544 lines
63 KiB
XML
544 lines
63 KiB
XML
<document id="A4152C16C6AA845EAC61DE8B57BCFDB0" ID-DOI="10.5070/P9401462457" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913330" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1712167746961" checkinUser="felipe" docAuthor="Wheeler, Elisabeth A., Manchester, Steven R. & Baas, Pieter" docDate="2023" docId="038AF505A31B960654ACFBF5FFA89819" docLanguage="en" docName="PaleoBios.40.14.1-55.pdf" docOrigin="PaleoBios 40 (14)" docSource="http://dx.doi.org/10.5070/p9401462457" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Klaassenoxylon WILKINSONII 2023, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="38" masterDocId="FFB38D7DA33F9621547BFFC0FFD49B4F" masterDocTitle="A late Eocene wood assemblage from the Crooked River Basin, Oregon, USA" masterLastPageNumber="55" masterPageNumber="1" pageNumber="35" updateTime="1728052592909" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:titleInfo id="B8045BFD2DC552A4ADA9D4CDFF552FF7">
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<mods:title id="023D7408F2103210989635C30924C098">A late Eocene wood assemblage from the Crooked River Basin, Oregon, USA</mods:title>
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<mods:name id="46D9C2FE9E6B4EFF026FCFD92EDCF01B" type="personal">
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<mods:roleTerm id="434ED1C89E4449E36A214A5B887A3FE2">Author</mods:roleTerm>
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<mods:namePart id="364354E76A6EFB03B4587AE86289A2FA">Wheeler, Elisabeth A.</mods:namePart>
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<mods:roleTerm id="788728068887CFB9029908A318E1DB4E">Author</mods:roleTerm>
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<mods:namePart id="5373E53CE19BEFC3464FD6ED8FE66011">Manchester, Steven R.</mods:namePart>
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<mods:name id="531252BFE982BA0EEB51E8FBAFCB6F73" type="personal">
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<mods:roleTerm id="59C6283B3FA82BC1A31D4BFBF5EC3751">Author</mods:roleTerm>
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<mods:namePart id="9EC2C082A3F71406542320C248BCB01C">Baas, Pieter</mods:namePart>
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<mods:typeOfResource id="75F0EF4139FAD24396967F2E98A8B822">text</mods:typeOfResource>
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<mods:title id="356CDE31E407857D35AECDE1A058B6E4">PaleoBios</mods:title>
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<mods:part id="2CBA94A6914316339EB45BA55E1FF183">
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<mods:date id="369B81A301F83209BEAF27A5AE45E64D">2023</mods:date>
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<mods:number id="F0A142955F2C488351DD571811E8B1D4">2023-11-01</mods:number>
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<mods:number id="267465AFAEA6A1B8E57088CDD475F288">40</mods:number>
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<mods:detail id="A7A6771DD800B1766F085C122BCE7B70" type="issue">
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<mods:number id="766F7D81A43FC6B0BA45548F6A5A9012">14</mods:number>
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<mods:start id="5FBF419732F3AC0B9F562C785ED5BF07">1</mods:start>
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<mods:url id="CBEF9CA54D62960B03891CEF424F5D51">http://dx.doi.org/10.5070/p9401462457</mods:url>
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<mods:classification id="F05AC6AE665C9DF068A5E7DBF32DEF20">journal article</mods:classification>
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<mods:identifier id="1FCB4571E5F78B934F877A4D1372D7B4" type="DOI">10.5070/P9401462457</mods:identifier>
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<mods:identifier id="86E35934534EF495BEC77581CF714A35" type="ISSN">0031-0298</mods:identifier>
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</mods:mods>
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<treatment id="038AF505A31B960654ACFBF5FFA89819" LSID="urn:lsid:plazi:treatment:038AF505A31B960654ACFBF5FFA89819" httpUri="http://treatment.plazi.org/id/038AF505A31B960654ACFBF5FFA89819" lastPageId="39" lastPageNumber="38" pageId="36" pageNumber="35">
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<subSubSection id="C3391798A31B960554ACFBF5FD6E9F1F" box="[215,698,1077,1105]" pageId="36" pageNumber="35" type="nomenclature">
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<paragraph id="8B9C4413A31B960554ACFBF5FD6E9F1F" blockId="36.[215,698,1077,1105]" box="[215,698,1077,1105]" pageId="36" pageNumber="35">
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<heading id="D0D4F37FA31B960554ACFBF5FD6E9F1F" allCaps="true" box="[215,698,1077,1105]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="35" reason="2">
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<taxonomicName id="4C233F90A31B960554ACFBF5FDB59F1E" authority="WILKINSONII" authorityName="WILKINSONII" authorityYear="2023" box="[215,609,1077,1105]" class="Magnoliopsida" family="Sapindaceae" genus="Klaassenoxylon" kingdom="Plantae" order="Sapindales" pageId="36" pageNumber="35" phylum="Tracheophyta" rank="genus" status="sp. nov.">
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<emphasis id="B9579801A31B960554ACFBF5FDB59F1E" box="[215,609,1077,1105]" italics="true" pageId="36" pageNumber="35">KLAASSENOXYLON WILKINSONII</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A264257AA31B9605561CFBF8FD6E9F1F" box="[615,698,1080,1104]" pageId="36" pageNumber="35" rank="species">
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<collectionCode id="ED32DCD6A31B9605561CFBF8FD559F1F" box="[615,641,1080,1104]" country="Brazil" name="Instituto de Botânica" pageId="36" pageNumber="35" type="Herbarium">SP</collectionCode>
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. NOV.
