180 lines
19 KiB
XML
180 lines
19 KiB
XML
<document id="EA4DE4D784AED14617E24DB13826C126" ID-CLB-Dataset="22833" ID-DOI="10.11646/zootaxa.4766.1.1" ID-GBIF-Dataset="ed4d1bf6-284a-4898-952a-58943d5a3026" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763907" ID-ZooBank="urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1587670592452" checkinUser="carolina" docAuthor="Ketwetsuriya, Chatchalerm, Karapunar, Baran, Charoentitirat, Thasinee & Nützel, Al- Exander" docDate="2020" docId="03B587AB4F3B1573FF517F2EFAFDFB96" docLanguage="en" docName="zootaxa.4766.1.1.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Biarmeaspira mazaevi Ketwetsuriya & Karapunar & Charoentitirat & Nützel 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="14" masterDocId="FF8CFFD34F37157EFFC6787EFFCBFF85" masterDocTitle="Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane" masterLastPageNumber="47" masterPageNumber="1" pageNumber="13" updateTime="1698836971451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="A9B42E9C49DC755EC6A661F447D64895" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="09114DA26F049F42E61C374EDCC4EDBF">
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<mods:title id="D975A61914D8A3865DC85EE25EE64E29">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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</mods:titleInfo>
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<mods:name id="DCF2F1C60B5DD5AB1302349050A0F9CC" type="personal">
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<mods:roleTerm id="B0D88345A269530A5FE44342D206DABE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D29AD873F6932B4E5355DCFAA9710CC4">Ketwetsuriya, Chatchalerm</mods:namePart>
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</mods:name>
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<mods:name id="BE1D1E196047373F4E4655808DDA9CA6" type="personal">
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<mods:roleTerm id="274ED3C46271C7671B5AFBC75D097FC3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E95EB8D2692B4A2E43835D3D059C3C19">Karapunar, Baran</mods:namePart>
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</mods:name>
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<mods:name id="52B17C285B9B62AFC487E47C032FD12F" type="personal">
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<mods:roleTerm id="41A9A499328A8907820A9A01A3748FB6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5F8B78321DB479DBB9D8CCC12CD032AA">Charoentitirat, Thasinee</mods:namePart>
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</mods:name>
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<mods:name id="C1ECB64240DD34B764A3284D60982DC9" type="personal">
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<mods:role id="28421C192123AAE6AFDD6C12FA5D08D8">
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<mods:roleTerm id="9588ACBD16CC07DC035E93005D3401A1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9C504069A35159396E1EC52E2BC02B96">Nützel, Al- Exander</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="EEC3F5FEE8D40EC6666D45059136347C">text</mods:typeOfResource>
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<mods:title id="715FFC3DC2DB6FBE811D83EA8EB99E1C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="863ADC8E2871E49CA4B3276866DE58BB">
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<mods:date id="8F788A8AE7F0AB7625E0443C78C24477">2020</mods:date>
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<mods:detail id="4E6F663BFF04ACF8A2C7B7CD30F22CE9" type="pubDate">
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<mods:number id="F5059072F79149D79E5562517AEB7E20">2020-04-17</mods:number>
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</mods:detail>
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<mods:detail id="42BE0C0FF4BDC2F17C2D1166D2A10E27" type="volume">
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<mods:number id="F7E53923F8B479199AD33A3417E6947E">4766</mods:number>
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</mods:detail>
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<mods:detail id="6C6A9065BB464A95594F00D494765174" type="issue">
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<mods:number id="714405DE84C06E4982BB4EE434B2E5AD">1</mods:number>
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<mods:start id="8945EF717FCF13DBA9303B2C0FC3188E">1</mods:start>
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<mods:end id="C67E381E5E17B358C35359F4296EFDDA">47</mods:end>
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<mods:classification id="E7ED247BA869AC90686CC77C92E0A772">journal article</mods:classification>
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<mods:identifier id="5ECF01E0AA66BF8CD02B5F5DFB2F9364" type="CLB-Dataset">22833</mods:identifier>
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<mods:identifier id="486C1D75C3AD4BCBF0A16DCC14B2B26C" type="DOI">10.11646/zootaxa.4766.1.1</mods:identifier>
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<mods:identifier id="321C1D2BC1274C7441B7DC0502B194C7" type="GBIF-Dataset">ed4d1bf6-284a-4898-952a-58943d5a3026</mods:identifier>
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<mods:identifier id="ABB9FF786AF14445D76FD47CDF094A49" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="BA72D2E9B77ACDBEAC97AEFFE9274E22" type="Zenodo-Dep">3763907</mods:identifier>
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<mods:identifier id="A78086D0BFD4EE23B4F4093F0ED2A528" type="ZooBank">urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE</mods:identifier>
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</mods:mods>
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<treatment id="03B587AB4F3B1573FF517F2EFAFDFB96" ID-GBIF-Taxon="163737119" LSID="urn:lsid:plazi:treatment:03B587AB4F3B1573FF517F2EFAFDFB96" httpUri="http://treatment.plazi.org/id/03B587AB4F3B1573FF517F2EFAFDFB96" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<subSubSection id="C30665364F3B1572FF517F2EFDC1F8EF" box="[151,522,1872,1898]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BA336BD4F3B1572FF517F2EFDC1F8EF" blockId="12.[151,522,1872,1934]" box="[151,522,1872,1898]" pageId="12" pageNumber="13">
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<heading id="D0EB81D14F3B1572FF517F2EFDC1F8EF" bold="true" box="[151,522,1872,1898]" fontSize="10" level="2" pageId="12" pageNumber="13" reason="2">
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<emphasis id="B968EAAF4F3B1572FF517F2EFDC1F8EF" bold="true" box="[151,522,1872,1898]" pageId="12" pageNumber="13">
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<taxonomicName id="4C1C4D3E4F3B1572FF517F2EFE6EF8EF" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[151,421,1872,1898]" class="Gastropoda" family="Raphistomidae" genus="Biarmeaspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Euomphalina" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="mazaevi" status="sp. nov.">
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<emphasis id="B968EAAF4F3B1572FF517F2EFE6EF8EF" bold="true" box="[151,421,1872,1898]" italics="true" pageId="12" pageNumber="13">Biarmeaspira mazaevi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57D44F3B1572FE6A7F2EFDC1F8EF" box="[428,522,1872,1898]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F3B1572FF517F0AFF31F80B" box="[151,250,1908,1934]" pageId="12" pageNumber="13" type="description">
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<paragraph id="8BA336BD4F3B1572FF517F0AFF31F80B" blockId="12.[151,522,1872,1934]" box="[151,250,1908,1934]" pageId="12" pageNumber="13">
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(
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<figureCitation id="13272A384F3B1572FF667F0AFF3AF80B" box="[160,241,1908,1934]" captionStart="FIGURE 10" captionStartId="14.[151,250,989,1014]" captionTargetBox="[152,1436,453,964]" captionTargetId="figure@14.[152,1436,453,964]" captionTargetPageId="14" captionText="FIGURE 10. Biarmeaspira mazaevi sp. nov. ESKU-19-LP 1, holotype. All scale bars represent 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763927" httpUri="https://zenodo.org/record/3763927/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<paragraph id="8BA336BD4F3B1572FF517FC2FD6FF87F" blockId="12.[151,1432,1980,2042]" pageId="12" pageNumber="13">
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<subSubSection id="C30665364F3B1572FF517FC2FA53F853" box="[151,1432,1980,2006]" pageId="12" pageNumber="13" type="etymology">
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<emphasis id="B968EAAF4F3B1572FF517FC2FED5F853" bold="true" box="[151,286,1980,2006]" pageId="12" pageNumber="13">Etymology.</emphasis>
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After Alexey Mazaev for his work on the Late Palaeozoic gastropods and establishment of this genus.
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</subSubSection>
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<subSubSection id="C30665364F3B1572FF017F9EFD6FF87F" box="[199,676,2016,2042]" pageId="12" pageNumber="13" type="materials_examined">
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<materialsCitation id="3B743CE04F3B1572FF017F9EFD6FF87F" ID-GBIF-Occurrence="4433193301" box="[199,676,2016,2042]" collectionCode="ESKU" pageId="12" pageNumber="13" specimenCode="ESKU-19-LP 1" specimenCount="1" typeStatus="holotype">
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<emphasis id="B968EAAF4F3B1572FF017F9EFEF0F87F" bold="true" box="[199,315,2016,2042]" pageId="12" pageNumber="13">
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<typeStatus id="54A7881F4F3B1572FF017F9EFEFDF87F" box="[199,310,2016,2042]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCount id="9D1AFD344F3B1572FE847F9EFE2DF87F" box="[322,486,2016,2042]" pageId="12" pageNumber="13">One specimen</specimenCount>
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:
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<specimenCode id="DBBA9EC64F3B1572FE367F9EFD55F87F" box="[496,670,2016,2042]" pageId="12" pageNumber="13">ESKU-19-LP 1</specimenCode>
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.
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</materialsCitation>
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</subSubSection>
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</paragraph>
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<paragraph id="8BA336BD4F3A1573FF0178E7FB4CFF36" blockId="13.[151,1437,152,1043]" box="[199,1159,152,179]" pageId="13" pageNumber="14">
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Dimensions (mm): ESKU-19-LP 1: height = 11.0; width =
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<emphasis id="B968EAAF4F3A1573FC9A78E4FCA3FF36" box="[860,872,154,179]" italics="true" pageId="13" pageNumber="14">c</emphasis>
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. 9.5; apical angle = 100º.
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</paragraph>
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<subSubSection id="C30665364F3A1573FF0178C3FE74FE9A" pageId="13" pageNumber="14" type="distribution">
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<paragraph id="8BA336BD4F3A1573FF0178C3FE74FE9A" blockId="13.[151,1437,152,1043]" pageId="13" pageNumber="14">
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<emphasis id="B968EAAF4F3A1573FF0178C3FD6AFF52" bold="true" box="[199,673,188,215]" pageId="13" pageNumber="14">
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<typeStatus id="54A7881F4F3A1573FF0178C3FEC9FF52" box="[199,258,189,215]" pageId="13" pageNumber="14">Type</typeStatus>
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locality and stratigraphical range.
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</emphasis>
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Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about
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<quantity id="4CE49B584F3A1573FEF1789FFEB4FF7E" box="[311,383,225,251]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="14" unit="km" value="13.0">13 km</quantity>
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east of
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<collectingRegion id="49D8F85F4F3A1573FE1C789FFD69FF7E" box="[474,674,225,251]" country="Thailand" name="Lop Buri" pageId="13" pageNumber="14">Lopburi Province</collectingRegion>
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, Central
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<collectingCountry id="F30B762D4F3A1573FCCF789FFCA6FF7E" box="[777,877,225,251]" name="Thailand" pageId="13" pageNumber="14">Thailand</collectingCountry>
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(
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<figureCitation id="13272A384F3A1573FCBB789FFC0BFF7E" box="[893,960,225,251]" captionStart="FIGURE" captionStartId="1.[151,168,1975,2001]" captionTargetBox="[243,1345,1164,1941]" captionTargetId="figure@1.[236,1350,1160,1948]" captionTargetPageId="1" captionText="FIGURE 1. Geological map of the study location at Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about 13 km east of Lopburi Province, Central Thailand." figureDoi="http://doi.org/10.5281/zenodo.3763909" httpUri="https://zenodo.org/record/3763909/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
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), Khao Khad Formation,
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<collectingRegion id="49D8F85F4F3A1573FB21789FFA8DFF7E" box="[1255,1350,225,251]" country="Thailand" name="Sara Buri" pageId="13" pageNumber="14">Saraburi</collectingRegion>
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Group, Middle Permian, Roadian.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F3A1573FF017956FEA1FD2E" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BA336BD4F3A1573FF017956FEA1FD2E" blockId="13.[151,1437,152,1043]" pageId="13" pageNumber="14">
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<emphasis id="B968EAAF4F3A1573FF017956FE92FEC6" bold="true" box="[199,345,296,323]" pageId="13" pageNumber="14">Description.</emphasis>
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Shell moderately high-spired, turbiniform, earliest whorls missing, 4 whorls preserved; early teleoconch whorls convex, ornamented by up to four spiral lirae; later spire whorls with sharp angulation at mid whorl face; upper whorl face above angulation develops a concave subsutural ramp; ramp ornamented with several spiral threads; selenizone narrow, angulated, situated at middle angulation, ornamented with obscure lunulae, bordered by two distinct spiral cords; lower whorl face flat to concave, subvertical inclined about 10 degrees from axis, ornamented with up to eight widely spaced fine spiral lirae; lower whorl face twice as wide as upper whorl face; suture shallow and indistinct; lower part of the lower whorl face ornamented by two strongest spiral cords, representing periphery, interspace between the two cords markedly concave, wide and ornamented with additional 2 fine spiral lirae, lowest cord represents the basal edge; profile near basal edge swollen; base convex, ornamented by numerous widely spaced fine spiral cords that are stronger and more widely spaced than those on ramp and lower whorl face; aperture unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F3A1573FF017ACBFAFDFB96" pageId="13" pageNumber="14" type="discussion">
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<paragraph id="8BA336BD4F3A1573FF017ACBFCA8FC06" blockId="13.[151,1437,152,1043]" pageId="13" pageNumber="14">
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<emphasis id="B968EAAF4F3A1573FF017ACBFEF0FD4A" bold="true" box="[199,315,693,719]" pageId="13" pageNumber="14">Remarks.</emphasis>
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This single specimen is placed in the genus
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<taxonomicName id="4C1C4D3E4F3A1573FCF57AC8FC06FD4A" authorityName="Mazaev" authorityYear="2006" box="[819,973,694,719]" class="Gastropoda" family="Raphistomidae" genus="Biarmeaspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Euomphalina" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F3A1573FCF57AC8FC06FD4A" box="[819,973,694,719]" italics="true" pageId="13" pageNumber="14">Biarmeaspira</emphasis>
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</taxonomicName>
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due to the presence of an angular selenizone and a strong spiral ornament. The early teleoconch is absent and the growth lines and the aperture are poorly preserved so that this generic assignment is not entirely beyond doubt. Another genus having a prominent spiral ornament, a swollen basal edge and an angular selenizone is the Triassic genus
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<taxonomicName id="4C1C4D3E4F3A1573FBDE7B5FFBA4FCBE" box="[1048,1135,801,827]" class="Gastropoda" family="Pleurotomariidae" genus="Sisenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F3A1573FBDE7B5FFBA4FCBE" box="[1048,1135,801,827]" italics="true" pageId="13" pageNumber="14">Sisenna</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C1C4D3E4F3A1573FBB87B5FFB1EFCBE" box="[1150,1237,801,827]" class="Gastropoda" family="Pleurotomariidae" genus="Sisenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F3A1573FBB87B5FFB1EFCBE" box="[1150,1237,801,827]" italics="true" pageId="13" pageNumber="14">Sisenna</emphasis>
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</taxonomicName>
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has a lower spire compared to
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<taxonomicName id="4C1C4D3E4F3A1573FEEC7B38FE0FFCDA" authorityName="Mazaev" authorityYear="2006" box="[298,452,838,863]" class="Gastropoda" family="Raphistomidae" genus="Biarmeaspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Euomphalina" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F3A1573FEEC7B38FE0FFCDA" box="[298,452,838,863]" italics="true" pageId="13" pageNumber="14">Biarmeaspira</emphasis>
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</taxonomicName>
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and it possesses an angulation on the early whorl face which later turns into selenizone during ontogeny, a characteristic that is absent in
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<emphasis id="B968EAAF4F3A1573FD077B14FCA8FC06" box="[705,867,874,899]" italics="true" pageId="13" pageNumber="14">
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<taxonomicName id="4C1C4D3E4F3A1573FD077B14FC94FC06" authorityName="Mazaev" authorityYear="2006" box="[705,863,874,899]" class="Gastropoda" family="Raphistomidae" genus="Biarmeaspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Euomphalina" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">Biarmeaspira</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph id="8BA336BD4F3A1573FF017BF0FAFDFB96" blockId="13.[151,1437,152,1043]" pageId="13" pageNumber="14">
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<taxonomicName id="4C1C4D3E4F3A1573FF017BF0FE0FFC22" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[199,452,910,935]" class="Gastropoda" family="Raphistomidae" genus="Biarmeaspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Euomphalina" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="mazaevi" status="sp. nov.">
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<emphasis id="B968EAAF4F3A1573FF017BF0FE0FFC22" box="[199,452,910,935]" italics="true" pageId="13" pageNumber="14">Biarmeaspira mazaevi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57D44F3A1573FE0C7BF3FDEAFC22" box="[458,545,909,935]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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can be differentiated from other
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<taxonomicName id="4C1C4D3E4F3A1573FC497BF0FBE2FC22" authorityName="Mazaev" authorityYear="2006" box="[911,1065,910,935]" class="Gastropoda" family="Raphistomidae" genus="Biarmeaspira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Euomphalina" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F3A1573FC497BF0FBE2FC22" box="[911,1065,910,935]" italics="true" pageId="13" pageNumber="14">Biarmeaspira</emphasis>
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</taxonomicName>
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species by its ornamentation and by having a swollen basal edge.
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<emphasis id="B968EAAF4F3A1573FE387BCCFDC4FC4E" box="[510,527,946,971]" italics="true" pageId="13" pageNumber="14">B</emphasis>
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|
.?
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<emphasis id="B968EAAF4F3A1573FDEE7BCFFD65FC4E" box="[552,686,945,971]" italics="true" pageId="13" pageNumber="14">loatienensis</emphasis>
|
|
(
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|
<bibRefCitation id="EF8D4B4C4F3A1573FD7A7BCFFC97FC4E" author="Mansuy" box="[700,860,945,971]" firstAuthor="Mansuy" pageId="13" pageNumber="14" pagination="1 - 190" refId="ref26085" refString="Mansuy, H. (1914) Nouvelle contribution a la paleontologie du Yunnan. Memoires du Service Geologique de l'Indochine, 3 (2), 1 - 190." type="journal article" year="1914">Mansuy, 1914</bibRefCitation>
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|
) from the Carboniferous of Loatien,
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|
<collectingRegion id="49D8F85F4F3A1573FB307BCFFA87FC4E" box="[1270,1356,945,971]" country="China" name="Yunnan" pageId="13" pageNumber="14">Yunnan</collectingRegion>
|
|
is very similar in shape, but is ornamented with more prominent spiral ribs on the upper and lateral whorl face.
|
|
<emphasis id="B968EAAF4F3A1573FADB7BA8FAE5FC6A" box="[1309,1326,982,1007]" italics="true" pageId="13" pageNumber="14">B</emphasis>
|
|
.?
|
|
<emphasis id="B968EAAF4F3A1573FA8E7BABFECAFB96" italics="true" pageId="13" pageNumber="14">choueitangensis</emphasis>
|
|
(
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|
<bibRefCitation id="EF8D4B4C4F3A1573FED67B87FE78FB96" author="Mansuy" box="[272,435,1017,1043]" firstAuthor="Mansuy" pageId="13" pageNumber="14" pagination="1 - 146" refId="ref25972" refString="Mansuy, H. (1912) Paleontologie. In: Mansuy, H. & Deprat, J. (Eds.), Etude geologique du Yun-Nan oriental. Actes de la Societe Geologique de l'Indo-Chine, 1 (2), pp. 1 - 146." type="journal article" year="1912">Mansuy, 1912</bibRefCitation>
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|
, pl. 18, fig. 12) shows a similar whorl morphology and might be closely related.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |