treatments-xml/data/03/B7/87/03B787F6FFDD9B7E61C37956FE06F98C.xml

507 lines
57 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="C5DF183CBCE8FBD2CF57F82866BB13AD" ID-DOI="10.17348/jbrit.v15.i2.1206" ID-Zenodo-Dep="14111022" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_requiresApprovalFor="plazi" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1634280027544" checkinUser="donat" docAuthor="Fawcett, Susan &amp; Smith, Alan R." docDate="2021" docId="03B787F6FFDD9B7E61C37956FE06F98C" docLanguage="en" docName="2021_ThelypteridaceaeFerns_OACopy3.pdf.imf" docOrigin="Fort Worth, Texas, USA: BRIT Press" docTitle="Glaphyropteridopsis" docType="treatment" docVersion="4" lastPageNumber="43" masterDocId="FF8EFF8EFFE99B4B602B7D49FFA7FFBB" masterDocTitle="A Generic Classification of the Thelypteridaceae" pageNumber="42" updateTime="1731444927364" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
<mods:mods id="C00004F458EF5199372B50AE3E3F3B3A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0B37D9855769ECCC1F765B8C5707E80">
<mods:title id="E181EE3C711D5A7CB7DAA401DA0EE0E6">A Generic Classification of the Thelypteridaceae</mods:title>
</mods:titleInfo>
<mods:name id="DDA6F4F383FA047654E1D27EBEE99266" type="personal">
<mods:role id="BF084C53B246B57C3D82FED631FA9024">
<mods:roleTerm id="C2E23E7B305B35D41D5446BC27973BA2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA49815244D1FC8E741E2DFF616F9E34">Fawcett, Susan</mods:namePart>
</mods:name>
<mods:name id="9A0256425D1A2A6ADAC4674068D2703D" type="personal">
<mods:role id="5B6EF91F812C4685DC0AFA952FD5A30D">
<mods:roleTerm id="76858300B3145E3A5747228AA1667B53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B356734CF702CE645660D3BB4734181">Smith, Alan R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="969299A154371B6DA2D8EC812A902CA7">text</mods:typeOfResource>
<mods:originInfo id="5EA83F330B543AB833646574FEDE7386">
<mods:dateIssued id="95E78225F6EDD785CEE97CACAE125EF3">2021</mods:dateIssued>
<mods:publisher id="8578C5550562E947E878920CECE2CFC6">BRIT Press</mods:publisher>
<mods:place id="73E8FADC035A68F00B6EDC07163CE9E9">
<mods:placeTerm id="AC0353ACFA708FD51C68179896F87D51">Fort Worth, Texas, USA</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:classification id="7DD1D6EBAB521A1B463ACF6EE10554E2">book</mods:classification>
<mods:identifier id="41F9B1381A5DEB479D44B203FC35CA91" type="DOI">10.17348/jbrit.v15.i2.1206</mods:identifier>
<mods:identifier id="6E1235303618DEF5ADCDEB959F1EDF4C" type="Zenodo-Dep">14111022</mods:identifier>
</mods:mods>
<treatment id="03B787F6FFDD9B7E61C37956FE06F98C" ID-DOI="http://doi.org/10.5281/zenodo.14076332" ID-Zenodo-Dep="14076332" LSID="urn:lsid:plazi:treatment:03B787F6FFDD9B7E61C37956FE06F98C" httpUri="http://treatment.plazi.org/id/03B787F6FFDD9B7E61C37956FE06F98C" lastPageId="53" lastPageNumber="43" pageId="52" pageNumber="42">
<subSubSection id="C304656BFFDD9B7F61C37956FCE6FB82" box="[488,833,1055,1081]" pageId="52" pageNumber="42" type="nomenclature">
<paragraph id="8BA136E0FFDD9B7F61C37956FCE6FB82" blockId="52.[97,1227,1055,1143]" box="[488,833,1055,1081]" pageId="52" pageNumber="42">
<heading id="D0E9818CFFDD9B7F61C37956FCE6FB82" allCaps="true" bold="true" box="[488,833,1055,1081]" centered="true" fontSize="11" level="2" pageId="52" pageNumber="42" reason="0">
<taxonomicName id="4C1E4D63FFDD9B7F61C37956FCE6FB82" ID-CoL="4NSR" box="[488,833,1055,1081]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDD9B7F61C37956FCE6FB82" bold="true" box="[488,833,1055,1081]" pageId="52" pageNumber="42">GLAPHYROPTERIDOPSIS</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDD9B7F604A7901FE2EFBCC" pageId="52" pageNumber="42" type="reference_group">
<paragraph id="8BA136E0FFDD9B7F604A7901FE2EFBCC" blockId="52.[97,1227,1055,1143]" pageId="52" pageNumber="42">
<treatmentCitation id="0ABF10F1FFDD9B7F604A7901FD04FBE6" author="Glaphyropteridopsis Ching" box="[97,675,1096,1117]" journal="Acta Phytotax. Sin." page="320" pageId="52" pageNumber="42" score="3.863100345872848" volume="8" year="1963">
<taxonomicName id="4C1E4D63FFDD9B7F604A7901FD96FBE6" ID-CoL="4NSR" authority="Ching, Acta Phytotax. Sin." authorityName="Ching, Acta Phytotax. Sin." box="[97,561,1096,1117]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDD9B7F604A7901FE94FBE6" bold="true" box="[97,307,1096,1117]" pageId="52" pageNumber="42">Glaphyropteridopsis</emphasis>
Ching, Acta Phytotax. Sin.
</taxonomicName>
8:320. 1963
</treatmentCitation>
.
<emphasis id="B96AEAF2FFDD9B7F628D790EFD1AFBE7" box="[678,701,1095,1116]" italics="true" pageId="52" pageNumber="42"></emphasis>
<treatmentCitation id="0ABF10F1FFDD9B7F62967903FE22FBCC" author="K. Iwats." journal="Mem. Coll. Sci. Univ. Kyoto, ser. B" page="29" pageId="52" pageNumber="42" score="10.725328331126558" volume="31" year="1964">
<taxonomicName id="4C1E4D63FFDD9B7F62967903FF6CFBCC" authority="(Ching) K. Iwats., Mem. Coll. Sci. Univ." baseAuthorityName="Ching" class="Polypodiopsida" family="Thelypteridaceae" genus="Thelypteris" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="section" section="Glaphyropteridopsis">
<emphasis id="B96AEAF2FFDD9B7F62967903FCA8FBE0" box="[701,783,1098,1115]" italics="true" pageId="52" pageNumber="42">Thelypteris</emphasis>
sect.
<emphasis id="B96AEAF2FFDD9B7F63137903FC6BFBE0" box="[824,972,1098,1115]" italics="true" pageId="52" pageNumber="42">Glaphyropteridopsis</emphasis>
(Ching) K. Iwats., Mem. Coll.Sci. Univ.
</taxonomicName>
<collectingRegion id="49DAF802FFDD9B7F60E6792CFF5CFBCE" box="[205,251,1125,1141]" country="Japan" name="Kyoto" pageId="52" pageNumber="42">Kyoto</collectingRegion>
, ser.
<collectionCode id="ED0FAE25FFDD9B7F6135792CFE8DFBCE" box="[286,298,1125,1141]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="52" pageNumber="42" type="Herbarium">B</collectionCode>
, 31:29.1964
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDD9B7F61A3792FFBB5FBCC" box="[392,1042,1125,1143]" pageId="52" pageNumber="42" type="type_taxon">
<paragraph id="8BA136E0FFDD9B7F61A3792FFBB5FBCC" blockId="52.[97,1227,1055,1143]" box="[392,1042,1125,1143]" pageId="52" pageNumber="42">
<typeStatus id="54A58842FFDD9B7F61B7792CFE1CFBCE" box="[412,443,1125,1141]" pageId="52" pageNumber="42">
<collectionCode id="ED0FAE25FFDD9B7F61B7792CFE1CFBCE" box="[412,443,1125,1141]" pageId="52" pageNumber="42">TYPE</collectionCode>
</typeStatus>
:
<taxonomicName id="4C1E4D63FFDD9B7F61EF792CFCBBFBCC" ID-CoL="6KGYK" authority="(Hook.) Ching" authorityName="Ching" baseAuthorityName="Hook." box="[452,796,1125,1143]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDD9B7F61EF792CFD0DFBCD" box="[452,682,1125,1142]" italics="true" pageId="52" pageNumber="42">Glaphyropteridopsis erubescens</emphasis>
(Hook.) Ching
</taxonomicName>
[=
<taxonomicName id="4C1E4D63FFDD9B7F6318792CFBA8FBCC" ID-CoL="4LFD7" authority="Hook." authorityName="Hook." box="[819,1039,1125,1143]" class="Polypodiopsida" family="Thelypteridaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDD9B7F6318792CFC7DFBCD" box="[819,986,1125,1142]" italics="true" pageId="52" pageNumber="42">Polypodium erubescens</emphasis>
Hook.
</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDD9B7F60AA79C2FDC4FB27" box="[129,611,1163,1180]" pageId="52" pageNumber="42" type="reference_group">
<paragraph id="8BA136E0FFDD9B7F60AA79C2FDC4FB27" blockId="52.[129,611,1163,1180]" box="[129,611,1163,1180]" pageId="52" pageNumber="42">
For additional synonymy, see
<bibRefCitation id="EF8F4B11FFDD9B7F614A79C2FE62FB27" author="Ching" box="[353,453,1163,1180]" firstAuthor="Ching" journalOrPublisher="Acta Phytotax. Sin." pageId="52" pageNumber="42" pagination="289 - 335" part="8" refId="ref107523" refString="CHING, R. C. 1963. A reclassification of the family Thelypteridaceae from the mainland of Asia. Acta Phytotax. Sin. 8: 289 - 335." title="A reclassification of the family Thelypteridaceae from the mainland of Asia" type="journal article" year="1963">Ching (1963)</bibRefCitation>
and
<bibRefCitation id="EF8F4B11FFDD9B7F61C279C2FDF8FB27" author="Holttum" box="[489,607,1163,1180]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="52" pageNumber="42" pagination="17 - 52" part="19" refId="ref109463" refString="HOLTTUM, R. E. 1971. Studies in the family Thelypteridaceae III. A new system of genera in the Old World. Blumea 19: 17 - 52." title="Studies in the family Thelypteridaceae III. A new system of genera in the Old World" type="journal article" year="1971">Holttum (1971)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDD9B7F604A79F8FC19FABC" pageId="52" pageNumber="42" type="etymology">
<paragraph id="8BA136E0FFDD9B7F604A79F8FC19FABC" blockId="52.[97,1226,1201,1799]" pageId="52" pageNumber="42">
<emphasis id="B96AEAF2FFDD9B7F604A79F8FF47FB7D" box="[97,224,1201,1222]" italics="true" pageId="52" pageNumber="42">Etymology.—</emphasis>
From
<emphasis id="B96AEAF2FFDD9B7F613279F8FE03FB7D" box="[281,420,1201,1222]" italics="true" pageId="52" pageNumber="42">Glaphyropteris</emphasis>
(Gr.
<emphasis id="B96AEAF2FFDD9B7F61FA79F8FD8EFB7D" box="[465,553,1201,1222]" italics="true" pageId="52" pageNumber="42">glaphyros</emphasis>
, hollow +
<emphasis id="B96AEAF2FFDD9B7F62A079FBFD18FB7C" box="[651,703,1202,1223]" italics="true" pageId="52" pageNumber="42">pteris</emphasis>
, fern) +
<emphasis id="B96AEAF2FFDD9B7F632779FBFCE6FB7C" box="[780,833,1202,1223]" italics="true" pageId="52" pageNumber="42">-opsis</emphasis>
, like.
<bibRefCitation id="EF8F4B11FFDD9B7F635279FBFC5FFB7C" author="Ching" box="[889,1016,1202,1223]" firstAuthor="Ching" journalOrPublisher="Acta Phytotax. Sin." pageId="52" pageNumber="42" pagination="289 - 335" part="8" refId="ref107523" refString="CHING, R. C. 1963. A reclassification of the family Thelypteridaceae from the mainland of Asia. Acta Phytotax. Sin. 8: 289 - 335." title="A reclassification of the family Thelypteridaceae from the mainland of Asia" type="journal article" year="1963">Ching (1963)</bibRefCitation>
alluded to the general similarity in frond size and blade dissection of
<taxonomicName id="4C1E4D63FFDD9B7F62367998FD7FFB5D" box="[541,728,1233,1254]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDD9B7F62367998FD7FFB5D" box="[541,728,1233,1254]" italics="true" pageId="52" pageNumber="42">Glaphyropteridopsis</emphasis>
</taxonomicName>
to
<emphasis id="B96AEAF2FFDD9B7F62DE7998FC27FB5D" box="[757,896,1233,1254]" italics="true" pageId="52" pageNumber="42">Glaphyropteris</emphasis>
(=
<taxonomicName id="4C1E4D63FFDD9B7F63B6799BFBA1FB5C" box="[925,1030,1234,1255]" class="Polypodiopsida" family="Thelypteridaceae" genus="Steiropteris" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDD9B7F63B6799BFBA1FB5C" box="[925,1030,1234,1255]" italics="true" pageId="52" pageNumber="42">Steiropteris</emphasis>
</taxonomicName>
in our treatment) in the Neotropics. However, these two genera are not closely related, as understood by Ching.
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDD9B7E60A4785BFEE9FE8C" lastPageId="53" lastPageNumber="43" pageId="52" pageNumber="42" type="description">
<paragraph id="8BA136E0FFDD9B7E60A4785BFEE9FE8C" blockId="52.[97,1226,1201,1799]" lastBlockId="53.[121,1250,162,1591]" lastPageId="53" lastPageNumber="43" pageId="52" pageNumber="42">
Plants terrestrial or on (among) rocks, medium-sized to large, 50150(
<quantity id="4CE69B05FFDD9B7F6311785BFC25FA9C" box="[826,898,1298,1319]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="42" unit="cm" value="300.0">300 cm</quantity>
);
<emphasis id="B96AEAF2FFDD9B7F63B9785BFC57FA9C" bold="true" box="[914,1008,1298,1319]" pageId="52" pageNumber="42">rhizomes</emphasis>
short and thick, sometimes massive, short-creeping or ascending, with sparse, ± glabrous scales at apices;
<emphasis id="B96AEAF2FFDD9B7F63A1787BFC6AFAFC" bold="true" box="[906,973,1330,1351]" pageId="52" pageNumber="42">fronds</emphasis>
clustered or approximate, monomorphic, arching;
<emphasis id="B96AEAF2FFDD9B7F6160781BFE21FADC" bold="true" box="[331,390,1362,1383]" pageId="52" pageNumber="42">stipes</emphasis>
stramineous to tan, usually stout (2)410+ mm diam., 30100+ cm long, with sparse brown to tan, ovate, thin appressed scales at bases, with sparse acicular hairs or glabrescent;
<emphasis id="B96AEAF2FFDD9B7F63F97838FBB3FA3D" bold="true" box="[978,1044,1393,1414]" pageId="52" pageNumber="42">blades</emphasis>
herbaceous, chartaceous or leathery, drying yellowish green to reddish brown, elliptic, with truncate bases (lacking greatly reduced pinnae), pinnate-pinnatifid, lacking pustules or glands, and generally lacking hairs between veins;
<emphasis id="B96AEAF2FFDD9B7F63D978FBFBE0FA7C" bold="true" box="[1010,1095,1458,1479]" pageId="52" pageNumber="42">rachises</emphasis>
stramineous;
<emphasis id="B96AEAF2FFDD9B7F604A789BFF00FA5C" bold="true" box="[97,167,1490,1511]" pageId="52" pageNumber="42">pinnae</emphasis>
sessile, opposite or subopposite, often large (to 50 ×
<quantity id="4CE69B05FFDD9B7F62A4789BFD1AFA5C" box="[655,701,1490,1511]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="52" pageNumber="42" unit="cm" value="5.0">5 cm</quantity>
in
<taxonomicName id="4C1E4D63FFDD9B7F62F2789BFCF3FA5D" box="[729,852,1489,1511]" class="Polypodiopsida" family="Thelypteridaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="52" pageNumber="42" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDD9B7F62F2789BFCF3FA5D" box="[729,852,1489,1511]" italics="true" pageId="52" pageNumber="42">G. erubescens</emphasis>
</taxonomicName>
), linear-lanceolate, deeply pinnatifid or nearly pinnatisect (incised within 1(2) mm of costae), lowermost pinnae often narrowed at their bases, deflexed, costae grooved adaxially and sparsely to densely hairy abaxially; segments large, falcate-lanceolate, those at the base of the larger pinnae often somewhat elongate and/or toothed, and reflexed to overlap the rachis;
<emphasis id="B96AEAF2FFDD9B7F63947B78FB97F9FD" bold="true" box="[959,1072,1585,1606]" pageId="52" pageNumber="42">aerophores</emphasis>
absent at pinna bases or with only a small darkened patch in dried fronds;
<emphasis id="B96AEAF2FFDD9B7F62B97B1BFCFEF9DC" bold="true" box="[658,857,1618,1639]" pageId="52" pageNumber="42">indument abaxially</emphasis>
of sparse to dense acicular hairs along rachis, costae, costules, and veins, seldom with hairs between veins, glands (both sessile and stipitate) and scales lacking;
<emphasis id="B96AEAF2FFDD9B7F60A17BDBFEF6F91C" bold="true" box="[138,337,1682,1703]" pageId="52" pageNumber="42">indument adaxially</emphasis>
of usually dense hairs along costal grooves, costules, veins, and tissue between veins usually glabrous;
<emphasis id="B96AEAF2FFDD9B7F60957BFBFF54F97C" bold="true" box="[190,243,1714,1735]" pageId="52" pageNumber="42">veins</emphasis>
free, pinnate on segments, veinlets simple, to 25 pairs per segment, prominent and reaching margins, proximal pair running alongside transparent membrane below sinus or to sinus, or to margins just above sinus;
<emphasis id="B96AEAF2FFDD9B7F604A7BB8FF1FF8BD" bold="true" box="[97,184,1777,1798]" pageId="52" pageNumber="42">pustules</emphasis>
absent on laminar tissue;
<emphasis id="B96AEAF2FFDD9B7F61937BBBFE47F8BC" bold="true" box="[440,480,1778,1799]" pageId="52" pageNumber="42">sori</emphasis>
orbicular, exindusiate or with small indusia, these glabrous or with acicular hairs, subcostular and attached at bases of veinlets in a row on each side, close to or overlapping costules at maturity, usually confluent into a line when mature;
<emphasis id="B96AEAF2FFDC9B7E623E7D8BFDDDFF6C" bold="true" box="[533,634,194,215]" pageId="53" pageNumber="43">sporangia</emphasis>
glabrous or with hyaline setulae near annulus, these sometimes numerous;
<emphasis id="B96AEAF2FFDC9B7E60CF7DA8FE81FF4D" bold="true" box="[228,294,225,246]" pageId="53" pageNumber="43">spores</emphasis>
brown, elliptic, bilateral, irregularly reticulate-echinate, with perforations (
<bibRefCitation id="EF8F4B11FFDC9B7E63DB7DABFB7CFF4C" author="Tryon &amp; Lugardon" box="[1008,1243,226,247]" firstAuthor="Tryon" journalOrPublisher="Springer-Verlag, New York, U. S. A" pageId="53" pageNumber="43" refId="ref115218" refString="TRYON, A. F. &amp; B. LUGARDON. 1991. Spores of Pteridophyta. Springer-Verlag, New York, U. S. A." title="Spores of Pteridophyta" type="book" year="1991">Tryon &amp; Lugardon 1991</bibRefCitation>
) or with small tubercules on surfaces;
<emphasis id="B96AEAF2FFDC9B7E61CB7C4BFE4CFEAC" box="[480,491,258,279]" italics="true" pageId="53" pageNumber="43">x</emphasis>
= 36. Two species counted,
<taxonomicName id="4C1E4D63FFDC9B7E62DC7C4BFCD1FEAD" box="[759,886,257,279]" class="Polypodiopsida" family="Thelypteridaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDC9B7E62DC7C4BFCD1FEAD" box="[759,886,257,279]" italics="true" pageId="53" pageNumber="43">G. erubescens</emphasis>
</taxonomicName>
, both diploids and tetraploids, and
<taxonomicName id="4C1E4D63FFDC9B7E64E07C4BFF5EFE8D" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="rufostraminea">
<emphasis id="B96AEAF2FFDC9B7E64E07C4BFF5EFE8D" italics="true" pageId="53" pageNumber="43">G. rufostraminea</emphasis>
</taxonomicName>
, diploid.