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A31B960555F0FBA2FDD19F35" box="[395,517,1122,1146]" pageId="36" pageNumber="35" type="description">
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<paragraph id="8B9C4413A31B960555F0FBA2FDD19F35" blockId="36.[395,517,1122,1146]" box="[395,517,1122,1146]" pageId="36" pageNumber="35">
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<figureCitation id="13185896A31B960555F0FBA2FDD19F35" box="[395,517,1122,1146]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">FIG. 17A–K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A31B960554EBFB56FE919E8A" pageId="36" pageNumber="35" type="diagnosis">
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<paragraph id="8B9C4413A31B960554EBFB56FE919E8A" blockId="36.[112,801,1174,1964]" pageId="36" pageNumber="35">
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<emphasis id="B9579801A31B960554EBFB56FEE49FFF" bold="true" box="[144,304,1174,1200]" pageId="36" pageNumber="35">Diagnosis—</emphasis>
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Growth rings distinct to indistinct. Wood diffuse-porous. Narrow vessels solitary and in short radial multiples. Perforation plates exclusively solitary; intervessel pits alternate, minute to small; vessel-ray parenchyma pits similar to intervessel pits. Fibers with simple pits, septate and non-septate. Axial parenchyma rare, scanty paratracheal. Rays uniseriate (rarely biseriate), heterocellular, less than
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<quantity id="4CDBE9F6A31B96055599FA49FDFF9EEC" box="[482,555,1417,1443]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="35" unit="mm" value="1.0">1 mm</quantity>
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tall. Long chains of crystals in fibers,
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A31B960554EBFA0EFDB89D1E" pageId="36" pageNumber="35" type="materials_examined">
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<paragraph id="8B9C4413A31B960554EBFA0EFEF69D44" blockId="36.[112,801,1174,1964]" pageId="36" pageNumber="35">
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<materialsCitation id="3B4B4E4EA31B960554EBFA0EFEF69D44" collectionCode="UF" pageId="36" pageNumber="35" specimenCode="UF 278-84877" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9579801A31B960554EBFA0EFEFD9EA7" bold="true" box="[144,297,1486,1512]" pageId="36" pageNumber="35">
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<typeStatus id="5498FAB1A31B960554EBFA0EFEDD9EA7" box="[144,265,1486,1512]" pageId="36" pageNumber="35" type="holotype">Holotype</typeStatus>
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—
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</emphasis>
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<specimenCode id="DB85EC68A31B96055553FA0FFE369EA7" box="[296,482,1486,1513]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF 278-84877</specimenCode>
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, estimated maximum diameter
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<quantity id="4CDBE9F6A31B960554ABFA31FECB9D44" box="[208,287,1521,1547]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="35" unit="cm" value="12.0">12 cm</quantity>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9C4413A31B960554EBF9D3FDB89D1E" blockId="36.[112,801,1174,1964]" pageId="36" pageNumber="35">
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<materialsCitation id="3B4B4E4EA31B960554EBF9D3FDBC9D1E" collectionCode="UF" pageId="36" pageNumber="35" specimenCode="UF 278-84882, 84897" specimenCount="1" typeStatus="paratype">
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<emphasis id="B9579801A31B960554EBF9D3FEE79D62" bold="true" box="[144,307,1555,1581]" pageId="36" pageNumber="35">
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<typeStatus id="5498FAB1A31B960554EBF9D3FEC09D62" box="[144,276,1555,1581]" pageId="36" pageNumber="35" type="paratype">Paratypes</typeStatus>
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—
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</emphasis>
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<specimenCode id="DB85EC68A31B9605554FF9D4FE3F9D62" box="[308,491,1555,1582]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF 278-84882</specimenCode>
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,
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<specimenCode id="DB85EC68A31B9605558CF9D3FD9D9D62" box="[503,585,1555,1581]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">84897</specimenCode>
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, estimated maximum diameters
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||
<quantity id="4CDBE9F6A31B96055547F9F6FE5A9D1E" box="[316,398,1590,1617]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="36" pageNumber="35" unit="cm" value="2.2">2.2 cm</quantity>
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(with pith),
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<quantity id="4CDBE9F6A31B96055651F9F6FDBC9D1E" box="[554,616,1590,1617]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="35" unit="cm" value="4.0">4 cm</quantity>
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</materialsCitation>
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||
.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A31B960554EBF999FDEC9D94" pageId="36" pageNumber="35" type="etymology">
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<paragraph id="8B9C4413A31B960554EBF999FDD59DF6" blockId="36.[112,801,1174,1964]" pageId="36" pageNumber="35">
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<emphasis id="B9579801A31B960554EBF999FE979D3C" bold="true" box="[144,323,1625,1651]" pageId="36" pageNumber="35">Etymology—</emphasis>
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Named for Hazel Wilkinson (1932- 2019), a devoted plant anatomist with a long career at the Jodrell Laboratory, Kew,
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<collectingCountry id="F3340483A31B960555B4F95FFDD59DF6" box="[463,513,1695,1721]" name="United Kingdom" pageId="36" pageNumber="35">U.K.</collectingCountry>
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</paragraph>
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<paragraph id="8B9C4413A31B960554EBF900FDEC9D94" blockId="36.[112,801,1174,1964]" box="[144,568,1728,1755]" pageId="36" pageNumber="35">
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<emphasis id="B9579801A31B960554EBF900FE939D95" bold="true" box="[144,327,1728,1754]" pageId="36" pageNumber="35">Occurrence—</emphasis>
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Dietz Hill (
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<collectionCode id="ED32DCD6A31B960555B1F901FE249D94" box="[458,496,1729,1755]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
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278).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3391798A31B960654EBF923FFA89819" lastPageId="39" lastPageNumber="38" pageId="36" pageNumber="35" type="description">
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<paragraph id="8B9C4413A31B960554EBF923FE219C6E" blockId="36.[112,801,1174,1964]" pageId="36" pageNumber="35">
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<emphasis id="B9579801A31B960554EBF923FE9F9DB2" bold="true" box="[144,331,1763,1789]" pageId="36" pageNumber="35">Description—</emphasis>
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Growth rings present, marked by radially narrow fibers (
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<figureCitation id="13185896A31B96055526F8C6FE339C6E" box="[349,487,1798,1825]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17A, B</figureCitation>
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).
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</paragraph>
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<paragraph id="8B9C4413A31B960554EBF8E9FBBC99DA" blockId="36.[112,801,1174,1964]" lastBlockId="36.[832,1521,424,1964]" pageId="36" pageNumber="35">
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Wood diffuse-porous; vessels solitary 29% in
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<collectionCode id="ED32DCD6A31B960556C5F8E9FD359C0C" box="[702,737,1833,1859]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
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278- 84877, 64% in
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<collectionCode id="ED32DCD6A31B96055556F88CFE849C29" box="[301,336,1868,1894]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
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278-84897 and in radial multiples of 2–3(-4), solitary vessels round to oval in outline (
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<figureCitation id="13185896A31B9605568AF8AEFF1C9CE4" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17A, B</figureCitation>
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); mean tangential diameters 74 (16) µm in
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<collectionCode id="ED32DCD6A31B96055686F851FCF49CE4" box="[765,800,1937,1963]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
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278-84877, 67 (12) µm in
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<collectionCode id="ED32DCD6A31B960550E7FE68FB6B9A8D" box="[1180,1215,424,450]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
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278-83897, total range of both specimens 45–106 µm; 18–23–33 vessels/ mm
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<superScript id="7C56E95BA31B9605570AFE2EFCA8994E" attach="left" box="[881,892,494,513]" fontSize="8" pageId="36" pageNumber="35">2</superScript>
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. Perforation plates exclusively simple (
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<figureCitation id="13185896A31B96055105FE2EFC809963" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17C, F</figureCitation>
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); intervessel pits alternate, minute to small (
|
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<figureCitation id="13185896A31B9605510CFDD1FA359964" box="[1399,1505,529,555]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17C</figureCitation>
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); vessel-ray parenchyma pits similar to intervessel pits (
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<figureCitation id="13185896A31B96055732FD97FC63993D" box="[841,951,599,626]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17D</figureCitation>
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). Average vessel element length 350 (
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<collectionCode id="ED32DCD6A31B960551EBFD98FA6C993D" box="[1424,1464,600,626]" country="USA" lsid="urn:lsid:biocol.org:col:15659" name="San Diego Natural History Museum" pageId="36" pageNumber="35" type="Herbarium">SD</collectionCode>
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=71) µm, range 249–452 µm.
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</paragraph>
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<paragraph id="8B9C4413A31B9605571BFD5EFAAA9994" blockId="36.[832,1521,424,1964]" pageId="36" pageNumber="35">Axial parenchyma rare, scanty paratracheal, in 3–4-celled strands as viewed in radial section.</paragraph>
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<paragraph id="8B9C4413A31B9605571BFD24FADA986D" blockId="36.[832,1521,424,1964]" pageId="36" pageNumber="35">
|
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Fibers thin- to thick-walled, pitting not observed, septate and non-septate (
|
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<figureCitation id="13185896A31B96055007FCC7FAD6986D" box="[1148,1282,775,802]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17F, G</figureCitation>
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).
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</paragraph>
|
||
<paragraph id="8B9C4413A31B9605571BFCEBFC3798BA" blockId="36.[832,1521,424,1964]" pageId="36" pageNumber="35">
|
||
Rays uniseriate (
|
||
<figureCitation id="13185896A31B96055054FCEBFB60980A" box="[1071,1204,811,837]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17F, G</figureCitation>
|
||
), with intermixed weakly procumbent to square cells to weakly upright cells; in others a tendency for square to upright cells to be restricted to ray margins; average ray height 346 (
|
||
<collectionCode id="ED32DCD6A31B960551FAFC54FA7D98E0" box="[1409,1449,916,943]" country="USA" lsid="urn:lsid:biocol.org:col:15659" name="San Diego Natural History Museum" pageId="36" pageNumber="35" type="Herbarium">SD</collectionCode>
|
||
=154) µm in
|
||
<collectionCode id="ED32DCD6A31B960557F7FC78FC61989D" box="[908,949,952,978]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
|
||
.84877; 284 (127) µm in
|
||
<collectionCode id="ED32DCD6A31B960550A4FC78FAD6989D" box="[1247,1282,952,978]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
|
||
278-84897; 10–14 rays per mm.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A31B9605571BFC3EFA1B9F57" blockId="36.[832,1521,424,1964]" box="[864,1487,1022,1048]" pageId="36" pageNumber="35">Prismatic crystals in chambered fibrous elements.</paragraph>
|
||
<paragraph id="8B9C4413A31B9605571BFBE1FB679DDD" blockId="36.[832,1521,424,1964]" pageId="36" pageNumber="35">
|
||
<emphasis id="B9579801A31B9605571BFBE1FBA39F74" bold="true" box="[864,1143,1057,1083]" pageId="36" pageNumber="35">Description of pith—</emphasis>
|
||
Pith complex, surrounded by a sheath of small cells with dark contents and clusters and strands of fiber-sclereids and sclereids (
|
||
<figureCitation id="13185896A31B9605513BFBA7FA719FCD" box="[1344,1445,1127,1154]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17I</figureCitation>
|
||
). Pith proper composed of
|
||
<specimenCount id="9D258F9AA31B9605503DFB4BFB169FEA" box="[1094,1218,1163,1189]" count="2" pageId="36" pageNumber="35" type="generic" typeStatus="types">two types</specimenCount>
|
||
of parenchyma cells: 1) large “empty” rectangular cells that are in radial rows towards the periphery and clusters of isodiametric cells in the center, and 2) small cells with dark contents (tannins?) forming a three-dimensional network throughout the pith and partly segmenting it by concave arches as seen in cross section (
|
||
<figureCitation id="13185896A31B96055029FA9EFB6A9E37" box="[1106,1214,1374,1400]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17G</figureCitation>
|
||
). There are isodiametric empty cells and dark cells resembling irregular “rosettes” as described by
|
||
<bibRefCitation id="EFB239E2A31B96055077FA64FABA9EF0" author="Poole, I. & H. P. Wilkinson" box="[1036,1390,1444,1471]" pageId="36" pageNumber="35" pagination="65 - 75" refId="ref38390" refString="Poole, I. and H. P. Wilkinson. 1992. Two sapindaceous woods from the London Clay (Eocene) of southeast England. Review of Palaeobotany and Palynology 75: 65 - 75. [https: // doi. org / 10.1016 / 0034 - 6667 (92) 90150 - F]." type="journal article" year="1992">
|
||
<collectingRegion id="49E78AF1A31B96055077FA64FB869EF1" box="[1036,1106,1444,1470]" country="United Kingdom" name="Poole" pageId="36" pageNumber="35">Poole</collectingRegion>
|
||
and Wilkinson (1992)
|
||
</bibRefCitation>
|
||
for
|
||
<taxonomicName id="4C233F90A31B960551D9FA65FBFF9EAB" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="36" pageNumber="35" phylum="Tracheophyta" rank="species" species="guioaoides">
|
||
<emphasis id="B9579801A31B960551D9FA65FBFF9EAB" italics="true" pageId="36" pageNumber="35">Sapindoxylon guioaoides</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFB239E2A31B9605504FFA07FA4E9EAD" author="Poole, I. & H. P. Wilkinson" box="[1076,1434,1479,1506]" pageId="36" pageNumber="35" pagination="65 - 75" refId="ref38390" refString="Poole, I. and H. P. Wilkinson. 1992. Two sapindaceous woods from the London Clay (Eocene) of southeast England. Review of Palaeobotany and Palynology 75: 65 - 75. [https: // doi. org / 10.1016 / 0034 - 6667 (92) 90150 - F]." type="journal article" year="1992">
|
||
<collectingRegion id="49E78AF1A31B9605504FFA07FBAE9EAE" box="[1076,1146,1479,1505]" country="United Kingdom" name="Poole" pageId="36" pageNumber="35">Poole</collectingRegion>
|
||
and Wilkinson (1992)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C233F90A31B960551D2FA08FC1B9D4A" authority="Cav. (1797)" authorityName="Cav." authorityYear="1797" class="Magnoliopsida" family="Sapindaceae" genus="Guioa" kingdom="Plantae" order="Sapindales" pageId="36" pageNumber="35" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31B960551D2FA08FA249EAB" box="[1449,1520,1480,1508]" italics="true" pageId="36" pageNumber="35">Guioa</emphasis>
|
||
Cav. (1797)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C233F90A31B9605506AFA2BFA939D4A" authority="Gaertn. (1788)" authorityName="Gaertn." authorityYear="1788" box="[1041,1351,1515,1543]" class="Magnoliopsida" family="Sapindaceae" genus="Alectryon" kingdom="Plantae" order="Sapindales" pageId="36" pageNumber="35" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31B9605506AFA2BFB529D48" box="[1041,1158,1515,1543]" italics="true" pageId="36" pageNumber="35">Alectryon</emphasis>
|
||
Gaertn. (1788)
|
||
</taxonomicName>
|
||
. However,
|
||
<collectionCode id="ED32DCD6A31B960551B6FA2BFA249D4A" box="[1485,1520,1515,1541]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="36" pageNumber="35" type="Museum">UF</collectionCode>
|
||
278-84882’s pith differs significantly in having a distinct sheath of small-cells including some sclereids bordering the protoxylem. Prismatic crystals present in peripheral pith parenchyma (
|
||
<figureCitation id="13185896A31B96055058F9B7FB729DDD" box="[1059,1190,1655,1682]" captionStart="Figure 17" captionStartId="37.[151,226,1916,1940]" captionTargetBox="[155,1486,194,1902]" captionTargetId="figure-16@37.[155,1486,194,1906]" captionTargetPageId="37" captionText="Figure 17. Caption on pg. 37." figureDoi="http://doi.org/10.5281/zenodo.10913364" httpUri="https://zenodo.org/record/10913364/files/figure.png" pageId="36" pageNumber="35">Fig. 17J, K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A31B9607571BF95AFEBB9900" blockId="36.[832,1521,424,1964]" lastBlockId="38.[112,801,389,1966]" lastPageId="38" lastPageNumber="37" pageId="36" pageNumber="35">
|
||
<emphasis id="B9579801A31B9605571BF95AFB269DFA" bold="true" box="[864,1266,1690,1717]" pageId="36" pageNumber="35">Comparison to extant woods—</emphasis>
|
||
We initially searched the InsideWood database for diffuse-porous wood (5p), vessels solitary and in short multiples and not arranged in an obvious pattern (6–11a), simple perforation plates (13r), minute to small alternate intervessel pitting (22p, 26a, 27a), vessel-ray parenchyma pits similar to intervessel pits (30p), vessel tangential diameter averaging less than 200 µm (43a), vessel frequency between 5–40 per mm
|
||
<superScript id="7C56E95BA319960754DAFE45FF789AD7" attach="left" box="[161,172,389,408]" fontSize="8" pageId="38" pageNumber="37">2</superScript>
|
||
(46a, 49a, 50a), fibers with simple pits (61p), axial parenchyma not obvious (77a, 79–86a), rays exclusively uniseriate (96p), storied structure absent (118a, 120a), and prismatic crystals present in fibers (143p). This first search returned two
|
||
<taxonomicName id="4C233F90A31996075515FDD2FDF99963" authority="J. St." authorityName="J. St." box="[366,557,530,556]" class="Magnoliopsida" family="Lythraceae" kingdom="Plantae" order="Myrtales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Lythraceae J. St.</taxonomicName>
|
||
-Hil. (1805) records and 37
|
||
<taxonomicName id="4C233F90A319960754B7FDF5FEBF9900" authorityName="JUSSIEU" authorityYear="1789" box="[204,363,565,591]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<subSection id="E2AC5DF8A31A960454ECF8BCFE069CDB" box="[151,466,1916,1941]" pageId="37" pageNumber="36" type="multiple">
|
||
<paragraph id="8B9C4413A31A960454ECF8BCFE069CDB" blockId="37.[151,466,1916,1941]" box="[151,466,1916,1941]" pageId="37" pageNumber="36">
|
||
<emphasis id="B9579801A31A960454ECF8BCFED19CDB" bold="true" box="[151,261,1916,1940]" pageId="37" pageNumber="36">Figure 17</emphasis>
|
||
. Caption on pg. 37.
|
||
</paragraph>
|
||
</subSection>
|
||
<caption id="DF5C149BA3199607540BFF01FAB99A11" ID-DOI="http://doi.org/10.5281/zenodo.13890623" ID-Zenodo-Dep="13890623" httpUri="https://zenodo.org/record/13890623/files/figure.png" pageId="38" pageNumber="37" startId="38.[112,187,193,217]" targetBox="[155,1486,194,1902]" targetPageId="37" targetType="figure">
|
||
<paragraph id="8B9C4413A3199607540BFF01FAB99A11" blockId="38.[112,1520,193,351]" pageId="38" pageNumber="37">
|
||
<emphasis id="B9579801A3199607540BFF01FF0B9B96" bold="true" box="[112,223,193,217]" pageId="38" pageNumber="37">Figure 17</emphasis>
|
||
.
|
||
<taxonomicName id="4C233F90A31996075492FF01FEA69B96" authorityName="JUSSIEU" authorityYear="1789" box="[233,370,193,217]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C233F90A31996075500FF02FD749B95" authority="Wheeler & Manchester & Baas, 2023" authorityName="Wheeler & Manchester & Baas" authorityYear="2023" box="[379,672,194,218]" class="Magnoliopsida" family="Sapindaceae" genus="Klaassenoxylon" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="species" species="wilkinsonae" status="sp. nov.">
|
||
<emphasis id="B9579801A31996075500FF02FD749B95" box="[379,672,194,218]" italics="true" pageId="38" pageNumber="37">Klaassenoxylon wilkinsonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A264257AA319960756DDFF01FD229B97" box="[678,758,193,216]" pageId="38" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B9579801A31996075681FF01FCF29B96" bold="true" box="[762,806,193,217]" pageId="38" pageNumber="37">A, B</emphasis>
|
||
. Diffuse-porous wood. Vessels solitary and in short radial multiples, UF 278-84877, TS.
|
||
<emphasis id="B9579801A319960755FFFF1BFE479BBC" bold="true" box="[388,403,219,243]" pageId="38" pageNumber="37">C</emphasis>
|
||
. Crowded alternate intervessel pitting, some coalescent apertures, UF 278-84877, TLS.
|
||
<emphasis id="B9579801A31996075144FF1CFA859BBB" bold="true" box="[1343,1361,220,244]" pageId="38" pageNumber="37">D</emphasis>
|
||
. Vessel-ray parenchyma pitting similar to intervessel pitting, procumbent ray cells, UF 278-84877, RLS.
|
||
<emphasis id="B9579801A31996075036FF36FB889A41" bold="true" box="[1101,1116,246,270]" pageId="38" pageNumber="37">E</emphasis>
|
||
. Rays with barely procumbent and square cells, UF 278-84897, RLS.
|
||
<emphasis id="B9579801A319960755AEFED1FE379A66" bold="true" box="[469,483,273,297]" pageId="38" pageNumber="37">F</emphasis>
|
||
. Uniseriate rays, crystals in chambered fibers (C), UF 278-84877, TLS.
|
||
<emphasis id="B9579801A319960750A2FED1FB3D9A66" bold="true" box="[1241,1257,273,297]" pageId="38" pageNumber="37">G</emphasis>
|
||
. Uniseriate rays, septate fibers (S= septae), TLS.
|
||
<emphasis id="B9579801A31996075516FEECFEAB9A0B" bold="true" box="[365,383,300,324]" pageId="38" pageNumber="37">H</emphasis>
|
||
. Chambered crystals in fibers, UF 278-84897, TLS.
|
||
<emphasis id="B9579801A319960757DFFEECFC799A0B" bold="true" box="[932,941,300,324]" pageId="38" pageNumber="37">I</emphasis>
|
||
. Complex pith, UF 278-84882, TS.
|
||
<emphasis id="B9579801A31996075164FEECFA909A0B" bold="true" box="[1311,1348,300,324]" pageId="38" pageNumber="37">J, K</emphasis>
|
||
. Longitudinal sections ofpith, UF 278-84882. Scale bars=500 µm in A, I; 200 µm in B, J; 100 µm in E. F, K; 50 µm in G; 20 µm in C, D, H.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9C4413A319960754EBFD98FEC799F7" blockId="38.[112,801,389,1966]" pageId="38" pageNumber="37">
|
||
Search 2 included presence of both septate and non-septate fibers and yielded one
|
||
<taxonomicName id="4C233F90A3199607564EFDBBFD6A99DA" box="[565,702,635,661]" class="Magnoliopsida" family="Lythraceae" kingdom="Plantae" order="Myrtales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Lythraceae</taxonomicName>
|
||
and 32
|
||
<taxonomicName id="4C233F90A3199607540BFD5EFEDB99F7" authorityName="JUSSIEU" authorityYear="1789" box="[112,271,670,696]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A319960754EBFD02FD03989D" blockId="38.[112,801,389,1966]" pageId="38" pageNumber="37">
|
||
Search 3 added presence of rays with procumbent, square, and upright cells intermixed throughout the ray (109p) and absence of dimorphic fibers (67a) this narrowed the possibilities to seven sapindaceous genera, one of which,
|
||
<taxonomicName id="4C233F90A3199607556DFC8DFD249826" authority="Planch. ex Benth. (1849)" authorityName="Benth." authorityYear="1849" box="[278,752,845,873]" class="Magnoliopsida" family="Sapindaceae" genus="Lecaniodiscus" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A3199607556DFC8DFE6B9826" box="[278,447,845,873]" italics="true" pageId="38" pageNumber="37">Lecaniodiscus</emphasis>
|
||
Planch. ex Benth. (1849)
|
||
</taxonomicName>
|
||
has crystals in ray cells as a constant and so does not match the Dietz Hill (
|
||
<collectionCode id="ED32DCD6A31996075563FC55FEEA98E0" box="[280,318,917,943]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="38" pageNumber="37" type="Museum">UF</collectionCode>
|
||
278) woods. Consequently, we consider that these woods are allied with the
|
||
<taxonomicName id="4C233F90A3199607564FFC78FD07989D" authorityName="JUSSIEU" authorityYear="1789" box="[564,723,952,978]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A319960754EBFC1BFA259963" blockId="38.[112,801,389,1966]" lastBlockId="38.[833,1522,387,1966]" pageId="38" pageNumber="37">
|
||
<bibRefCitation id="EFB239E2A319960754EBFC1BFEB798BA" author="Klaassen, R." box="[144,355,987,1013]" pageId="38" pageNumber="37" pagination="1 - 214" refId="ref36287" refString="Klaassen, R. 1999. Wood anatomy of the Sapindaceae. IAWA Journal Supplement 2: 1 - 214." type="journal article" year="1999">Klaassen (1999</bibRefCitation>
|
||
: p. 174–175) surveyed the wood anatomy of the
|
||
<taxonomicName id="4C233F90A3199607554FFC3EFE049F57" authorityName="JUSSIEU" authorityYear="1789" box="[308,464,1022,1048]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
and recognized four wood anatomical
|
||
<typeStatus id="5498FAB1A31996075485FBE2FE909F73" box="[254,324,1058,1084]" pageId="38" pageNumber="37">types</typeStatus>
|
||
. These Dietz Hill (
|
||
<collectionCode id="ED32DCD6A31996075661FBE2FD949F73" box="[538,576,1058,1084]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="38" pageNumber="37" type="Museum">UF</collectionCode>
|
||
278) woods fit his
|
||
<typeStatus id="5498FAB1A3199607540BFB85FF7A9F10" box="[112,174,1093,1119]" pageId="38" pageNumber="37">Type</typeStatus>
|
||
IVa, which includes 26 genera that have intervessel pits of 3–6 µm, septate fibers, scanty paratracheal parenchyma, heterocellular uniseriate rays, and crystals in fibrous elements.
