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<document id="0609D82362C0A0C94D91FB69F117F958" ID-DOI="10.11646/zootaxa.5496.2.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="13346723" ID-ZooBank="970337FC-03E1-4468-A604-5AC14590F7B0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1724133169350" checkinUser="plazi" docAuthor="Na, Sol-Moon &amp; Bae, Yang-Seop" docDate="2024" docId="736887809902627F5BCF7CE64CB0200A" docLanguage="en" docName="zootaxa.5496.2.9.pdf" docOrigin="Zootaxa 5496 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5496.2.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atteva wallengreni Sohn &amp; Wu 2013" docType="treatment" docVersion="2" lastPageNumber="292" masterDocId="8F51FFF89900627D5B587C714C612608" masterDocTitle="First report of the family Attevidae (Lepidoptera, Yponomeutoidea) from Cambodia, with description of a new species" masterLastPageNumber="294" masterPageNumber="290" pageNumber="292" updateTime="1724133637781" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="08B5EF8010E6A0A478418EB2964572D3">First report of the family Attevidae (Lepidoptera, Yponomeutoidea) from Cambodia, with description of a new species</mods:title>
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<mods:namePart id="3B528F19E2B31C62D995133D8B95655A">Na, Sol-Moon</mods:namePart>
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<mods:affiliation id="7CCC847EB60F73F3F2770A4A4F38A106">Yeongnam Regional Office, Animal and Plant Quarantine Agency, Busan, 39660, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="4175B8D72826257F121A90A2E6BF6B90" type="email">nasom86@gmail.com</mods:nameIdentifier>
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<mods:namePart id="81A91D6B261CED380965CD65B52AF089">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="4F3997E0BCD4D027FA26DFD4A7EC85AB" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="7AE4D355642857A0E5651F6ED9140507">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy-ro, Yeonsu-gu, Incheon, 22012, Republic of Korea &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo-dong, Incheon, 22012, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="FC7540EBE43B44C67F65F4DAB26B06E4" type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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<mods:title id="78CC8543B8E1B9830ADFBC1FF0FB8CE5">Zootaxa</mods:title>
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<mods:date id="83C56CCF1FD7B54780320A3C56B92643">2024</mods:date>
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<subSubSection id="B3DB651D9902627F5BCF7CE64E3E26B9" box="[151,607,151,178]" pageId="2" pageNumber="292" type="nomenclature">
<paragraph id="FB7E36969902627F5BCF7CE64E3E26B9" blockId="2.[151,607,151,213]" box="[151,607,151,178]" pageId="2" pageNumber="292">
<heading id="A03681FA9902627F5BCF7CE64E3E26B9" bold="true" box="[151,607,151,178]" fontSize="11" level="1" pageId="2" pageNumber="292" reason="1">
<taxonomicName id="3CC14D159902627F5BCF7CE64E3E26B9" ID-CoL="BWKPX" authority="Sohn &amp; Wu, 2013" authorityName="Sohn &amp; Wu" authorityYear="2013" box="[151,607,151,178]" class="Insecta" family="Attevidae" genus="Atteva" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="wallengreni">
<emphasis id="C9B5EA849902627F5BCF7CE64E3E26B9" bold="true" box="[151,607,151,178]" pageId="2" pageNumber="292">
<emphasis id="C9B5EA849902627F5BCF7CE64D1B26BA" bold="true" box="[151,378,151,178]" italics="true" pageId="2" pageNumber="292">Atteva wallengreni</emphasis>
<bibRefCitation id="9F504B679902627F5ADA7CE64E3E26B9" author="Sohn, J. C. &amp; Wu, C. S." box="[386,607,151,177]" pageId="2" pageNumber="292" pagination="1 - 16" refId="ref3156" refString="Sohn, J. C. &amp; Wu, C. S. (2013) Taxonomic review of Attevidae from China with descriptions of two new species and a revised identity of the Ailanthus webworm moth, Atteva fabriciella, from the Asian tropics. Journal of Insect Science, 13 (66), 1 - 16. https: // doi. org / 10.1673 / 031.013.6601" type="journal article" year="2013">Sohn &amp; Wu, 2013</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B3DB651D9902627F5BCF7CCA4D0926DD" box="[151,360,187,213]" pageId="2" pageNumber="292" type="description">
<paragraph id="FB7E36969902627F5BCF7CCA4D0926DD" blockId="2.[151,607,151,213]" box="[151,360,187,213]" pageId="2" pageNumber="292">
(
<figureCitation id="63FA2A139902627F5BC77CCA4D6026DD" box="[159,257,187,213]" captionStart="FIGURES 1" captionStartId="1.[151,262,1339,1363]" captionTargetBox="[151,1437,1021,1315]" captionTargetId="figure-143@1.[151,1437,1021,1315]" captionTargetPageId="1" captionText="FIGURES 1. Adults of Atteva spp.: A. A. fabriciella Swederus; B. A. wallengreni Sohn &amp; Wu; C. A. chalco Bae &amp; Na sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.13346725" httpUri="https://zenodo.org/record/13346725/files/figure.png" pageId="2" pageNumber="292">Figs. 1B</figureCitation>
,
<figureCitation id="63FA2A139902627F5A557CCA4D5326DD" box="[269,306,187,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,938,962]" captionTargetBox="[161,1425,305,904]" captionTargetId="figure-37@2.[151,1436,304,915]" captionTargetPageId="2" captionText="FIGURE 2. Male genitalia of Atteva spp.: A. A. wallengreni Sohn &amp; Wu, genitalia slide no. QNA-30; B. A. chalco sp. nov. Bae &amp; Na, genitalia slide no.QNA-28 (Paratype) (scale bar=1mm)." figureDoi="http://doi.org/10.5281/zenodo.13346727" httpUri="https://zenodo.org/record/13346727/files/figure.png" pageId="2" pageNumber="292">2A</figureCitation>
,
<figureCitation id="63FA2A139902627F5A667CCA4D3E26DD" box="[318,351,187,213]" captionStart="FIGURES 3" captionStartId="3.[151,264,1004,1028]" captionTargetBox="[161,1425,191,961]" captionTargetId="figure-22@3.[151,1436,181,980]" captionTargetPageId="3" captionText="FIGURES 3. Female genitalia of Atteva spp.: A. A. fabriciella Swederus, genitalia slide no. QNA-32; B. A. wallengreni Sohn &amp; Wu, genitalia slide no. QNA-33; C. A. chalco sp. nov., genitalia slide no. QNA-25 (Paratype) (scale bar=1mm)." figureDoi="http://doi.org/10.5281/zenodo.13346729" httpUri="https://zenodo.org/record/13346729/files/figure.png" pageId="2" pageNumber="292">3B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B3DB651D9902627F5BCF7D714F572710" box="[151,822,256,280]" pageId="2" pageNumber="292" type="reference_group">
<paragraph id="FB7E36969902627F5BCF7D714F572710" blockId="2.[151,822,256,280]" box="[151,822,256,280]" pageId="2" pageNumber="292">
<treatmentCitationGroup id="DBD111B89902627F5BCF7D714F572710" box="[151,822,256,280]" pageId="2" pageNumber="292">
<treatmentCitation id="7A6010879902627F5BCF7D714E51271F" author="Sohn, J. C. &amp; Wu, C. S." box="[151,560,256,280]" page="7" pageId="2" pageNumber="292" year="2013">
<taxonomicName id="3CC14D159902627F5BCF7D714E51271F" ID-CoL="BWKPX" authority="Sohn &amp; Wu, 2013: 7" authorityName="Sohn &amp; Wu" authorityPageNumber="7" authorityYear="2013" box="[151,560,256,280]" class="Insecta" family="Attevidae" genus="Atteva" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="wallengreni">
<emphasis id="C9B5EA849902627F5BCF7D714D372710" box="[151,342,256,280]" italics="true" pageId="2" pageNumber="292">Atteva wallengreni</emphasis>
<bibRefCitation id="9F504B679902627F5A057D714E51271F" author="Sohn, J. C. &amp; Wu, C. S." box="[349,560,256,280]" pageId="2" pageNumber="292" pagination="1 - 16" refId="ref3156" refString="Sohn, J. C. &amp; Wu, C. S. (2013) Taxonomic review of Attevidae from China with descriptions of two new species and a revised identity of the Ailanthus webworm moth, Atteva fabriciella, from the Asian tropics. Journal of Insect Science, 13 (66), 1 - 16. https: // doi. org / 10.1673 / 031.013.6601" type="journal article" year="2013">Sohn &amp; Wu, 2013: 7</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. TL:
<collectingCountry id="83D676069902627F593E7D714EC22710" box="[614,675,256,280]" name="China" pageId="2" pageNumber="292">China</collectingCountry>
(Guangzhou).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="AFBE661E9902627F5BCF7FDB4F7125EE" ID-DOI="http://doi.org/10.5281/zenodo.13346727" ID-Zenodo-Dep="13346727" httpUri="https://zenodo.org/record/13346727/files/figure.png" pageId="2" pageNumber="292" startId="2.[151,250,938,962]" targetBox="[161,1425,305,904]" targetPageId="2" targetType="figure">
<paragraph id="FB7E36969902627F5BCF7FDB4F7125EE" blockId="2.[151,1436,938,998]" pageId="2" pageNumber="292">
<emphasis id="C9B5EA849902627F5BCF7FDB4D7325CA" bold="true" box="[151,274,938,962]" pageId="2" pageNumber="292">FIGURE 2.</emphasis>
Male genitalia of
<taxonomicName id="3CC14D159902627F5A927FDB4E5C25CA" box="[458,573,938,962]" class="Insecta" family="Attevidae" genus="Atteva" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C9B5EA849902627F5A927FDB4E6A25C9" box="[458,523,938,961]" italics="true" pageId="2" pageNumber="292">Atteva</emphasis>
spp.
</taxonomicName>
:
<emphasis id="C9B5EA849902627F591E7FDB4E3E25CA" bold="true" box="[582,607,938,962]" pageId="2" pageNumber="292">A.</emphasis>
<taxonomicName id="3CC14D159902627F593C7FDB4F1725CA" authority="Sohn &amp; Wu" authorityName="Sohn &amp; Wu" authorityYear="2013" box="[612,886,938,962]" class="Insecta" family="Attevidae" genus="Atteva" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="wallengreni">
<emphasis id="C9B5EA849902627F593C7FDB4E9925CA" box="[612,760,938,962]" italics="true" pageId="2" pageNumber="292">A. wallengreni</emphasis>
Sohn &amp; Wu
</taxonomicName>
, genitalia slide no. QNA-30;
<emphasis id="C9B5EA849902627F5FC57FDB48D525CA" bold="true" box="[1181,1204,938,962]" pageId="2" pageNumber="292">B.</emphasis>
<taxonomicName id="3CC14D159902627F5FE27FDB497B25CA" authorityName="Na &amp; Bae" authorityYear="2024" box="[1210,1306,938,962]" class="Insecta" family="Attevidae" genus="Atteva" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="chalco" status="sp. nov.">
<emphasis id="C9B5EA849902627F5FE27FDB497B25CA" box="[1210,1306,938,962]" italics="true" pageId="2" pageNumber="292">A. chalco</emphasis>
</taxonomicName>
<emphasis id="C9B5EA849902627F5E477FDB490E25CA" bold="true" box="[1311,1391,938,962]" pageId="2" pageNumber="292">
<taxonomicNameLabel id="D28657FF9902627F5E477FDB490E25CA" box="[1311,1391,938,962]" pageId="2" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Bae &amp; Na, genitalia slide no.QNA-28 (Paratype) (scale bar=1mm).
</paragraph>
</caption>
<subSubSection id="B3DB651D9902627F5BCF78494F9122B6" pageId="2" pageNumber="292" type="diagnosis">
<paragraph id="FB7E36969902627F5BCF78494F9122B6" blockId="2.[151,1437,1080,1538]" pageId="2" pageNumber="292">
<emphasis id="C9B5EA849902627F5BCF78494D73225A" bold="true" box="[151,274,1080,1106]" pageId="2" pageNumber="292">Diagnosis.</emphasis>
This species is superficially similar to
<taxonomicName id="3CC14D159902627F599E78484841225A" authority="Swerderus, 1787" authorityName="Swerderus" authorityYear="1787" box="[710,1056,1080,1107]" class="Insecta" family="Attevidae" genus="Phalaena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="fabriciella">
<emphasis id="C9B5EA849902627F599E78484F3B225A" box="[710,858,1080,1107]" italics="true" pageId="2" pageNumber="292">A. fabriciella</emphasis>
Swerderus, 1787
</taxonomicName>
, but it can be distinguished by the following characters: frequently more than 40 white dots on the forewing; a comb-like spinose on socii and apex of uncus larger and slenderer respectively; costa of valva without protrusion; corpus bursae without conspicuous pleats, signum smaller than that of
<taxonomicName id="3CC14D159902627F597878D44ED522B6" authorityName="Swerderus" authorityYear="1787" box="[544,692,1188,1215]" class="Insecta" family="Attevidae" genus="Phalaena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="fabriciella">
<emphasis id="C9B5EA849902627F597878D44ED522B6" box="[544,692,1188,1215]" italics="true" pageId="2" pageNumber="292">A. fabriciella</emphasis>
</taxonomicName>
(refer to
<bibRefCitation id="9F504B679902627F584578D54F8422B6" author="Sohn, J. C. &amp; Wu, C. S." box="[797,997,1188,1215]" pageId="2" pageNumber="292" pagination="1 - 16" refId="ref3156" refString="Sohn, J. C. &amp; Wu, C. S. (2013) Taxonomic review of Attevidae from China with descriptions of two new species and a revised identity of the Ailanthus webworm moth, Atteva fabriciella, from the Asian tropics. Journal of Insect Science, 13 (66), 1 - 16. https: // doi. org / 10.1673 / 031.013.6601" type="journal article" year="2013">Sohn &amp; Wu 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B3DB651D9902627F5B9F78B94F09230E" pageId="2" pageNumber="292" type="distribution">
<paragraph id="FB7E36969902627F5B9F78B94F09230E" blockId="2.[151,1437,1080,1538]" pageId="2" pageNumber="292">
<emphasis id="C9B5EA849902627F5B9F78B94D0022EA" bold="true" box="[199,353,1224,1250]" pageId="2" pageNumber="292">Distribution.</emphasis>
<collectingCountry id="83D676069902627F5A3178B94DBD22EA" box="[361,476,1224,1250]" name="Cambodia" pageId="2" pageNumber="292">Cambodia</collectingCountry>
(Present study),
<collectingCountry id="83D676069902627F59C678B94E8022EA" box="[670,737,1224,1250]" name="China" pageId="2" pageNumber="292">China</collectingCountry>
(
<collectingRegion id="3905F8749902627F59AA78B94F1822EA" box="[754,889,1224,1250]" country="China" name="Guangdong" pageId="2" pageNumber="292">Guangdong</collectingRegion>
,
<collectingRegion id="3905F8749902627F58DD78B94FB822EA" box="[901,985,1224,1250]" country="China" name="Hainan" pageId="2" pageNumber="292">Hainan</collectingRegion>
,
<collectingCountry id="83D676069902627F58BE78B9483E22EA" box="[998,1119,1224,1250]" name="Hong Kong" pageId="2" pageNumber="292">Hongkong</collectingCountry>
, Shaanxi),
<collectingCountry id="83D676069902627F5F8778B9492D22EA" box="[1247,1356,1224,1250]" name="Indonesia" pageId="2" pageNumber="292">Indonesia</collectingCountry>
(
<collectingRegion id="3905F8749902627F5E0478B949EE22EA" box="[1372,1423,1224,1250]" country="Indonesia" name="Bali" pageId="2" pageNumber="292">Bali</collectingRegion>
),
<collectingCountry id="83D676069902627F5BCF789D4C9F230E" box="[151,254,1260,1286]" name="Malaysia" pageId="2" pageNumber="292">Malaysia</collectingCountry>
(
<collectingRegion id="3905F8749902627F5A55789D4D32230E" box="[269,339,1260,1286]" country="Malaysia" name="Perak" pageId="2" pageNumber="292">Perak</collectingRegion>
),
<collectingCountry id="83D676069902627F5A3D789D4DA8230E" box="[357,457,1260,1286]" name="Thailand" pageId="2" pageNumber="292">Thailand</collectingCountry>
(Chiangmai),
<collectingCountry id="83D676069902627F5932789D4EA8230E" box="[618,713,1260,1286]" name="Vietnam" pageId="2" pageNumber="292">Vietnam</collectingCountry>
(Hochiminh).
</paragraph>
</subSubSection>
<subSubSection id="B3DB651D9902627F5B9F79614DA82322" box="[199,457,1296,1322]" pageId="2" pageNumber="292" type="biology_ecology">
<paragraph id="FB7E36969902627F5B9F79614DA82322" blockId="2.[151,1437,1080,1538]" box="[199,457,1296,1322]" pageId="2" pageNumber="292">
<emphasis id="C9B5EA849902627F5B9F79614D2A2322" bold="true" box="[199,331,1296,1322]" pageId="2" pageNumber="292">Host plant.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B3DB651D9902627F5B9F79454817239E" pageId="2" pageNumber="292" type="materials_examined">
<paragraph id="FB7E36969902627F5B9F79454817239E" blockId="2.[151,1437,1080,1538]" pageId="2" pageNumber="292">
<emphasis id="C9B5EA849902627F5B9F79454DD02346" bold="true" box="[199,433,1332,1358]" pageId="2" pageNumber="292">Material examined.</emphasis>
[
<collectionCode id="9DD0AE539902627F5AE779444E682347" box="[447,521,1333,1359]" pageId="2" pageNumber="292">INUB</collectionCode>
]
<specimenCount id="EDC7FD1F9902627F594E79444E4A2347" box="[534,555,1333,1359]" count="1" pageId="2" pageNumber="292" type="female"></specimenCount>
, Seima (
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,
<geoCoordinate id="9EF550519902627F587D79454FD82347" box="[805,953,1332,1359]" degrees="107" direction="east" minutes="03" orientation="longitude" pageId="2" pageNumber="292" precision="15" seconds="15" value="107.05417">E107º0315”</geoCoordinate>
),
<date id="8F7F10569902627F58967945482A2346" box="[974,1099,1332,1358]" pageId="2" pageNumber="292" value="2011-11-11">11.XI.2011</date>
(Bae Y.S., Kim Y.K., Ju Y.D., Park B.S., Lee H.J.), genitalia slide no. QNA-33.
<specimenCount id="EDC7FD1F9902627F59E079284EB3237B" box="[696,722,1369,1395]" count="1" pageId="2" pageNumber="292" type="male"></specimenCount>
<specimenCount id="EDC7FD1F9902627F598A79284E85237B" box="[722,740,1369,1395]" count="1" pageId="2" pageNumber="292" type="female"></specimenCount>
, Samkos (
<geoCoordinate id="9EF550519902627F580E79294F94237A" box="[854,1013,1368,1394]" degrees="12" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="292" precision="1" seconds="38.8" value="12.210777">N12º1238.8”</geoCoordinate>
,
<geoCoordinate id="9EF550519902627F58A7792948C7237B" box="[1023,1190,1368,1395]" degrees="102" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="292" precision="1" seconds="54.3" value="102.898415">E102º5354.3”</geoCoordinate>
Alt.
<quantity id="3C399B739902627F5F8279294942237A" box="[1242,1315,1368,1394]" metricMagnitude="2" metricUnit="m" metricValue="9.34" pageId="2" pageNumber="292" unit="m" value="934.0">934 m</quantity>
),
<date id="8F7F10569902627F5E6E792949FD237B" box="[1334,1436,1368,1395]" pageId="2" pageNumber="292" value="2015-02-08">8.II.2015</date>
(Bae Y.S., Ju Y.D., Park B.S., Na S.M., Kim J.W., Lee D.J.), genitalia slide no. QNA-31.
</paragraph>
</subSubSection>
<subSubSection id="B3DB651D9902627F5B9F79D14CB0200A" pageId="2" pageNumber="292" type="discussion">
<paragraph id="FB7E36969902627F5B9F79D14CB0200A" blockId="2.[151,1437,1080,1538]" pageId="2" pageNumber="292">
<emphasis id="C9B5EA849902627F5B9F79D14D5523B2" bold="true" box="[199,308,1440,1466]" pageId="2" pageNumber="292">Remarks</emphasis>
.
<bibRefCitation id="9F504B679902627F5A1B79D14E7A23B2" author="Sohn, J. C. &amp; Wu, C. S." box="[323,539,1440,1467]" pageId="2" pageNumber="292" pagination="1 - 16" refId="ref3156" refString="Sohn, J. C. &amp; Wu, C. S. (2013) Taxonomic review of Attevidae from China with descriptions of two new species and a revised identity of the Ailanthus webworm moth, Atteva fabriciella, from the Asian tropics. Journal of Insect Science, 13 (66), 1 - 16. https: // doi. org / 10.1673 / 031.013.6601" type="journal article" year="2013">Sohn &amp; Wu (2013)</bibRefCitation>
described
<taxonomicName id="3CC14D159902627F59CE79D04F5A23B2" authorityName="Sohn &amp; Wu" authorityYear="2013" box="[662,827,1440,1467]" class="Insecta" family="Attevidae" genus="Atteva" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="292" phylum="Arthropoda" rank="species" species="wallengreni">
<emphasis id="C9B5EA849902627F59CE79D04F5A23B2" box="[662,827,1440,1467]" italics="true" pageId="2" pageNumber="292">A. wallengreni</emphasis>
</taxonomicName>
with or without triangular black scales on their frons. We collected only full white frons individuals in this study. This species is reported from
<collectingCountry id="83D676069902627F5FC679B5497023D6" box="[1182,1297,1476,1502]" name="Cambodia" pageId="2" pageNumber="292">Cambodia</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>