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDC9B7E60827C0BFBC7FC8C" pageId="53" pageNumber="43" type="diagnosis">
<paragraph id="8BA136E0FFDC9B7E60827C0BFBC7FC8C" blockId="53.[121,1250,162,1591]" pageId="53" pageNumber="43">
<emphasis id="B96AEAF2FFDC9B7E60827C0BFE7BFEED" box="[169,476,321,343]" italics="true" pageId="53" pageNumber="43">
Diagnosis.—
<taxonomicName id="4C1E4D63FFDC9B7E610A7C08FE7BFEED" box="[289,476,321,342]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">Glaphyropteridopsis</taxonomicName>
</emphasis>
resembles
<taxonomicName id="4C1E4D63FFDC9B7E626C7C08FD37FEED" box="[583,656,321,342]" class="Polypodiopsida" family="Thelypteridaceae" genus="Chingia" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E626C7C08FD37FEED" box="[583,656,321,342]" italics="true" pageId="53" pageNumber="43">Chingia</emphasis>
</taxonomicName>
in frond
<taxonomicName id="4C1E4D63FFDC9B7E62C37C0BFCE7FEEC" box="[744,832,322,343]" form="and" pageId="53" pageNumber="43" rank="form">form and</taxonomicName>
size, the truncate blade bases, and position of the sori, but the veins of the former are always free (vs. anastomosing below sinuses, except in
<taxonomicName id="4C1E4D63FFDC9B7E64007C2BFBDEFECC" box="[1067,1145,354,375]" class="Polypodiopsida" family="Thelypteridaceae" genus="Chingia" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="pricei">
<emphasis id="B96AEAF2FFDC9B7E64007C2BFBDEFECC" box="[1067,1145,354,375]" italics="true" pageId="53" pageNumber="43">C. pricei</emphasis>
</taxonomicName>
). Scales of
<taxonomicName id="4C1E4D63FFDC9B7E60527CC8FF65FE2D" box="[121,194,385,406]" class="Polypodiopsida" family="Thelypteridaceae" genus="Chingia" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60527CC8FF65FE2D" box="[121,194,385,406]" italics="true" pageId="53" pageNumber="43">Chingia</emphasis>
</taxonomicName>
are bristly, numerous, and spreading (vs. appressed, tan, ovate in
<taxonomicName id="4C1E4D63FFDC9B7E636D7CC8FBA7FE2D" box="[838,1024,385,406]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E636D7CC8FBA7FE2D" box="[838,1024,385,406]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
), and spores are sometimes reticulate and perforate (vs. echinate) (
<bibRefCitation id="EF8F4B11FFDC9B7E620B7CEBFD0FFE0C" author="Holttum" box="[544,680,418,439]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="53" pageNumber="43" pagination="17 - 52" part="19" refId="ref109463" refString="HOLTTUM, R. E. 1971. Studies in the family Thelypteridaceae III. A new system of genera in the Old World. Blumea 19: 17 - 52." title="Studies in the family Thelypteridaceae III. A new system of genera in the Old World" type="journal article" year="1971">Holttum 1971</bibRefCitation>
,
<bibRefCitation id="EF8F4B11FFDC9B7E629B7CEBFD45FE0C" author="Holttum" box="[688,738,418,439]" firstAuthor="Holttum" journalOrPublisher="Flora Malesiana" pageId="53" pageNumber="43" pagination="334 - 560" part="1" refId="ref109955" refString="HOLTTUM, R. E. 1982. Thelypteridaceae. Flora Malesiana, Ser. II, 1 (5): 334 - 560." title="Thelypteridaceae" type="journal article" year="1982">1982</bibRefCitation>
). All species of
<taxonomicName id="4C1E4D63FFDC9B7E63597CE8FB8AFE0D" box="[882,1069,417,438]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E63597CE8FB8AFE0D" box="[882,1069,417,438]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
except
<taxonomicName id="4C1E4D63FFDC9B7E645E7CEBFF14FE6C" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="jinfushanensis">
<emphasis id="B96AEAF2FFDC9B7E645E7CEBFF14FE6C" italics="true" pageId="53" pageNumber="43">G. jinfushanensis</emphasis>
</taxonomicName>
have subcostular sori, a character that sets it apart from most other thelypterid species and genera, except for some species of
<taxonomicName id="4C1E4D63FFDC9B7E61207CA8FEF3FE4D" box="[267,340,481,502]" class="Polypodiopsida" family="Thelypteridaceae" genus="Chingia" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E61207CA8FEF3FE4D" box="[267,340,481,502]" italics="true" pageId="53" pageNumber="43">Chingia</emphasis>
</taxonomicName>
; however,this tendency for the sori to overlap the costules is most extreme in
<taxonomicName id="4C1E4D63FFDC9B7E640B7CA8FB7DFE4D" box="[1056,1242,481,502]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E640B7CA8FB7DFE4D" box="[1056,1242,481,502]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
. Another strong synapomorphy of the genus is the tendency to have strictly opposite or at least subopposite pinnae. This feature applies not only to the proximal pinnae but also to distal pinnae and is rather striking compared to the generally alternate pinnae (at least in the middle and distal parts of the blade) in most cyclosoroid genera. Species of
<taxonomicName id="4C1E4D63FFDC9B7E60527F28FE93FDCD" box="[121,308,609,630]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60527F28FE93FDCD" box="[121,308,609,630]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
also resemble some neotropical
<taxonomicName id="4C1E4D63FFDC9B7E62447F28FCB0FDCC" box="[623,791,609,631]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pelazoneuron" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B96AEAF2FFDC9B7E62447F28FD4DFDCD" box="[623,746,609,630]" italics="true" pageId="53" pageNumber="43">Pelazoneuron</emphasis>
spp.
</taxonomicName>
, just outside the large clade containing the former, but
<taxonomicName id="4C1E4D63FFDC9B7E60FA7FC8FEEAFD2D" box="[209,333,641,662]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pelazoneuron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60FA7FC8FEEAFD2D" box="[209,333,641,662]" italics="true" pageId="53" pageNumber="43">Pelazoneuron</emphasis>
</taxonomicName>
has medial sori, more substantial indusia, more lanceolate, spreading, hairy stipe base scales, alternate pinnae above the blade base, and cristate or echinulate spores, rather than the reticulate spores of
<taxonomicName id="4C1E4D63FFDC9B7E60527F88FDA5FD6C" authority="(Patel et al. 2019 a)" baseAuthorityName="Patel" baseAuthorityYear="2019" box="[121,514,705,727]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60527F88FE9AFD6D" box="[121,317,705,726]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
(
<bibRefCitation id="EF8F4B11FFDC9B7E61617F8BFE5CFD6C" author="Patel" box="[330,507,706,727]" etAl="et al." firstAuthor="Patel" journalOrPublisher="Int. J. Pl. Sci." pageId="53" pageNumber="43" pagination="1016 - 1035" part="180" refId="ref112729" refString="PATEL, N., S. FAWCETT, M. SUNDUE, &amp; J. BUDKE. 2019 a. Evolution of perine morphology in Thelypteridaceae. Int. J. Pl. Sci. 180: 1016 - 1035." title="Evolution of perine morphology in Thelypteridaceae" type="journal article" year="2019" yearSuffix="a">Patel et al. 2019a</bibRefCitation>
)
</taxonomicName>
. The superficial resemblance of
<taxonomicName id="4C1E4D63FFDC9B7E63617F88FBA9FD6D" box="[842,1038,705,726]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E63617F88FBA9FD6D" box="[842,1038,705,726]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
to certain species of
<taxonomicName id="4C1E4D63FFDC9B7E60527FA8FF45FD4D" box="[121,226,737,758]" class="Polypodiopsida" family="Thelypteridaceae" genus="Steiropteris" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60527FA8FF45FD4D" box="[121,226,737,758]" italics="true" pageId="53" pageNumber="43">Steiropteris</emphasis>
</taxonomicName>
(the subgroup
<emphasis id="B96AEAF2FFDC9B7E615E7FA8FE59FD4D" box="[373,510,737,758]" italics="true" pageId="53" pageNumber="43">Glaphyropteris</emphasis>
), particularly
<taxonomicName id="4C1E4D63FFDC9B7E62AC7FA8FD50FD4D" box="[647,759,737,758]" class="Polypodiopsida" family="Thelypteridaceae" genus="Steiropteris" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="decussata">
<emphasis id="B96AEAF2FFDC9B7E62AC7FA8FD50FD4D" box="[647,759,737,758]" italics="true" pageId="53" pageNumber="43">S. decussata</emphasis>
</taxonomicName>
and allies (
<bibRefCitation id="EF8F4B11FFDC9B7E634E7FABFC7FFD4C" author="Smith" box="[869,984,738,759]" firstAuthor="Smith" journalOrPublisher="Univ. Calif. Publ. Bot." pageId="53" pageNumber="43" pagination="1 - 38" part="76" refId="ref114270" refString="SMITH, A. R. 1980. Taxonomy of Thelypteris subgenus Steiropteris, including Glaphyropteris (Pteridophyta). Univ. Calif. Publ. Bot. 76: 1 - 38." title="Taxonomy of Thelypteris subgenus Steiropteris, including Glaphyropteris (Pteridophyta)" type="journal article" year="1980">
<collectingRegion id="49DAF802FFDC9B7E634E7FABFC07FD4C" box="[869,928,738,759]" country="Bermuda" name="Smith's" pageId="53" pageNumber="43">Smith</collectingRegion>
1980
</bibRefCitation>
), is not indicative of a close phylogenetic relationship; that species and relatives differ in having pronounced elongate aerophores, mucilaginous croziers, non-falcate segments with rounded apices, resinous laminar glands, and broadly winged spores.
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDC9B7E60827E08FB23FC4C" pageId="53" pageNumber="43" type="biology_ecology">
<paragraph id="8BA136E0FFDC9B7E60827E08FB23FC4C" blockId="53.[121,1250,162,1591]" pageId="53" pageNumber="43">
<emphasis id="B96AEAF2FFDC9B7E60827E08FDD7FCED" box="[169,624,833,854]" italics="true" pageId="53" pageNumber="43">
Biogeography and ecology.—
<taxonomicName id="4C1E4D63FFDC9B7E619F7E08FDD7FCED" box="[436,624,833,854]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">Glaphyropteridopsis</taxonomicName>
</emphasis>
is centered in, and nearly restricted to, South
<collectingCountry id="F3097670FFDC9B7E641A7E0BFBC8FCEC" box="[1073,1135,834,855]" name="China" pageId="53" pageNumber="43">China</collectingCountry>
, with all 11 known species occurring there, ten of them endemic, and several known only from the
<typeStatus id="54A58842FFDC9B7E63E37E2BFC5AFCCC" box="[968,1021,866,887]" pageId="53" pageNumber="43">types</typeStatus>
or very few collections. The only species ranging outside of
<collectingCountry id="F3097670FFDC9B7E61FF7ECBFDB7FC2C" box="[468,528,898,919]" name="China" pageId="53" pageNumber="43">China</collectingCountry>
is the
<typeStatus id="54A58842FFDC9B7E627A7ECBFDD9FC2C" box="[593,638,898,919]" pageId="53" pageNumber="43">type</typeStatus>
,
<taxonomicName id="4C1E4D63FFDC9B7E62AC7EC8FCA1FC2D" box="[647,774,897,918]" class="Polypodiopsida" family="Thelypteridaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDC9B7E62AC7EC8FCA1FC2D" box="[647,774,897,918]" italics="true" pageId="53" pageNumber="43">G. erubescens</emphasis>
</taxonomicName>
, whose distribution extends into northern
<collectingCountry id="F3097670FFDC9B7E64837ECBFB7AFC2C" box="[1192,1245,898,919]" name="India" pageId="53" pageNumber="43">India</collectingCountry>
,
<collectingCountry id="F3097670FFDC9B7E60527EEBFF13FC0C" box="[121,180,930,951]" name="Nepal" pageId="53" pageNumber="43">Nepal</collectingCountry>
,
<collectingCountry id="F3097670FFDC9B7E60977EEBFEA3FC0C" box="[188,260,930,951]" name="Bhutan" pageId="53" pageNumber="43">Bhutan</collectingCountry>
, northern
<collectingCountry id="F3097670FFDC9B7E61437EEBFE60FC0C" box="[360,455,930,951]" name="Myanmar" pageId="53" pageNumber="43">Myanmar</collectingCountry>
,
<collectingCountry id="F3097670FFDC9B7E61E47EEBFD83FC0C" box="[463,548,930,951]" name="Vietnam" pageId="53" pageNumber="43">Vietnam</collectingCountry>
,
<collectingCountry id="F3097670FFDC9B7E62067EEBFDD1FC0C" box="[557,630,930,951]" name="Taiwan" pageId="53" pageNumber="43">Taiwan</collectingCountry>
, and in
<collectingCountry id="F3097670FFDC9B7E62EB7EEBFCADFC0C" box="[704,778,930,951]" name="Malaysia" pageId="53" pageNumber="43">Malesia</collectingCountry>
only in the
<collectingCountry id="F3097670FFDC9B7E63507EEBFC4CFC0C" box="[891,1003,930,951]" name="Philippines" pageId="53" pageNumber="43">Philippines</collectingCountry>
. The genus is absent from the rest of
<collectingCountry id="F3097670FFDC9B7E60F67E8BFE8EFC6C" box="[221,297,962,983]" name="Malaysia" pageId="53" pageNumber="43">Malesia</collectingCountry>
, Melanesia, and Polynesia, as well as Australasia and Africa and the New World. Species occur from
<quantity id="4CE69B05FFDC9B7E60527EABFF51FC4C" box="[121,246,994,1015]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="2.0" metricValueMin="0.6" pageId="53" pageNumber="43" unit="m" value="1300.0" valueMax="2000.0" valueMin="600.0">6002000 m</quantity>
. They typically occur in forests or at forest margins, along streams, on rocks, or along roadsides.
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDC9B7E6082794BFDD7FA0C" pageId="53" pageNumber="43" type="discussion">
<paragraph id="8BA136E0FFDC9B7E6082794BFBC4FB4C" blockId="53.[121,1250,162,1591]" pageId="53" pageNumber="43">
<emphasis id="B96AEAF2FFDC9B7E6082794BFE4FFBAD" box="[169,488,1025,1047]" italics="true" pageId="53" pageNumber="43">Taxonomic and phylogenetic study.</emphasis>
—Datasets based primarily on cpDNA sequences (
<bibRefCitation id="EF8F4B11FFDC9B7E63E3794BFBDFFBAC" author="He &amp; Zhang" box="[968,1144,1026,1047]" firstAuthor="He" journalOrPublisher="Molec. Phylogen. Evol." pageId="53" pageNumber="43" pagination="757 - 764" part="65" refId="ref109255" refString="HE, L. J. &amp; X. C. ZHANG. 2012. Exploring generic delimitation within the fern family Thelypteridaceae. Molec. Phylogen. Evol. 65: 757 - 764." title="Exploring generic delimitation within the fern family Thelypteridaceae" type="journal article" year="2012">He &amp; Zhang 2012</bibRefCitation>
;
<bibRefCitation id="EF8F4B11FFDC9B7E64AB794BFF14FB8C" author="Patel" etAl="et al." firstAuthor="Patel" journalOrPublisher="Int. J. Pl. Sci." pageId="53" pageNumber="43" pagination="1016 - 1035" part="180" refId="ref112729" refString="PATEL, N., S. FAWCETT, M. SUNDUE, &amp; J. BUDKE. 2019 a. Evolution of perine morphology in Thelypteridaceae. Int. J. Pl. Sci. 180: 1016 - 1035." title="Evolution of perine morphology in Thelypteridaceae" type="journal article" year="2019" yearSuffix="a">Patel et al. 2019a</bibRefCitation>
) and coalescent nuclear phylogenomic analyses (Fawcett et al. in press) infer
<taxonomicName id="4C1E4D63FFDC9B7E638D7968FBC6FB8D" box="[934,1121,1057,1078]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E638D7968FBC6FB8D" box="[934,1121,1057,1078]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
as sister to a large clade of strictly paleotropical cyclosoroids—including
<taxonomicName id="4C1E4D63FFDC9B7E62877908FC90FBED" box="[684,823,1089,1110]" class="Polypodiopsida" family="Thelypteridaceae" genus="Amblovenatum" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E62877908FC90FBED" box="[684,823,1089,1110]" italics="true" pageId="53" pageNumber="43">Amblovenatum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E636B7908FC2EFBED" box="[832,905,1089,1110]" class="Polypodiopsida" family="Thelypteridaceae" genus="Chingia" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E636B7908FC2EFBED" box="[832,905,1089,1110]" italics="true" pageId="53" pageNumber="43">Chingia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E63B97908FC4BFBED" box="[914,1004,1089,1110]" class="Polypodiopsida" family="Thelypteridaceae" genus="Christella" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E63B97908FC4BFBED" box="[914,1004,1089,1110]" italics="true" pageId="53" pageNumber="43">Christella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E63DE7908FBFAFBED" box="[1013,1117,1089,1110]" class="Polypodiopsida" family="Thelypteridaceae" genus="Grypothrix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E63DE7908FBFAFBED" box="[1013,1117,1089,1110]" italics="true" pageId="53" pageNumber="43">Grypothrix</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E644C790BFB7CFBEC" box="[1127,1243,1090,1111]" class="Polypodiopsida" family="Thelypteridaceae" genus="Menisciopsis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E644C790BFB7CFBEC" box="[1127,1243,1090,1111]" italics="true" pageId="53" pageNumber="43">Menisciopsis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E60527928FF53FBCD" box="[121,244,1121,1142]" class="Polypodiopsida" family="Thelypteridaceae" genus="Plesioneuron" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60527928FF53FBCD" box="[121,244,1121,1142]" italics="true" pageId="53" pageNumber="43">Plesioneuron</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E6129792BFE3DFBCC" authorityName="sensu Holttum. In" authorityYear="1982" box="[258,410,1122,1143]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E6129792BFE3DFBCC" box="[258,410,1122,1143]" italics="true" pageId="53" pageNumber="43">Pneumatopteris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E61837928FD81FBCD" authorityName="sensu Holttum. These" authorityYear="1982" box="[424,550,1121,1142]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pronephrium" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E61837928FD81FBCD" box="[424,550,1121,1142]" italics="true" pageId="53" pageNumber="43">Pronephrium</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E621F7928FD7BFBCD" box="[564,732,1121,1142]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E621F7928FD7BFBCD" box="[564,732,1121,1142]" italics="true" pageId="53" pageNumber="43">Pseudocyclosorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E62C27928FCF9FBCD" box="[745,862,1121,1142]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E62C27928FCF9FBCD" box="[745,862,1121,1142]" italics="true" pageId="53" pageNumber="43">Reholttumia</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1E4D63FFDC9B7E63B17928FBE0FBCD" baseAuthorityName="Holttum" baseAuthorityYear="1982" box="[922,1095,1121,1142]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E63B17928FBE0FBCD" box="[922,1095,1121,1142]" italics="true" pageId="53" pageNumber="43">Sphaerostephanos</emphasis>
</taxonomicName>
, but excluding
<taxonomicName id="4C1E4D63FFDC9B7E605279C8FF7BFB2D" box="[121,220,1153,1174]" class="Polypodiopsida" family="Thelypteridaceae" genus="Cyclosorus" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus" sensu="stricto">
<emphasis id="B96AEAF2FFDC9B7E605279C8FF7BFB2D" box="[121,220,1153,1174]" italics="true" pageId="53" pageNumber="43">Cyclosorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595789FFDC9B7E60CB79CBFF5CFB2C" box="[224,251,1154,1175]" pageId="53" pageNumber="43" sensu="stricto">
<emphasis id="B96AEAF2FFDC9B7E60CB79CBFF52FB2C" box="[224,245,1154,1175]" italics="true" pageId="53" pageNumber="43">s.s</emphasis>
.
</taxonomicNameLabel>
and also excluding all neotropical cyclosoroid genera, except
<emphasis id="B96AEAF2FFDC9B7E636B79C8FC06FB2D" box="[832,929,1153,1174]" italics="true" pageId="53" pageNumber="43">
<taxonomicName id="4C1E4D63FFDC9B7E636B79C8FC39FB2D" box="[832,926,1153,1174]" class="Polypodiopsida" family="Thelypteridaceae" genus="Christella" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">Christella</taxonomicName>
.
</emphasis>
However, a concatenated analysis of the same nuclear dataset (Fawcett et al. in press) places
<taxonomicName id="4C1E4D63FFDC9B7E628D79E8FCC6FB0D" box="[678,865,1185,1206]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E628D79E8FCC6FB0D" box="[678,865,1185,1206]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
within the chingioid subclade (
<taxonomicName id="4C1E4D63FFDC9B7E64B979E8FB7CFB0D" box="[1170,1243,1185,1206]" class="Polypodiopsida" family="Thelypteridaceae" genus="Chingia" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E64B979E8FB7CFB0D" box="[1170,1243,1185,1206]" italics="true" pageId="53" pageNumber="43">Chingia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E60527988FF44FB6D" box="[121,227,1217,1238]" class="Polypodiopsida" family="Thelypteridaceae" genus="Grypothrix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60527988FF44FB6D" box="[121,227,1217,1238]" italics="true" pageId="53" pageNumber="43">Grypothrix</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D63FFDC9B7E60C4798BFEC3FB6C" box="[239,356,1218,1239]" class="Polypodiopsida" family="Thelypteridaceae" genus="Menisciopsis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E60C4798BFEC3FB6C" box="[239,356,1218,1239]" italics="true" pageId="53" pageNumber="43">Menisciopsis</emphasis>
</taxonomicName>
, and
<emphasis id="B96AEAF2FFDC9B7E61B77988FDBEFB6D" box="[412,537,1217,1238]" italics="true" pageId="53" pageNumber="43">
<taxonomicName id="4C1E4D63FFDC9B7E61B77988FDB4FB6D" box="[412,531,1217,1238]" class="Polypodiopsida" family="Thelypteridaceae" genus="Plesioneuron" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">Plesioneuron</taxonomicName>
;
</emphasis>
<figureCitation id="13252A65FFDC9B7E6234798BFDF2FB6C" box="[543,597,1218,1239]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="53" pageNumber="43">Fig. 1</figureCitation>
). Low or conflicting support along backbone nodes in plastid and nuclear analyses suggest that the precise phylogenetic position of this lineage be interpreted with caution.
</paragraph>
<paragraph id="8BA136E0FFDC9B7E6082784BFB32FACC" blockId="53.[121,1250,162,1591]" pageId="53" pageNumber="43">
<emphasis id="B96AEAF2FFDC9B7E6082784BFF7AFAAC" box="[169,221,1282,1303]" italics="true" pageId="53" pageNumber="43">Notes</emphasis>
.—
<bibRefCitation id="EF8F4B11FFDC9B7E60D2784BFEDAFAAC" author="Ching" box="[249,381,1282,1303]" firstAuthor="Ching" journalOrPublisher="Acta Phytotax. Sin." pageId="53" pageNumber="43" pagination="289 - 335" part="8" refId="ref107523" refString="CHING, R. C. 1963. A reclassification of the family Thelypteridaceae from the mainland of Asia. Acta Phytotax. Sin. 8: 289 - 335." title="A reclassification of the family Thelypteridaceae from the mainland of Asia" type="journal article" year="1963">Ching (1963)</bibRefCitation>
, when describing the genus, surmised that his
<taxonomicName id="4C1E4D63FFDC9B7E63657848FC63FAAC" box="[846,964,1281,1303]" class="Polypodiopsida" family="Thelypteridaceae" genus="Grypothrix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="eriocarpa">
<emphasis id="B96AEAF2FFDC9B7E63657848FC63FAAC" box="[846,964,1281,1303]" italics="true" pageId="53" pageNumber="43">G. eriocarpa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E4D63FFDC9B7E63DD7848FBCAFAAD" box="[1014,1133,1281,1302]" class="Polypodiopsida" family="Thelypteridaceae" genus="Grypothrix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="splendens">
<emphasis id="B96AEAF2FFDC9B7E63DD7848FBCAFAAD" box="[1014,1133,1281,1302]" italics="true" pageId="53" pageNumber="43">G. splendens</emphasis>
</taxonomicName>
could be of hybrid origin, since they were rare and came from the same locality as
<taxonomicName id="4C1E4D63FFDC9B7E63387868FC35FA8D" box="[787,914,1313,1334]" class="Polypodiopsida" family="Thelypteridaceae" genus="Polypodium" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDC9B7E63387868FC35FA8D" box="[787,914,1313,1334]" italics="true" pageId="53" pageNumber="43">G. erubescens</emphasis>
</taxonomicName>
and
<emphasis id="B96AEAF2FFDC9B7E63947868FBC7FA8D" box="[959,1120,1313,1334]" italics="true" pageId="53" pageNumber="43">
<taxonomicName id="4C1E4D63FFDC9B7E63947868FBFAFA8D" box="[959,1117,1313,1334]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="rufostraminea">G. rufostraminea</taxonomicName>
.
</emphasis>
This hypothesis has not been tested.
<bibRefCitation id="EF8F4B11FFDC9B7E6148780BFD84FAEC" author="Shieh &amp; Tsai" box="[355,547,1346,1367]" firstAuthor="Shieh" journalOrPublisher="J. Sci. Engin. (Natl. Chung-Hsing Univ.)" pageId="53" pageNumber="43" pagination="1 - 12" part="24" refId="ref114027" refString="SHIEH, W. C. &amp; J. L. TSAI. 1987. Three natural hybrids of the Thelypteridaceae in Taiwan. J. Sci. Engin. (Natl. Chung-Hsing Univ.) 24: 1 - 12." title="Three natural hybrids of the Thelypteridaceae in Taiwan" type="journal article" year="1987">Shieh &amp; Tsai (1987)</bibRefCitation>
described
<taxonomicName id="4C1E4D63FFDC9B7E62A17808FC29FAED" box="[650,910,1345,1366]" class="Polypodiopsida" family="Thelypteridaceae" genus="Thelypteris" isHybrid="true" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="erubesquirolica">
<emphasis id="B96AEAF2FFDC9B7E62A17808FD55FAED" box="[650,754,1345,1366]" italics="true" pageId="53" pageNumber="43">Thelypteris</emphasis>
×
<emphasis id="B96AEAF2FFDC9B7E63297808FC29FAED" box="[770,910,1345,1366]" italics="true" pageId="53" pageNumber="43">erubesquirolica</emphasis>
</taxonomicName>
, which they believed to be a hybrid between
<taxonomicName id="4C1E4D63FFDC9B7E60E57828FE56FACD" box="[206,497,1377,1398]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDC9B7E60E57828FE56FACD" box="[206,497,1377,1398]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis erubescens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E4D63FFDC9B7E62357828FCB2FACD" box="[542,789,1377,1398]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pseudocyclosorus" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="esquirolii">
<emphasis id="B96AEAF2FFDC9B7E62357828FCB2FACD" box="[542,789,1377,1398]" italics="true" pageId="53" pageNumber="43">Pseudocyclosorus esquirolii</emphasis>
</taxonomicName>
. This hypothesis needs re-examination.
</paragraph>
<paragraph id="8BA136E0FFDC9B7E608278CBFDD7FA0C" blockId="53.[121,1250,162,1591]" pageId="53" pageNumber="43">
About half of the known species are illustrated by
<bibRefCitation id="EF8F4B11FFDC9B7E62B578CBFCA0FA2C" author="Lin" box="[670,775,1410,1431]" editor="Shing, K." firstAuthor="Lin" journalOrPublisher="Science Press, Beijing, China. Pp" pageId="53" pageNumber="43" pagination="113 - 317" refId="ref111203" refString="LIN, Y. X. 1999. Thelypteridaceae. In Shing, K., ed., Flora Reipublicae Popularis Sinicae. Science Press, Beijing, China. Pp. 113 - 317." title="Thelypteridaceae" type="book chapter" volumeTitle="Flora Reipublicae Popularis Sinicae" year="1999">Lin (1999)</bibRefCitation>
. All necessary combinations have been made in
<taxonomicName id="4C1E4D63FFDC9B7E605278E8FE94FA0D" box="[121,307,1441,1462]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAF2FFDC9B7E605278E8FE94FA0D" box="[121,307,1441,1462]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis</emphasis>
</taxonomicName>
, by
<bibRefCitation id="EF8F4B11FFDC9B7E617278EBFE7EFA0C" author="Ching" box="[345,473,1442,1463]" firstAuthor="Ching" journalOrPublisher="Acta Phytotax. Sin." pageId="53" pageNumber="43" pagination="289 - 335" part="8" refId="ref107523" refString="CHING, R. C. 1963. A reclassification of the family Thelypteridaceae from the mainland of Asia. Acta Phytotax. Sin. 8: 289 - 335." title="A reclassification of the family Thelypteridaceae from the mainland of Asia" type="journal article" year="1963">Ching (1963)</bibRefCitation>
and
<bibRefCitation id="EF8F4B11FFDC9B7E622D78EBFDCBFA0C" author="Lin" box="[518,620,1442,1463]" editor="Shing, K." firstAuthor="Lin" journalOrPublisher="Science Press, Beijing, China. Pp" pageId="53" pageNumber="43" pagination="113 - 317" refId="ref111203" refString="LIN, Y. X. 1999. Thelypteridaceae. In Shing, K., ed., Flora Reipublicae Popularis Sinicae. Science Press, Beijing, China. Pp. 113 - 317." title="Thelypteridaceae" type="book chapter" volumeTitle="Flora Reipublicae Popularis Sinicae" year="1999">Lin (1999)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304656BFFDC9B7E60827888FE06F98C" pageId="53" pageNumber="43" type="taxon_list">
<paragraph id="8BA136E0FFDC9B7E60827888FE06F98C" blockId="53.[121,1250,162,1591]" pageId="53" pageNumber="43">
<emphasis id="B96AEAF2FFDC9B7E60827888FEFFFA6C" box="[169,344,1473,1495]" italics="true" pageId="53" pageNumber="43">Constituent species</emphasis>
.—
<taxonomicName id="4C1E4D63FFDC9B7E615D7888FD42FA6C" authority="Y. X. Lin" authorityName="Y. X. Lin" box="[374,741,1473,1495]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="emeiensis">
<emphasis id="B96AEAF2FFDC9B7E615D7888FD28FA6C" box="[374,655,1473,1495]" italics="true" pageId="53" pageNumber="43">Glaphyropteridopsis emeiensis</emphasis>
Y.X. Lin
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E62C57888FC01FA6C" authority="Ching" authorityName="Ching" box="[750,934,1473,1495]" class="Polypodiopsida" family="Thelypteridaceae" genus="Grypothrix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="eriocarpa">
<emphasis id="B96AEAF2FFDC9B7E62C57888FCC4FA6C" box="[750,867,1473,1495]" italics="true" pageId="53" pageNumber="43">G. eriocarpa</emphasis>
Ching
</taxonomicName>
;
<emphasis id="B96AEAF2FFDC9B7E639B7888FC1DFA6D" box="[944,954,1473,1494]" italics="true" pageId="53" pageNumber="43">*</emphasis>
<taxonomicName id="4C1E4D63FFDC9B7E63917888FF10FA4C" authority="(Wall. ex Hook.) Ching" authorityName="Ching" baseAuthorityName="Hook." class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="erubescens">
<emphasis id="B96AEAF2FFDC9B7E63917888FB9DFA6D" box="[954,1082,1473,1494]" italics="true" pageId="53" pageNumber="43">G. erubescens</emphasis>
(Wall. ex Hook.) Ching
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E60EB78A8FDFBFA4C" authority="Ching &amp; W. M. Chu ex Y. X. Lin" authorityName="Y. X. Lin" box="[192,604,1505,1527]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="glabrata">
<emphasis id="B96AEAF2FFDC9B7E60EB78A8FE8FFA4D" box="[192,296,1505,1526]" italics="true" pageId="53" pageNumber="43">G. glabrata</emphasis>
Ching &amp; W.M. Chu ex Y.X. Lin
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E624E78A8FC13FA4C" authority="Ching ex Y. X. Lin" authorityName="Y. X. Lin" box="[613,948,1505,1527]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="jinfushanensis">
<emphasis id="B96AEAF2FFDC9B7E624E78A8FCA4FA4D" box="[613,771,1505,1526]" italics="true" pageId="53" pageNumber="43">G. jinfushanensis</emphasis>
Ching ex Y.X. Lin
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E639578A8FB66FA4C" authority="Ching ex Y. X. Lin" authorityName="Y. X. Lin" box="[958,1217,1505,1527]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="B96AEAF2FFDC9B7E639578A8FBA8FA4D" box="[958,1039,1505,1526]" italics="true" pageId="53" pageNumber="43">G. mollis</emphasis>
Ching ex Y.X. Lin
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E64E078A8FED4F9AC" authority="Ching ex Y. X. Lin" authorityName="Y. X. Lin" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="pallida">
<emphasis id="B96AEAF2FFDC9B7E64E078A8FF1DF9AD" italics="true" pageId="53" pageNumber="43">G. pallida</emphasis>
Ching ex Y.X. Lin
</taxonomicName>
; *
<taxonomicName id="4C1E4D63FFDC9B7E61A37B48FD66F9AC" authority="(Christ) Ching" authorityName="Ching" baseAuthorityName="Christ" box="[392,705,1537,1559]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="rufostraminea">
<emphasis id="B96AEAF2FFDC9B7E61A37B48FD80F9AD" box="[392,551,1537,1558]" italics="true" pageId="53" pageNumber="43">G. rufostraminea</emphasis>
(Christ) Ching
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E62E77B48FC11F9AC" authority="Y. X. Lin" authorityName="Y. X. Lin" box="[716,950,1537,1559]" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="sichuanensis">
<emphasis id="B96AEAF2FFDC9B7E62E77B48FCF9F9AD" box="[716,862,1537,1558]" italics="true" pageId="53" pageNumber="43">G. sichuanensis</emphasis>
Y.X. Lin
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E63EA7B48FBD9F9AC" authority="Ching" authorityName="Ching" box="[961,1150,1537,1559]" class="Polypodiopsida" family="Thelypteridaceae" genus="Grypothrix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="splendens">
<emphasis id="B96AEAF2FFDC9B7E63EA7B48FB9FF9AD" box="[961,1080,1537,1558]" italics="true" pageId="53" pageNumber="43">G. splendens</emphasis>
Ching
</taxonomicName>
;
<taxonomicName id="4C1E4D63FFDC9B7E64A37B48FE06F98C" authority="Ching &amp; W. M. Chu ex Y. X. Lin" class="Polypodiopsida" family="Thelypteridaceae" genus="Glaphyropteridopsis" kingdom="Plantae" order="Polypodiales" pageId="53" pageNumber="43" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="B96AEAF2FFDC9B7E64A37B48FB46F9AD" box="[1160,1249,1537,1558]" italics="true" pageId="53" pageNumber="43">G. villosa</emphasis>
Ching &amp; W.M. Chu ex Y.X. Lin
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>