|
||
<taxonomicName id="4C233F90A31996075514FB6FFD349F86" authority="Adema (1993)" authorityName="Adema" authorityYear="1993" box="[367,736,1198,1225]" class="Magnoliopsida" family="Sapindaceae" genus="Cnesmocarpon" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075514FB6FFDF79F86" box="[367,547,1199,1225]" italics="true" pageId="38" pageNumber="37">Cnesmocarpon</emphasis>
|
||
Adema (1993)
|
||
</taxonomicName>
|
||
New
|
||
<collectingCountry id="F3340483A3199607540BFB12FF1D9FA3" box="[112,201,1234,1260]" name="Guinea-Bissau" pageId="38" pageNumber="37">Guinea</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3340483A319960754A8FB12FE939FA2" box="[211,327,1234,1261]" name="Australia" pageId="38" pageNumber="37">Australia</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C233F90A31996075521FB12FD5D9FA2" authority="Franch. (1886)" authorityName="Franch." authorityYear="1886" box="[346,649,1234,1262]" class="Magnoliopsida" family="Sapindaceae" genus="Delavaya" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075521FB12FE1F9FA1" box="[346,459,1234,1262]" italics="true" pageId="38" pageNumber="37">Delavaya</emphasis>
|
||
Franch. (1886)
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F3340483A319960756ECFB12FD309FA2" box="[663,740,1234,1261]" name="China" pageId="38" pageNumber="37">China</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C233F90A31996075683FB12FE3D9E40" authority="Buerki and Callm. (2010)" authorityName="Buerki and Callm." authorityYear="2010" class="Magnoliopsida" family="Sapindaceae" genus="Gereaua" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075683FB12FF629E40" italics="true" pageId="38" pageNumber="37">Gereaua</emphasis>
|
||
Buerki and Callm. (2010)
|
||
</taxonomicName>
|
||
(syn.
|
||
<taxonomicName id="4C233F90A31996075657FB35FF659E7D" authority="Radlk., 1878" authorityName="Radlk." authorityYear="1878" class="Magnoliopsida" family="Sapindaceae" genus="Haplocoelum" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075657FB35FD1E9E40" box="[556,714,1269,1295]" italics="true" pageId="38" pageNumber="37">Haplocoelum</emphasis>
|
||
Radlk., 1878
|
||
</taxonomicName>
|
||
; Africa),
|
||
<taxonomicName id="4C233F90A31996075558FAD7FD5C9E7D" authority="Blume (1848)" authorityName="Blume" authorityYear="1848" box="[291,648,1303,1331]" class="Magnoliopsida" family="Sapindaceae" genus="Macphersonia" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075558FAD7FE049E7C" box="[291,464,1303,1331]" italics="true" pageId="38" pageNumber="37">Macphersonia</emphasis>
|
||
Blume (1848)
|
||
</taxonomicName>
|
||
(Tropical
|
||
<collectionCode id="ED32DCD6A31996075772FAD9FCC89E7C" box="[777,796,1305,1331]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="38" pageNumber="37" type="Herbarium">E</collectionCode>
|
||
. Africa,
|
||
<collectingCountry id="F3340483A319960754B1FAFBFEB09E19" box="[202,356,1339,1366]" name="Madagascar" pageId="38" pageNumber="37">Madagascar</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C233F90A31996075502FAFAFD699E1A" authority="Baill. (1874)" authorityName="Baill." authorityYear="1874" box="[377,701,1338,1366]" class="Magnoliopsida" family="Sapindaceae" genus="Pseudopteris" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075502FAFAFDC29E19" box="[377,534,1338,1366]" italics="true" pageId="38" pageNumber="37">Pseudopteris</emphasis>
|
||
Baill. (1874)
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F3340483A319960756B5FAFBFF179E36" name="Madagascar" pageId="38" pageNumber="37">Madagascar</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C233F90A319960754A3FA9FFE429E37" authority="Endl." authorityName="Endl." box="[216,406,1374,1403]" class="Magnoliopsida" family="Sapindaceae" genus="Ungnadia" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960754A3FA9FFE9B9E34" box="[216,335,1375,1403]" italics="true" pageId="38" pageNumber="37">Ungnadia</emphasis>
|
||
Endl.
|
||
</taxonomicName>
|
||
(1833[1835];
|
||
<collectingCountry id="F3340483A31996075637FA9EFD729E37" box="[588,678,1374,1400]" name="Mexico" pageId="38" pageNumber="37">Mexico</collectingCountry>
|
||
, sw
|
||
<collectingCountry id="F3340483A319960756A0FA9FFCC19E36" box="[731,789,1375,1401]" name="United States of America" pageId="38" pageNumber="37">USA</collectingCountry>
|
||
). Klaassen’s synoptical key with twenty-six features is slightly awkward to use. We excluded genera that have as constant features ones not observed in these fossils (vessels in a radial pattern, mean intervessel pit diameter>7 µm, mean vessel diameter>120 µm, fiber dimorphism present, apotracheal parenchyma abundant, complete sheaths of vasicentric parenchyma, aliform to confluent parenchyma, banded non-marginal parenchyma, idioblastic axial cells, bi- and tri-seriate rays common, mean ray height>
|
||
<quantity id="4CDBE9F6A31996075526F97FFE709D96" box="[349,420,1727,1753]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="37" unit="mm" value="1.0">1 mm</quantity>
|
||
, rays all homocellular, rays of two distinct sizes, crystals in ray cells). In addition to the six genera listed above as sharing features with
|
||
<collectionCode id="ED32DCD6A31996075686F8C5FCF49C50" box="[765,800,1797,1823]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="38" pageNumber="37" type="Museum">UF</collectionCode>
|
||
278-84887, Klaassen’s key indicates the following genera also have similarity:
|
||
<taxonomicName id="4C233F90A319960755D8F88CFD6C9C2A" authority="Blume (1847)" authorityName="Blume" authorityYear="1847" box="[419,696,1867,1896]" class="Magnoliopsida" family="Sapindaceae" genus="Arytera" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960755D8F88CFDD49C27" box="[419,512,1868,1896]" italics="true" pageId="38" pageNumber="37">Arytera</emphasis>
|
||
Blume (1847)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A319960756BEF88CFEEE9CC6" authority="Capuron (1969)" authorityName="Capuron" authorityYear="1969" class="Magnoliopsida" family="Sapindaceae" genus="Beguea" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960756BEF88CFCF49C27" box="[709,800,1868,1896]" italics="true" pageId="38" pageNumber="37">Beguea</emphasis>
|
||
Capuron (1969)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A31996075533F8AFFE799CC4" box="[328,429,1903,1931]" class="Magnoliopsida" family="Sapindaceae" genus="Cupania" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075533F8AFFE799CC4" box="[328,429,1903,1931]" italics="true" pageId="38" pageNumber="37">Cupania</emphasis>
|
||
</taxonomicName>
|
||
<collectionCode id="ED32DCD6A319960755CEF8AFFE139CC6" box="[437,455,1903,1929]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="38" pageNumber="37" type="Herbarium">L</collectionCode>
|
||
. (1753),
|
||
<taxonomicName id="4C233F90A31996075643F8AFFCF49CC7" authority="Radlk." authorityName="Radlk." box="[568,800,1902,1931]" class="Magnoliopsida" family="Sapindaceae" genus="Cupaniopsis" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075643F8AFFD1E9CC4" box="[568,714,1903,1931]" italics="true" pageId="38" pageNumber="37">Cupaniopsis</emphasis>
|
||
Radlk.
|
||
</taxonomicName>
|
||
(1879/1880),
|
||
<taxonomicName id="4C233F90A3199607555BF852FC599AEF" authority="Hook. f. (Bentham and Hooker, 1862)" authorityName="Hook. f. (Bentham and Hooker" authorityYear="1862" box="[288,909,389,1966]" class="Magnoliopsida" family="Sapindaceae" genus="Diploglottis" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A3199607555BF852FE799CE1" box="[288,429,1938,1966]" italics="true" pageId="38" pageNumber="37">Diploglottis</emphasis>
|
||
Hook.f. (Bentham and Hooker, 1862)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A319960757E7FE43FB1B9AEF" authority="Capuron (1969)" authorityName="Capuron" authorityYear="1969" box="[924,1231,387,416]" class="Magnoliopsida" family="Sapindaceae" genus="Neotina" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960757E7FE43FC299AD0" box="[924,1021,387,415]" italics="true" pageId="38" pageNumber="37">Neotina</emphasis>
|
||
Capuron (1969)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A319960750A6FE43FA3F9AD0" authority="Poit. (1804)" authorityName="Poit." authorityYear="1804" box="[1245,1515,387,415]" class="Magnoliopsida" family="Sapindaceae" genus="Thouinia" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960750A6FE43FA9F9AD0" box="[1245,1355,387,415]" italics="true" pageId="38" pageNumber="37">Thouinia</emphasis>
|
||
Poit. (1804)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C233F90A3199607570FFE68FBEE9A8D" authority="Radlk." authorityName="Radlk." box="[884,1082,424,450]" class="Magnoliopsida" family="Sapindaceae" genus="Toechima" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A3199607570FFE68FC3C9A8D" box="[884,1000,424,450]" italics="true" pageId="38" pageNumber="37">Toechima</emphasis>
|
||
Radlk.
|
||
</taxonomicName>
|
||
(1877[1879]) of
|
||
<typeStatus id="5498FAB1A31996075172FE69FA929A8C" box="[1289,1350,425,451]" pageId="38" pageNumber="37">Type</typeStatus>
|
||
IVa;
|
||
<taxonomicName id="4C233F90A31996075105FE68FC239AA9" authority="Gaertn. (1788)" authorityName="Gaertn." authorityYear="1788" class="Magnoliopsida" family="Sapindaceae" genus="Alectryon" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075105FE68FA269A8B" box="[1406,1522,424,452]" italics="true" pageId="38" pageNumber="37">Alectryon</emphasis>
|
||
Gaertn. (1788)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A31996075079FE0CFADA9AA9" authority="Aubl. (1775)" authorityName="Aubl." authorityYear="1775" box="[1026,1294,459,488]" class="Magnoliopsida" family="Sapindaceae" genus="Matayba" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075079FE0CFBB99AA7" box="[1026,1133,460,488]" italics="true" pageId="38" pageNumber="37">Matayba</emphasis>
|
||
Aubl. (1775)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A31996075163FE0CFC439946" authority="Willd. (1805)" authorityName="Willd." authorityYear="1805" class="Magnoliopsida" family="Sapindaceae" genus="Schleichera" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075163FE0CFA779AA7" box="[1304,1443,460,488]" italics="true" pageId="38" pageNumber="37">Schleichera</emphasis>
|
||
Willd. (1805)
|
||
</taxonomicName>
|
||
of
|
||
<typeStatus id="5498FAB1A319960757C3FE2FFC219946" box="[952,1013,495,521]" pageId="38" pageNumber="37">Type</typeStatus>
|
||
IVb; and
|
||
<taxonomicName id="4C233F90A31996075018FE2DFA3F9946" authority="Rehder (1935)" authorityName="Rehder" authorityYear="1935" box="[1123,1515,493,521]" class="Magnoliopsida" family="Sapindaceae" genus="Handeliodendron" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31996075018FE2DFAE79946" box="[1123,1331,493,521]" italics="true" pageId="38" pageNumber="37">Handeliodendron</emphasis>
|
||
Rehder (1935)
|
||
</taxonomicName>
|
||
, which did not fit in any of Klaassen’s wood
|
||
<typeStatus id="5498FAB1A3199607512CFDD2FA5A9963" box="[1367,1422,530,556]" pageId="38" pageNumber="37">type</typeStatus>
|
||
groups.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A3199607571AFDF5FAA19809" blockId="38.[833,1522,387,1966]" pageId="38" pageNumber="37">
|
||
Buerki et al.’s (2021) recent analysis of
|
||
<taxonomicName id="4C233F90A3199607512EFDF5FA259900" authorityName="JUSSIEU" authorityYear="1789" box="[1365,1521,565,591]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
recognized four subfamilies and twenty tribes. All six genera from InsideWood Search 3 belong to subfamily
|
||
<taxonomicName id="4C233F90A31996075713FD5EFB0E99F6" authority="Burnett (1835)" authorityName="Burnett" authorityYear="1835" box="[872,1242,670,697]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="subFamily" subFamily="Sapindoideae">Sapindoideae Burnett (1835)</taxonomicName>
|
||
, but to three different tribes:
|
||
<taxonomicName id="4C233F90A319960757E3FD02FBF89993" box="[920,1068,706,732]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="tribe" tribe="Ungnadieae">Ungnadieae</taxonomicName>
|
||
Buerki and Callm. (2021) (
|
||
<emphasis id="B9579801A31996075100FD02FC6C984E" italics="true" pageId="38" pageNumber="37">
|
||
<taxonomicName id="4C233F90A31996075100FD02FA399991" box="[1403,1517,706,734]" class="Magnoliopsida" family="Sapindaceae" genus="Delavaya" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">Delavaya</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A3199607573AFD25FC6C984E" box="[833,952,741,769]" class="Magnoliopsida" family="Sapindaceae" genus="Ungnadia" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">Ungnadia</taxonomicName>
|
||
</emphasis>
|
||
), Stadmanieae Buerki and Callm. (2021) (
|
||
<taxonomicName id="4C233F90A319960751B2FD25FB15986D" authority="Buerki and Callm. (2010)" authorityName="Buerki and Callm." authorityYear="2010" class="Magnoliopsida" family="Sapindaceae" genus="Gereaua" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960751B2FD25FC5C986C" italics="true" pageId="38" pageNumber="37">Gereaua</emphasis>
|
||
Buerki and Callm. (2010)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9579801A319960750B7FCC7FCA39809" italics="true" pageId="38" pageNumber="37">
|
||
<taxonomicName id="4C233F90A319960750B7FCC7FAAD986C" authorityName="Blume" authorityYear="1848" box="[1228,1401,775,803]" class="Magnoliopsida" family="Sapindaceae" genus="Macphersonia" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">Macphersonia</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C233F90A319960751F9FCC7FCA39809" class="Magnoliopsida" family="Sapindaceae" genus="Pseudopteris" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">Pseudopteris</taxonomicName>
|
||
</emphasis>
|
||
), and Cupanieae Blume (
|
||
<taxonomicName id="4C233F90A319960750CBFCECFAB09809" authorityName="Adema" authorityYear="1993" box="[1200,1380,812,838]" class="Magnoliopsida" family="Sapindaceae" genus="Cnesmocarpon" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960750CBFCECFAB09809" box="[1200,1380,812,838]" italics="true" pageId="38" pageNumber="37">Cnesmocarpon</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A3199607571AFC8FFAB39E40" blockId="38.[833,1522,387,1966]" pageId="38" pageNumber="37">
|
||
It is tempting to suggest, based on present day geographic distribution as well as shared anatomical features, that these Dietz Hill sapindaceous woods might be related to tribe
|
||
<taxonomicName id="4C233F90A31996075079FC78FB4C989D" box="[1026,1176,952,978]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="tribe" tribe="Ungnadieae">Ungnadieae</taxonomicName>
|
||
. However, as pointed out by
|
||
<bibRefCitation id="EFB239E2A3199607573AFC1BFBD998B9" author="Klaassen, R." box="[833,1037,987,1014]" pageId="38" pageNumber="37" pagination="1 - 214" refId="ref36287" refString="Klaassen, R. 1999. Wood anatomy of the Sapindaceae. IAWA Journal Supplement 2: 1 - 214." type="journal article" year="1999">Klaassen (1999)</bibRefCitation>
|
||
, wood anatomical differences between the genera of groups IVa and IVb are small and could easily break down upon examination of more species per genus and more specimens per species. The main conclusion of the searches reported above and assorted alternative searches we did consistently yielded multiple
|
||
<taxonomicName id="4C233F90A3199607573AFB6EFC349F87" authorityName="JUSSIEU" authorityYear="1789" box="[833,992,1198,1224]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
. We consider the similarity in the complex pith anatomy with some modern and fossil
|
||
<taxonomicName id="4C233F90A3199607512EFB12FA259FA3" authorityName="JUSSIEU" authorityYear="1789" box="[1365,1521,1234,1260]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
|
||
as strong support for the family assignment.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A3199607571AFAD8FBCA9DB3" blockId="38.[833,1522,387,1966]" pageId="38" pageNumber="37">
|
||
<emphasis id="B9579801A3199607571AFAD8FB349E7D" bold="true" box="[865,1248,1304,1330]" pageId="38" pageNumber="37">Comparison to fossil woods—</emphasis>
|
||
The diagnosis of
|
||
<taxonomicName id="4C233F90A319960751DEFAD8FC749E17" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A319960751DEFAD8FC749E17" italics="true" pageId="38" pageNumber="37">Sapindoxylon</emphasis>
|
||
</taxonomicName>
|
||
as adapted by
|
||
<bibRefCitation id="EFB239E2A3199607502FFAFBFA7B9E19" author="Poole, I. & H. P. Wilkinson" box="[1108,1455,1339,1366]" pageId="38" pageNumber="37" pagination="65 - 75" refId="ref38390" refString="Poole, I. and H. P. Wilkinson. 1992. Two sapindaceous woods from the London Clay (Eocene) of southeast England. Review of Palaeobotany and Palynology 75: 65 - 75. [https: // doi. org / 10.1016 / 0034 - 6667 (92) 90150 - F]." type="journal article" year="1992">
|
||
<collectingRegion id="49E78AF1A3199607502FFAFBFB4E9E1A" box="[1108,1178,1339,1365]" country="United Kingdom" name="Poole" pageId="38" pageNumber="37">Poole</collectingRegion>
|
||
and Wilkinson (1992)
|
||
</bibRefCitation>
|
||
from a translation of Kraüsel’s work (1922) follows: “
|
||
<emphasis id="B9579801A319960751DFFA9DFA3E9E36" box="[1444,1514,1373,1401]" italics="true" pageId="38" pageNumber="37">Wood</emphasis>
|
||
: Diffuse porous.
|
||
<emphasis id="B9579801A31996075070FA42FB479ED3" box="[1035,1171,1408,1438]" italics="true" pageId="38" pageNumber="37">Growth rin</emphasis>
|
||
gs: lacking.
|
||
<emphasis id="B9579801A31996075152FA40FAA89ED3" box="[1321,1404,1408,1436]" italics="true" pageId="38" pageNumber="37">Vessels</emphasis>
|
||
: 2–8 per mm
|
||
<superScript id="7C56E95BA31996075709FA65FCA99EF7" attach="left" box="[882,893,1445,1464]" fontSize="8" pageId="38" pageNumber="37">2</superScript>
|
||
, mostly solitary or paired, seldom in threes, when abutting tangentially flattened, otherwise circular or elliptical. Perforation plates simple with mainly horizontal or slightly oblique end walls.
|
||
<emphasis id="B9579801A319960750D4F9CDFB2E9D66" box="[1199,1274,1549,1577]" italics="true" pageId="38" pageNumber="37">Fibres</emphasis>
|
||
: Libriform, forming the groundmass in regularly arranged rows; occasionally septate.
|
||
<emphasis id="B9579801A319960757D0F995FB4C9D20" box="[939,1176,1621,1647]" italics="true" pageId="38" pageNumber="37">Wood parenchyma:</emphasis>
|
||
Surrounding the vessels in one or two layers [vasicentric], sometimes also scattered.
|
||
<emphasis id="B9579801A3199607573AF95CFC569DF7" box="[833,898,1692,1720]" italics="true" pageId="38" pageNumber="37">Rays:</emphasis>
|
||
Uniseriate, occasionally biseriate, 3–30 cells high, consisting of one cell
|
||
<typeStatus id="5498FAB1A3199607503BF97FFBAD9D96" box="[1088,1145,1727,1753]" pageId="38" pageNumber="37">type</typeStatus>
|
||
; the marginal cells occasionally contain a crystal.”
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A3199606571AF8C5FEDB99E9" blockId="38.[833,1522,387,1966]" lastBlockId="39.[112,800,193,854]" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">
|
||
It is unfortunate that the vessel frequency was so narrowly defined. Some
|
||
<taxonomicName id="4C233F90A31996075042F8E8FAE19C0D" box="[1081,1333,1832,1860]" pageId="38" pageNumber="37">
|
||
<emphasis id="B9579801A31996075042F8E8FB0C9C0B" box="[1081,1240,1832,1860]" italics="true" pageId="38" pageNumber="37">Sapindoxylon</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
do not conform to the diagnosis in that feature, e.g.,
|
||
<taxonomicName id="4C233F90A3199607517CF88CFC5B9CC6" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="species" species="guioaoides">
|
||
<emphasis id="B9579801A3199607517CF88CFC5B9CC6" italics="true" pageId="38" pageNumber="37">Sapindoxylon guioaoides</emphasis>
|
||
</taxonomicName>
|
||
(average of 40 vessels mm
|
||
<superScript id="7C56E95BA3199607509BF8AEFB3F9CCE" attach="left" box="[1248,1259,1902,1921]" fontSize="8" pageId="38" pageNumber="37">2</superScript>
|
||
),
|
||
<taxonomicName id="4C233F90A31996075178F8AFFB519CE2" authority="Grambast-Fessard (1966)" authorityName="Grambast-Fessard" authorityYear="1966" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="species" species="elattostachyoides">
|
||
<emphasis id="B9579801A31996075178F8AFFA269CC4" box="[1283,1522,1903,1931]" italics="true" pageId="38" pageNumber="37">S. elattostachyoides</emphasis>
|
||
<bibRefCitation id="EFB239E2A3199607573AF852FB519CE2" author="Grambast-Fessard, N." box="[833,1157,1938,1965]" pageId="38" pageNumber="37" pagination="130 - 146" refId="ref35193" refString="Grambast-Fessard, N. 1966. Contribution a l'etude des flores tertiares des regiones provencales et alpines: deux bois nouveaux de dicotyledons de Pontien de Castellane. Memoires de la Societe geologique de France 105: 130 - 146, [+ pl. XI-XIV]" type="journal article" year="1966">Grambast-Fessard (1966)</bibRefCitation>
|
||
</taxonomicName>
|
||
, (20–40 vessels per mm
|
||
<superScript id="7C56E95BA319960751C2F851FA109CEB" attach="left" box="[1465,1476,1937,1956]" fontSize="8" pageId="38" pageNumber="37">2</superScript>
|
||
),
|
||
<taxonomicName id="4C233F90A319960651A6F852FEF99B92" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" lastPageId="39" lastPageNumber="38" order="Sapindales" pageId="38" pageNumber="37" phylum="Tracheophyta" rank="species" species="koelreuteroides">
|
||
<emphasis id="B9579801A319960651A6F852FEF99B92" italics="true" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">S. koelreuteroides</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFB239E2A31896065549FF02FD589B92" author="Poole, I. & H. P. Wilkinson" box="[306,652,194,221]" pageId="39" pageNumber="38" pagination="65 - 75" refId="ref38390" refString="Poole, I. and H. P. Wilkinson. 1992. Two sapindaceous woods from the London Clay (Eocene) of southeast England. Review of Palaeobotany and Palynology 75: 65 - 75. [https: // doi. org / 10.1016 / 0034 - 6667 (92) 90150 - F]." type="journal article" year="1992">
|
||
<collectingRegion id="49E78AF1A31896065549FF02FEAC9B93" box="[306,376,194,220]" country="United Kingdom" name="Poole" pageId="39" pageNumber="38">Poole</collectingRegion>
|
||
and Wilkinson (1992)
|
||
</bibRefCitation>
|
||
(average of 43 vessels mm
|
||
<superScript id="7C56E95BA31896065559FF25FEF99BB7" attach="left" box="[290,301,229,248]" fontSize="8" pageId="39" pageNumber="38">2</superScript>
|
||
). These Dietz Hill (
|
||
<collectionCode id="ED32DCD6A3189606566BFF26FDE29A4F" box="[528,566,230,256]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="39" pageNumber="38" type="Museum">UF</collectionCode>
|
||
278) samples have double the number of vessels per mm
|
||
<superScript id="7C56E95BA31896065631FEC8FD819A54" attach="left" box="[586,597,264,283]" fontSize="8" pageId="39" pageNumber="38">2</superScript>
|
||
of
|
||
<taxonomicName id="4C233F90A31896065606FEC9FEC39A09" authority="Krausel. They" authorityName="Krausel. They" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="38" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A31896065606FEC9FCF49A6A" box="[637,800,265,293]" italics="true" pageId="39" pageNumber="38">Sapindoxylon</emphasis>
|
||
Kraüsel. They
|
||
</taxonomicName>
|
||
also differ in 1) not having vasicentric axial parenchyma, but just scanty paratracheal; 2) heterocellular rays, rather than homocellular; 3) crystals in fibers/ chambered axial parenchyma, rather than in marginal ray parenchyma cells. These three features were used in
|
||
<bibRefCitation id="EFB239E2A3189606540BFE1CFE9B9AB9" author="Klaassen, R." box="[112,335,476,502]" pageId="39" pageNumber="38" pagination="1 - 214" refId="ref36287" refString="Klaassen, R. 1999. Wood anatomy of the Sapindaceae. IAWA Journal Supplement 2: 1 - 214." type="journal article" year="1999">Klaassen’s (1999)</bibRefCitation>
|
||
synoptic key and so were considered important in recognizing genera.
|
||
<taxonomicName id="4C233F90A3189606565FFE3FFD9F9972" authority="Wheeler and Manchester (2002)" authorityName="Wheeler" authorityYear="2002" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="38" phylum="Tracheophyta" rank="species" species="klaassenii">
|
||
<emphasis id="B9579801A3189606565FFE3FFF7C9972" italics="true" pageId="39" pageNumber="38">Sapindoxylon klaassenii</emphasis>
|
||
<bibRefCitation id="EFB239E2A318960654CBFDE2FD9F9972" author="Wheeler, E. A. & S. R. Manchester" box="[176,587,546,573]" pageId="39" pageNumber="38" pagination="1 - 188" refId="ref41110" refString="Wheeler, E. A. and S. R. Manchester. 2002. Woods of the Eocene Nut Beds flora, Clarno Formation, Oregon, USA. IAWA Journal Supplement 3: 1 - 188." type="journal article" year="2002">Wheeler and Manchester (2002)</bibRefCitation>
|
||
</taxonomicName>
|
||
from the middle Eocene Clarno Nut Beds differs from the
|
||
<bibRefCitation id="EFB239E2A3189606562EFD85FCF4992F" author="Krausel, R." box="[597,800,581,608]" pageId="39" pageNumber="38" pagination="231 - 287" refId="ref36362" refString="Krausel, R. 1922. Fossile holzer aus dem Tertiar von sud-Sumatra. Verhandlungen Geologischen Mijnbouwkd. Genoot. Nederland Kolon. (Geol. Ser.) 5: 231 - 287." type="journal article" year="1922">Kraüsel’s (1922)</bibRefCitation>
|
||
diagnosis in having rare axial parenchyma and heterocellular rays.
|
||
</paragraph>
|
||
<paragraph id="8B9C4413A318960654EBFD6FFFA89819" blockId="39.[112,800,193,854]" pageId="39" pageNumber="38">
|
||
Consequently, we choose not to assign
|
||
<collectionCode id="ED32DCD6A31896065617FD70FD5B9985" box="[620,655,688,714]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="39" pageNumber="38" type="Museum">UF</collectionCode>
|
||
278-84887 to
|
||
<taxonomicName id="4C233F90A318960654E9FD13FE4A99A3" authority="Krausel" authorityName="Krausel" box="[146,414,722,751]" class="Magnoliopsida" family="Sapindaceae" genus="Sapindoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="38" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A318960654E9FD13FEE199A0" box="[146,309,723,751]" italics="true" pageId="39" pageNumber="38">Sapindoxylon</emphasis>
|
||
Kraüsel
|
||
</taxonomicName>
|
||
and propose the genus
|
||
<taxonomicName id="4C233F90A318960656A8FD11FF3C985D" authorityName="WILKINSONII" authorityYear="2023" class="Magnoliopsida" family="Sapindaceae" genus="Klaassenoxylon" kingdom="Plantae" order="Sapindales" pageId="39" pageNumber="38" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B9579801A318960656A8FD11FF3C985D" italics="true" pageId="39" pageNumber="38">Klaassenoxylon</emphasis>
|
||
</taxonomicName>
|
||
to accommodate these Eocene sapindaceous woods from
|
||
<collectingRegion id="49E78AF1A31896065571FCD9FEBC987C" box="[266,360,793,819]" country="United States of America" name="Oregon" pageId="39" pageNumber="38">Oregon</collectingRegion>
|
||
. We also propose the new combination:
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |