E6FF06FACF4D80" box="[1216,1361,187,213]" name="South Africa" pageId="8" pageNumber="99">South Africa</collectingCountry>
,
<collectingCounty id="62474E1A6437834F257AFF06FED34DAC" pageId="8" pageNumber="99">Amathole mountains</collectingCounty>
in the
<collectingRegion id="495DF8746437834F2186FF62FD384DAC" box="[416,678,223,249]" country="South Africa" name="Eastern Cape" pageId="8" pageNumber="99">Eastern Cape Province</collectingRegion>
,
<collectingMunicipality id="6B42ACEC6437834F2292FF62FC2A4DAC" box="[692,948,223,249]" pageId="8" pageNumber="99">Hogsback State Forest</collectingMunicipality>
at the village of
<location id="8E46604D6437834F245AFF62FB714DAC" LSID="urn:lsid:plazi:treatment:033087806439834F20B1FA90FE444F9B:8E46604D6437834F245AFF62FB714DAC" box="[1148,1263,223,249]" country="South Africa" county="Amathole mountains" latitude="-32.589333" longLatPrecision="1" longitude="26.960695" municipality="Hogsback State Forest" name="Hogsback" pageId="8" pageNumber="99" stateProvince="Eastern Cape">Hogsback</location>
,
<geoCoordinate id="EEAD50516437834F24DAFF63FA064DAC" box="[1276,1432,222,249]" degrees="32" direction="south" minutes="35" orientation="latitude" pageId="8" pageNumber="99" precision="1" seconds="21.6" value="-32.589333">32°3521.6”S</geoCoordinate>
,
<geoCoordinate id="EEAD50516437834F20B1FEBFFEA64C48" box="[151,312,258,285]" degrees="26" direction="east" minutes="57" orientation="longitude" pageId="8" pageNumber="99" precision="1" seconds="38.5" value="26.960695">26°5738.5E</geoCoordinate>
, indigenous
<location id="8E46604D6437834F21E5FEBEFDFC4C48" LSID="urn:lsid:plazi:treatment:033087806439834F20B1FA90FE444F9B:8E46604D6437834F21E5FEBEFDFC4C48" box="[451,610,259,285]" country="South Africa" county="Amathole mountains" latitude="-32.589333" longLatPrecision="1" longitude="26.960695" municipality="Hogsback State Forest" name="Afro-montane" pageId="8" pageNumber="99" stateProvince="Eastern Cape">Afro-montane</location>
mixed forest, in coniferous litter,
<date id="FF2710566437834F23FEFEBEFBC94C48" box="[984,1111,259,285]" pageId="8" pageNumber="99" value="2019-09-14">
<collectingDate id="EF63E9BE6437834F23FEFEBEFBC94C48" box="[984,1111,259,285]" pageId="8" pageNumber="99" value="2019-09-14">14.IX.2019</collectingDate>
</date>
(collected by
<collectionCode id="ED88AE536437834F24D5FEBEFA974C48" box="[1267,1289,259,285]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="99" type="Museum">V</collectionCode>
.
<collectorName id="266C53406437834F252AFEBEFA064C48" box="[1292,1432,259,285]" pageId="8" pageNumber="99">A. Khaustov</collectorName>
,
<collectorName id="266C53406437834F20B1FE9AFEB34C14" box="[151,301,295,321]" pageId="8" pageNumber="99">S.G. Ermilov</collectorName>
,
<collectorName id="266C53406437834F211EFE9AFE434C14" box="[312,477,295,321]" pageId="8" pageNumber="99">J.A. Neethling</collectorName>
,
<collectorName id="266C53406437834F21CEFE9AFD564C14" box="[488,712,295,321]" pageId="8" pageNumber="99">E.A. Hugo-Coetzee</collectorName>
, and
<collectorName id="266C53406437834F2326FE9AFC314C17" box="[768,943,295,322]" pageId="8" pageNumber="99">A.A. Khaustov</collectorName>
).
</materialsCitation>
</paragraph>
<paragraph id="8B2636966437834F20E1FEF6FAD24CF8" blockId="8.[151,1437,151,718]" pageId="8" pageNumber="99">
<emphasis id="B9EDEA846437834F20E1FEF6FE104C31" bold="true" box="[199,398,330,357]" pageId="8" pageNumber="99">Type deposition.</emphasis>
<materialsCitation id="3BF13CCB6437834F21B1FEF6FAD64CF8" ID-GBIF-Occurrence="3028881303" collectionCode="NMB, TSUMZ" country="South Africa" county="National Museum Bloemfontein" location="All" municipality="University Museum of Zoology" pageId="8" pageNumber="99" specimenCount="2" stateProvince="Tyumen'" typeStatus="holotype">
The
<typeStatus id="542288346437834F21EAFEF6FDB04C30" box="[460,558,331,357]" pageId="8" pageNumber="99" type="holotype">holotype</typeStatus>
is deposited in the collection of the
<collectingCounty id="62474E1A6437834F23FCFEF6FAD34C30" box="[986,1357,331,357]" pageId="8" pageNumber="99">National Museum Bloemfontein</collectingCounty>
,
<collectingCountry id="F38E76066437834F257DFEF6FF414CDC" name="South Africa" pageId="8" pageNumber="99">South Africa</collectingCountry>
(
<collectionCode id="ED88AE536437834F20C8FED2FEAD4CDC" box="[238,307,367,393]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="8" pageNumber="99">NMB</collectionCode>
); three
<typeStatus id="542288346437834F21A3FED2FE6E4CDC" box="[389,496,367,393]" pageId="8" pageNumber="99" type="paratype">paratypes</typeStatus>
are deposited in the collection of the
<collectingRegion id="495DF8746437834F23B5FED2FBB04CDC" box="[915,1070,367,393]" country="Russia" name="Tyumen'" pageId="8" pageNumber="99">Tyumen State</collectingRegion>
<collectingMunicipality id="6B42ACEC6437834F2412FED2FA064CDC" box="[1076,1432,367,393]" pageId="8" pageNumber="99">University Museum of Zoology</collectingMunicipality>
,
<collectingRegion id="495DF8746437834F20B1FE2EFF6B4CF8" box="[151,245,403,429]" country="Russia" name="Tyumen'" pageId="8" pageNumber="99">Tyumen</collectingRegion>
,
<collectingCountry id="F38E76066437834F2127FE2EFED24CF8" box="[257,332,403,429]" name="Russia" pageId="8" pageNumber="99">Russia</collectingCountry>
(
<collectionCode id="ED88AE536437834F217CFE2EFE214CF8" box="[346,447,403,429]" pageId="8" pageNumber="99">TSUMZ</collectionCode>
).
<location id="8E46604D6437834F21F6FE2EFE6A4CF8" LSID="urn:lsid:plazi:treatment:033087806439834F20B1FA90FE444F9B:8E46604D6437834F21F6FE2EFE6A4CF8" box="[464,500,403,429]" country="South Africa" county="National Museum Bloemfontein" municipality="University Museum of Zoology" name="All" pageId="8" pageNumber="99" stateProvince="Tyumen'">All</location>
specimens are preserved in 70% solution of ethanol with a drop of glycerol
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C383651D6437834F20E1FE0AFF4A4F4C" pageId="8" pageNumber="99" type="etymology">
<paragraph id="8B2636966437834F20E1FE0AFF4A4F4C" blockId="8.[151,1437,151,718]" pageId="8" pageNumber="99">
<emphasis id="B9EDEA846437834F20E1FE0AFED04C84" bold="true" box="[199,334,439,465]" pageId="8" pageNumber="99">Etymology.</emphasis>
The specific name
<taxonomicName id="4C994D156437834F2216FE0AFD364C84" authorityName="Ermilov &amp; Hugo-Coetzee &amp; Khaustov" authorityYear="2021" box="[560,680,439,465]" class="Arachnida" family="Galumnidae" genus="Stictozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="ihaguensis">
<emphasis id="B9EDEA846437834F2216FE0AFD364C84" box="[560,680,439,465]" italics="true" pageId="8" pageNumber="99">ihaguensis</emphasis>
</taxonomicName>
, is derived from the Xhosa word, ihagu, meaning pig or hog. Hogsback is supposedly named after one of the mountain peaks overlooking the town which resembles a hogs back.
</paragraph>
</subSubSection>
<subSubSection id="C383651D6437834F20E1FD9EFE444F9B" pageId="8" pageNumber="99" type="discussion">
<paragraph id="8B2636966437834F20E1FD9EFE444F9B" blockId="8.[151,1437,151,718]" pageId="8" pageNumber="99">
<emphasis id="B9EDEA846437834F20E1FD9EFEA54F68" bold="true" box="[199,315,547,573]" pageId="8" pageNumber="99">Remarks.</emphasis>
<taxonomicName id="4C994D156437834F2164FD9EFDA84F68" authority="Ermilov &amp; Hugo-Coetzee &amp; Khaustov, 2021" authorityName="Ermilov &amp; Hugo-Coetzee &amp; Khaustov" authorityYear="2021" box="[322,566,547,573]" class="Arachnida" family="Galumnidae" genus="Stictozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="ihaguensis" status="sp. nov.">
<emphasis id="B9EDEA846437834F2164FD9EFDA84F68" box="[322,566,547,573]" italics="true" pageId="8" pageNumber="99">Stictozetes ihaguensis</emphasis>
</taxonomicName>
<emphasis id="B9EDEA846437834F221BFD9EFD084F68" bold="true" box="[573,662,547,573]" pageId="8" pageNumber="99">
<taxonomicNameLabel id="A2DE57FF6437834F221BFD9EFD084F68" box="[573,662,547,573]" pageId="8" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically most similar to
<taxonomicName id="4C994D156437834F2404FD9EFA024F6B" authority="(Berlese, 1916)" baseAuthorityName="Berlese" baseAuthorityYear="1916" box="[1058,1436,547,574]" class="Arachnida" family="Galumnidae" genus="Stictozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="99" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="B9EDEA846437834F2404FD9EFB744F68" box="[1058,1258,547,573]" italics="true" pageId="8" pageNumber="99">Stictozetes scaber</emphasis>
(
<bibRefCitation id="EF084B676437834F24DEFD9EFA0A4F6B" author="Berlese, A." box="[1272,1428,547,574]" pageId="8" pageNumber="99" pagination="19 - 67" refId="ref6325" refString="Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12, 19 - 67." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
)
</taxonomicName>
from
<collectingRegion id="495DF8746437834F20F2FDFAFEBD4F34" box="[212,291,583,609]" country="Ethiopia" name="Sumale" pageId="8" pageNumber="99">Somali</collectingRegion>
in the presence of bothridial seta with dilated head and minute lamellar seta, and the absence of reticulate pteromorph and sexual dimorphism in presence of median pore and notogastral porose areas, but differs from the latter by the bothridial seta with narrowly dilated bothridial head (versus broadly dilated) and the presence of median pore (versus absent).
</paragraph>
</subSubSection>
</treatment>
</document>