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<document id="0CB4250ADA8E849107F3C5AE332CA275" ID-CLB-Dataset="306451" ID-DOI="10.11646/zootaxa.5551.1.7" ID-GBIF-Dataset="1c81b294-3149-4e1d-b6e1-83315506956f" ID-ISSN="1175-5326" ID-Zenodo-Dep="14390215" ID-ZooBank="43517112-40DA-4E69-8BFB-BEC55AC5AAF2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733924615010" checkinUser="plazi" docAuthor="Delgado-Blas, Vi ́ Ctor Hugo, Díaz-Díaz, Oscar &amp; Rozbaczylo, Nicolás" docDate="2024" docId="0381CB35A128FF8DFF50FF0FFC70A63F" docLanguage="en" docName="zootaxa.5551.1.7.pdf" docOrigin="Zootaxa 5551 (1)" docSource="https://doi.org/10.11646/zootaxa.5551.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.20:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="20" docTitle="Laonice ayseniensis Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo 2024" docType="treatment" docVersion="4" lastPageNumber="165" masterDocId="FFB8B34DA12DFF85FFC7FF98FFABA45E" masterDocTitle="Laonice Malmgren, 1867 (Annelida: Spionidae) from the Aysén region in southern Chile with description of a new species" masterLastPageNumber="166" masterPageNumber="157" pageNumber="162" updateTime="1734091358508" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E9124B64C28940184E84DBD4F04CFA98">Laonice Malmgren, 1867 (Annelida: Spionidae) from the Aysén region in southern Chile with description of a new species</mods:title>
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<mods:namePart id="F7516EDF361A0989464E7D3673718000">Delgado-Blas, Vi ́ Ctor Hugo</mods:namePart>
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<mods:affiliation id="6AD0C1FAACEE75757EA18DFB0F978A0A">Universidad Auto ́ noma de Quintana Roo, Divisio ́ n de Ciencias, Ingenieri ́ a y Tecnologia, Chetumal, Quintana Roo, 77019, Me ́ xico</mods:affiliation>
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<taxonomicName id="4C2801A0A128FF80FF50FF0FFE22A4EF" authority="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo, 2024" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[151,393,151,177]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="5" pageNumber="162" phylum="Annelida" rank="species" species="ayseniensis">
<emphasis id="B95CA631A128FF80FF50FF0FFE22A4EF" bold="true" box="[151,393,151,177]" italics="true" pageId="5" pageNumber="162">Laonice ayseniensis</emphasis>
</taxonomicName>
n. sp
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</paragraph>
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<figureCitation id="131366A6A128FF80FF50FF23FE87A48B" box="[151,300,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Figure 2AQ</figureCitation>
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<paragraph id="8B977A23A128FF80FF50FE9BFD7BA5D7" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
<emphasis id="B95CA631A128FF80FF50FE9BFE95A543" bold="true" box="[151,318,259,285]" pageId="5" pageNumber="162">Type material</emphasis>
.
<materialsCitation id="3B40707EA128FF80FE94FE9BFC0AA51C" collectedFrom="slightly gravelly sand" collectingDate="2013-12-10" collectionCode="MNHNCL" collectorName="S. Martinez" county="Pichicolo" elevation="30" location="Hualaihue" municipality="Palena" pageId="5" pageNumber="162" specimenCode="MNHNCL ANN-15311" specimenCount="1" stateProvince="Los Lagos region" typeStatus="holotype">
<collectingCounty id="62F602AFA128FF80FE94FE9BFE6BA543" box="[339,448,259,285]" pageId="5" pageNumber="162">Pichicolo</collectingCounty>
,
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,
<location id="8EF72CF8A128FF80FDE4FE9BFD32A543" LSID="urn:lsid:plazi:treatment:0381CB35A128FF8DFF50FF0FFC70A63F:8EF72CF8A128FF80FDE4FE9BFD32A543" box="[547,665,259,285]" county="Pichicolo" latitude="-44.316666" longLatPrecision="1137" longitude="-72.55" municipality="Palena" name="Hualaihue" pageId="5" pageNumber="162" stateProvince="Los Lagos region">Hualaihue</location>
,
<collectingRegion id="49ECB4C1A128FF80FD63FE9CFCC2A543" box="[676,873,259,286]" country="Chile" name="Los Lagos" pageId="5" pageNumber="162">Los Lagos region</collectingRegion>
<date id="FF965CE3A128FF80FCA8FE9BFC59A543" box="[879,1010,259,285]" pageId="5" pageNumber="162" value="2013-12-10">
<collectingDate id="EFD2A50BA128FF80FCA8FE9BFC59A543" box="[879,1010,259,285]" pageId="5" pageNumber="162" value="2013-12-10">10/12/2013</collectingDate>
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,
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,
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, coll.
<collectorName id="26DD1FF5A128FF80FAD4FE9BFA33A543" box="[1299,1432,259,285]" pageId="5" pageNumber="162">S. Martínez</collectorName>
,
<collectedFrom id="383439DEA128FF80FF50FEBFFE21A51F" box="[151,394,295,321]" pageId="5" pageNumber="162">slightly gravelly sand</collectedFrom>
,
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depth
</elevation>
,
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(
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)
</materialsCitation>
and
<materialsCitation id="3B40707EA128FF80FC11FEBFFCDAA53B" collectionCode="MNHNCL" elevation="30" latitude="-44.316666" location="Canal Puyuhuapi" longLatPrecision="1137" longitude="-72.55" pageId="5" pageNumber="162" specimenCode="MNHNCL ANN-15395" specimenCount="1" stateProvince="Aysen Region" typeStatus="paratype">
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<typeStatus id="5493C481A128FF80FBCFFEB0FBD8A51C" box="[1032,1139,296,322]" pageId="5" pageNumber="162">paratypes</typeStatus>
(
<specimenCode id="DB8ED258A128FF80FB46FEBFFA3AA51C" box="[1153,1425,295,322]" pageId="5" pageNumber="162">MNHNCL ANN-15395</specimenCode>
).
<location id="8EF72CF8A128FF80FF50FED3FEF2A53B" LSID="urn:lsid:plazi:treatment:0381CB35A128FF8DFF50FF0FFC70A63F:8EF72CF8A128FF80FF50FED3FEF2A53B" box="[151,345,331,357]" county="Pichicolo" latitude="-44.316666" longLatPrecision="1137" longitude="-72.55" municipality="Palena" name="Canal Puyuhuapi" pageId="5" pageNumber="162" stateProvince="Aysen Region">Canal Puyuhuapi</location>
,
<collectingRegion id="49ECB4C1A128FF80FEA6FED3FE56A53B" box="[353,509,331,357]" country="Chile" name="Aysen" pageId="5" pageNumber="162">Aysén Region</collectingRegion>
(
<geoCoordinate id="EE1C1CE4A128FF80FDCBFED3FDC0A53B" box="[524,619,331,357]" degrees="44" direction="south" minutes="19" orientation="latitude" pageId="5" pageNumber="162" precision="925" value="-44.316666">44°19S</geoCoordinate>
,
<geoCoordinate id="EE1C1CE4A128FF80FDB2FED3FD74A53B" box="[629,735,331,357]" degrees="72" direction="west" minutes="33" orientation="longitude" pageId="5" pageNumber="162" precision="925" value="-72.55">72°33`W</geoCoordinate>
),
<elevation id="00059D10A128FF80FD35FED3FCDAA53B" box="[754,881,331,358]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="162" unit="m" value="30.0">
<quantity id="4CD0D7C6A128FF80FD35FED3FC81A538" box="[754,810,331,358]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="162" unit="m" value="30.0">30 m</quantity>
depth
</elevation>
</materialsCitation>
,
<materialsCitation id="3B40707EA128FF80FCBCFED3FD67A5D7" collectedFrom="gravelly-muddy sediments" collectingDate="2014-03-09" collectionCode="MNHNCL" collectorName="C. Carreno" pageId="5" pageNumber="162" specimenCode="MNHNCL ANN-15396" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D2EB1AAA128FF80FCBCFED3FC18A53B" box="[891,947,331,357]" pageId="5" pageNumber="162" type="generic">eight</specimenCount>
<typeStatus id="5493C481A128FF80FC7FFED4FB88A538" box="[952,1059,332,358]" pageId="5" pageNumber="162">paratypes</typeStatus>
(
<specimenCode id="DB8ED258A128FF80FBF6FED3FAEBA538" box="[1073,1344,331,358]" pageId="5" pageNumber="162">MNHNCL ANN-15396</specimenCode>
),
<date id="FF965CE3A128FF80FA95FED3FF79A5D7" pageId="5" pageNumber="162" value="2014-03-09">
<collectingDate id="EFD2A50BA128FF80FA95FED3FF79A5D7" pageId="5" pageNumber="162" value="2014-03-09">09-03- 2014</collectingDate>
</date>
, coll.
<collectorName id="26DD1FF5A128FF80FED2FEF7FE38A5D7" box="[277,403,367,393]" pageId="5" pageNumber="162">C. Carreño</collectorName>
,
<collectedFrom id="383439DEA128FF80FE59FEF7FD67A5D7" box="[414,716,367,393]" pageId="5" pageNumber="162">gravelly-muddy sediments</collectedFrom>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A128FF82FF00FE0BFBD1A48B" lastPageId="7" lastPageNumber="164" pageId="5" pageNumber="162" type="description">
<paragraph id="8B977A23A128FF80FF00FE0BFEC7A5A8" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
<emphasis id="B95CA631A128FF80FF00FE0BFEFAA5F3" bold="true" box="[199,337,403,429]" pageId="5" pageNumber="162">Description</emphasis>
.
<typeStatus id="5493C481A128FF80FEA5FE0BFE61A5F3" box="[354,458,403,429]" pageId="5" pageNumber="162" type="holotype">Holotype</typeStatus>
complete: 147 chaetigers, length
<quantity id="4CD0D7C6A128FF80FC96FE0BFC08A5F0" box="[849,931,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="162" unit="mm" value="48.0">48 mm</quantity>
and width
<quantity id="4CD0D7C6A128FF80FBE0FE0BFB29A5F0" box="[1063,1154,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="162" unit="mm" value="1.6">1.6 mm</quantity>
;
<typeStatus id="5493C481A128FF80FB57FE0CFB5BA5F0" box="[1168,1264,404,430]" pageId="5" pageNumber="162" type="paratype">paratype</typeStatus>
complete: 139 chaetigers, length
<quantity id="4CD0D7C6A128FF80FEA0FE2FFE1CA58C" box="[359,439,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="162" unit="mm" value="38.0">38 mm</quantity>
and width
<quantity id="4CD0D7C6A128FF80FDFEFE2FFD38A58C" box="[569,659,439,466]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="162" unit="mm" value="1.9">1.9 mm</quantity>
; 12 incomplete
<typeStatus id="5493C481A128FF80FC8EFE20FC12A58C" box="[841,953,440,466]" pageId="5" pageNumber="162" type="paratype">paratypes</typeStatus>
: 59106 chaetigers, length
<quantity id="4CD0D7C6A128FF80FB36FE2FFAC7A58C" box="[1265,1388,439,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.6" metricValueMin="1.0" pageId="5" pageNumber="162" unit="mm" value="18.0" valueMax="26.0" valueMin="10.0">1026 mm</quantity>
and width
<quantity id="4CD0D7C6A128FF80FF19FE43FEC3A5A8" box="[222,360,475,502]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.5" metricValueMin="1.6" pageId="5" pageNumber="162" unit="mm" value="2.05" valueMax="2.5" valueMin="1.6">1.62.5 mm</quantity>
.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FE67FC20A6AF" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Prostomium and peristomium fused anteriorly, giving a bicornate, cap-shaped appearance (
<figureCitation id="131366A6A128FF80FB6CFE67FAA8A647" box="[1195,1283,511,537]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
); prostomium triangular in the unfused region of the prostomium and peristomium. Peristomium well developed and free from notopodial lamellae (
<figureCitation id="131366A6A128FF80FE48FDDFFE47A63F" box="[399,492,583,609]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2B</figureCitation>
). One pair of large brownish eyespots, thicker on the lateral region, giving the appearance of raccoon spots on the posterior part of the prostomium (
<figureCitation id="131366A6A128FF80FC6DFDF3FBAEA6DB" box="[938,1029,619,645]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
). Occipital tentacle short, thick, and positioned slightly beyond the notopodial lamellae. Caruncle well developed, extending together with the nuchal organs that start between the notopodium and the prostomium, and accompany the caruncle until chaetigers 1416 (
<quantity id="4CD0D7C6A128FF80FF67FD4FFF71A6AF" box="[160,218,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="5" pageNumber="162" unit="in" value="16.0">16 in</quantity>
<typeStatus id="5493C481A128FF80FF27FD4FFEEFA6AF" box="[224,324,727,753]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) (
<figureCitation id="131366A6A128FF80FE9CFD4FFE1EA6AF" box="[347,437,727,753]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
). Palps absent in all specimens examined.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FD63FC2DA7B3" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Branchiae starting from chaetiger 2, continuing up to chaetigers 3538 (chaetiger
<quantity id="4CD0D7C6A128FF80FB91FD63FB25A74B" box="[1110,1166,763,790]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="5" pageNumber="162" unit="in" value="35.0">35 in</quantity>
<typeStatus id="5493C481A128FF80FB53FD63FB51A74B" box="[1172,1274,763,789]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
), all branchiae basally free of the notopodial lamellae (
<figureCitation id="131366A6A128FF80FD94FC87FD4CA767" box="[595,743,799,826]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E, F, H</figureCitation>
). All branchiae digitiform, long, thick, with spear-shaped tips, and heavily ciliated; first two pairs of branchiae short (
<figureCitation id="131366A6A128FF80FCD6FCDBFC21A703" box="[785,906,835,861]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E, F</figureCitation>
); first pair slightly shorter than the notopodial postchaetal lamellae (
<figureCitation id="131366A6A128FF80FE41FCFFFE71A7DF" box="[390,474,871,897]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E</figureCitation>
), increasing in length significantly just after the third pair of notolamellae and becoming twice as long as the notopodial postchaetal lamellae from chaetiger 6 (
<figureCitation id="131366A6A128FF80FC64FC13FC57A7FB" box="[931,1020,907,933]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A</figureCitation>
). From chaetiger 10, the length of the longest branchiae equal to the total width of the body (
<figureCitation id="131366A6A128FF80FD36FC37FCC1A797" box="[753,874,943,969]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A, G</figureCitation>
), thereafter branchiae length gradually decreasing, the last three pairs so small that they are hidden in the parapodium.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FC6FFE44A16F" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Notopodial postchaetal lamellae of chaetiger 1 very small, subtriangular, shorter than neurolamellae and occipital nuchal tentacle (
<figureCitation id="131366A6A128FF80FE70FB83FDBAA06B" box="[439,529,1051,1077]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2D</figureCitation>
). From pairs 23, lamellae increasing in size to form a broad subtriangular shape (
<figureCitation id="131366A6A128FF80FF58FBA7FEB0A007" box="[159,283,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2E, F</figureCitation>
); fourth pair and subsequent pairs even larger and wider, leaf-shaped, the fourth pair a third larger than the third pair, with the tips of the notopodial lamellae touching; fourth pair the largest notopodial lamellae, notopodial lamellae equal in size on chaetigers 1314, thereafter gradually decreasing in size, becoming rounded from chaetiger 37; from around chaetiger 44 lamellae becoming triangular and gradually decreasing in size until the end of the body. Notopodial prechaetal lamellae rounded, moderately developed (
<figureCitation id="131366A6A128FF80FB92FB57FB5AA0B7" box="[1109,1265,1231,1257]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2A, EH</figureCitation>
), postbranchial lamellae reduced. Dorsal transverse ridges very low, connecting notopodial postchaetal lamellae on chaetigers 37 48 (
<figureCitation id="131366A6A128FF80FF05FA8FFEB0A16F" box="[194,283,1303,1329]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2C</figureCitation>
), absent thereafter.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FAA3FD8BA1BB" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Neuropodial postchaetal lamellae of chaetiger 1 rectangular with rounded edges (
<figureCitation id="131366A6A128FF80FB98FAA3FB12A10B" box="[1119,1209,1339,1365]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2D</figureCitation>
), oval on chaetigers 28, with well developed ventral edges in the form of an earlobe, (
<figureCitation id="131366A6A128FF80FCA8FAC7FC4CA127" box="[879,999,1375,1401]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2EG</figureCitation>
), increasing gradually in size. Lamellae on chaetiger 9 without ventral lobe (
<figureCitation id="131366A6A128FF80FD84FA1BFD25A1C3" box="[579,654,1411,1437]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig 2I</figureCitation>
), and on chaetiger 25 lamellae subtriangular, wide, small, until the appearance of the ventrolateral pouches when they become rounded, gradually decreasing in size on posterior chaetigers, with low, rounded lobe.
</paragraph>
<paragraph id="8B977A23A128FF80FF00FA77FD3BA20F" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Ventrolateral pouches starting from chaetigers 3548 (
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<typeStatus id="5493C481A128FF80FCB6FA77FC7EA257" box="[881,981,1519,1545]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) and continuing up to chaetiger 57. The first, very small, pouch emerges from the dorsal part of the prechaetal neuropodial lamellae and the posterior dorsal base of the postchaetal neuropodial lamellae.
</paragraph>
<paragraph id="8B977A23A128FF80FF00F9C3FA91A3CB" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
All anterior parapodia with capillaries arranged in two rows. Long, thin capillary chaetae in the upper part of the notopodium; notopodial chaetae from chaetiger 5 up to chaetiger 22 with chaetae arranged in two rows, anterior row with short, striate and bilimbate chaetae (
<figureCitation id="131366A6A128FF80FD5AF93BFD45A2E3" box="[669,750,1699,1725]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2J</figureCitation>
), posterior row with long, striate and bilimbate chaetae (
<figureCitation id="131366A6A128FF80FAA8F93BFF16A2BF" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2K</figureCitation>
); from chaetiger 23 capillary chaetae arranged in two rows, long, thin, striate unilimbate (
<figureCitation id="131366A6A128FF80FB61F95FFB56A2BF" box="[1190,1277,1735,1761]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2L</figureCitation>
). Neuropodial chaetae similar to notopodial chaetae, arranged in two rows, but shorter and thinner. Neuropodial hooks from chaetigers 3538, (
<quantity id="4CD0D7C6A128FF80FEADF897FE08A377" box="[362,419,1807,1834]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="5" pageNumber="162" unit="in" value="35.0">35 in</quantity>
<typeStatus id="5493C481A128FF80FE6EF897FDA6A377" box="[425,525,1807,1833]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) with 612 hooks per fascicle; hooded hooks tridentate with two teeth on the main fang, giving a bidentate appearance in the lateral view, hoods with distal opening (
<figureCitation id="131366A6A128FF80FBEEF8ABFB0FA313" box="[1065,1188,1843,1869]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2N, O</figureCitation>
), companion capillary chaetae present (
<figureCitation id="131366A6A128FF80FE95F8CFFE0CA32F" box="[338,423,1879,1905]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2P</figureCitation>
). Sabre chaetae from chaetigers 2428 (
<quantity id="4CD0D7C6A128FF80FCA2F8CFFC34A32F" box="[869,927,1879,1905]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="5" pageNumber="162" unit="in" value="28.0">28 in</quantity>
<typeStatus id="5493C481A128FF80FC62F8CFFBA2A32F" box="[933,1033,1879,1905]" pageId="5" pageNumber="162" type="holotype">holotype</typeStatus>
) numbering up to three per fascicle; sabre chaetae heavily striate and granulate (
<figureCitation id="131366A6A128FF80FD47F8E3FD74A3CB" box="[640,735,1915,1941]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2M</figureCitation>
), from chaetiger 15 numbering up to two per fascicle.
</paragraph>
<paragraph id="8B977A23A128FF80FF00F807FDC1A383" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="162">
Pygidium with two small ventral papilliform cirri arranged closely to each other, and five long cirri: two on one side and three on the other side (
<figureCitation id="131366A6A128FF80FDC4F85BFDF6A383" box="[515,605,1987,2013]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="5" pageNumber="162">Fig. 2Q</figureCitation>
).
</paragraph>
<caption id="DF572AABA12BFF83FF50F963FE4EA3B5" ID-DOI="http://doi.org/10.5281/zenodo.14390223" ID-Zenodo-Dep="14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="6" pageNumber="163" startId="6.[151,250,1787,1811]" targetBox="[250,1337,190,1752]" targetPageId="6" targetType="figure">
<paragraph id="8B977A23A12BFF83FF50F963FE4EA3B5" blockId="6.[151,1437,1787,2027]" pageId="6" pageNumber="163">
<emphasis id="B95CA631A12BFF83FF50F963FEBEA34D" bold="true" box="[151,277,1787,1811]" pageId="6" pageNumber="163">FIGURE 2.</emphasis>
<taxonomicName id="4C2801A0A12BFF83FEDAF964FE41A34D" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[285,490,1788,1811]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="6" pageNumber="163" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12BFF83FEDAF964FE41A34D" box="[285,490,1788,1811]" italics="true" pageId="6" pageNumber="163">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12BFF83FE34F963FDEDA34D" bold="true" box="[499,582,1787,1811]" pageId="6" pageNumber="163">
<taxonomicNameLabel id="A26F1B4AA12BFF83FE34F963FDEDA34D" box="[499,582,1787,1811]" pageId="6" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp).
</paragraph>
</caption>
<paragraph id="8B977A23A12AFF82FF00FF0EFBD1A48B" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FF0EFE5FA4EE" bold="true" box="[199,500,150,177]" pageId="7" pageNumber="164">Staining in Methyl Green</emphasis>
. Peristomium stained profusely dorsally and ventrally. Anterior segments with outer margin of both notopodial and neuropodial postchaetal lamellae stained (
<figureCitation id="131366A6A12AFF82FC0EFF23FB8AA48B" box="[969,1057,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1787,1811]" captionTargetBox="[250,1337,190,1752]" captionTargetId="figure-21@6.[239,1348,181,1763]" captionTargetPageId="6" captionText="FIGURE 2. Laonice ayseniensis sp. nov. paratype (MNHNCL ANN-15395): (A) anterior region, dorsal view; (A´) anterior region, lateral-ventral view; (B) anterior region, lateral view; (C) dorsal ribs; (D) parapodium, chaetiger 1; (E) parapodium, chaetiger 2; (F) parapodium, chaetiger 3; (G) notopodium, chaetiger 7; (H) parapodium, chaetiger 15; (I) parapodium, chaetiger 25; (J) capillary notochaeta, anterior row of anterior chaetiger; (K) capillary notochaeta, posterior row of anterior chaetiger; (L) capillary chaeta of chaetiger 23; (M) sabre chaeta; (N) neuropodial hooded hook in frontal view; (O) same in lateral view; (P) companion capillary; (Q) pygidium in lateral view. Abbreviations: dorsal ribs (Dr), lateral membrane (LM), occipital tentacle (Ot), ventrolateral pouches (Vp)." figureDoi="http://doi.org/10.5281/zenodo.14390223" httpUri="https://zenodo.org/record/14390223/files/figure.png" pageId="7" pageNumber="164">Fig. 2A</figureCitation>
, A´, B).
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A12AFF82FF00FF47FE91A543" pageId="7" pageNumber="164" type="etymology">
<paragraph id="8B977A23A12AFF82FF00FF47FE91A543" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FF47FEE5A4A7" bold="true" box="[199,334,223,249]" pageId="7" pageNumber="164">Etymology.</emphasis>
The specific name
<taxonomicName id="4C2801A0A12AFF82FDEBFF47FD07A4A6" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[556,684,223,248]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis">
<emphasis id="B95CA631A12AFF82FDEBFF47FD07A4A6" box="[556,684,223,248]" italics="true" pageId="7" pageNumber="164">ayseniensis</emphasis>
</taxonomicName>
is derived from the name of a geographical zone in the
<collectingRegion id="49ECB4C1A12AFF82FAE1FF47FF40A543" country="Chile" name="Los Lagos" pageId="7" pageNumber="164">Los Lagos Region</collectingRegion>
,
<collectingCountry id="F33F3AB3A12AFF82FF31FE9BFE9DA543" box="[246,310,259,285]" name="Chile" pageId="7" pageNumber="164">Chile</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A12AFF82FF00FEBEFD6AA51F" box="[199,705,294,321]" pageId="7" pageNumber="164" type="distribution">
<paragraph id="8B977A23A12AFF82FF00FEBEFD6AA51F" blockId="7.[151,1437,150,2013]" box="[199,705,294,321]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FEBEFEF2A51E" bold="true" box="[199,345,294,320]" pageId="7" pageNumber="164">Distribution</emphasis>
Only known from
<typeStatus id="5493C481A12AFF82FDF7FEBFFDCBA51F" box="[560,608,295,321]" pageId="7" pageNumber="164">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33229A8A12AFF8DFF00FED3FC70A63F" lastPageId="8" lastPageNumber="165" pageId="7" pageNumber="164" type="discussion">
<paragraph id="8B977A23A12AFF82FF00FED3FB3EA647" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<emphasis id="B95CA631A12AFF82FF00FED3FE90A53B" bold="true" box="[199,315,331,357]" pageId="7" pageNumber="164">Remarks.</emphasis>
The
<emphasis id="B95CA631A12AFF82FEB8FED3FD9CA53B" box="[383,567,331,357]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FEB8FED3FE72A53A" authorityName="Malmgren" authorityYear="1867" box="[383,473,331,356]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="genus">Laonice</taxonomicName>
cirrata
</emphasis>
group (
<bibRefCitation id="EFB907D2A12AFF82FD48FED3FC4CA53B" author="Sikorski, A. &amp; Pavlova, L. V." box="[655,999,331,357]" pageId="7" pageNumber="164" pagination="353 - 368" refId="ref8162" refString="Sikorski, A. &amp; Pavlova, L. V. (2016) Three new species of Laonice (Polychaete: Spionidae) from West and Southwest Africa. Zootaxa, 4097 (3), 353 - 368. https: // doi. org / 10.11646 / zootaxa. 4097.3.4" type="journal article" year="2016">Sikorski &amp; Pavlova 2016: 361</bibRefCitation>
) can easily be distinguished by having a prostomium clearly fused with the peristomium on the anterior margin; capillaries on the anterior part of the body arranged in two vertical rows; and by having quite variable numerical characters. Nine species belong to this group:
<taxonomicName id="4C2801A0A12AFF82FF20FE2FFDAEA58F" authority="(M. Sars, 1851)" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[231,517,438,465]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FF20FE2FFEF9A58E" box="[231,338,439,464]" italics="true" pageId="7" pageNumber="164">L. cirrata</emphasis>
(M.Sars, 1851)
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FDD7FE2FFCC2A58F" authority="(Kinberg, 1866)" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[528,873,439,465]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">
<emphasis id="B95CA631A12AFF82FDD7FE2FFD07A58F" box="[528,684,439,465]" italics="true" pageId="7" pageNumber="164">L. brevicornis</emphasis>
(Kinberg, 1866)
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FCB2FE2FFB4EA58F" authority="Soderstrom, 1920" authorityName="Soderstrom" authorityYear="1920" box="[885,1253,439,465]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FCB2FE2FFBBEA58F" box="[885,1045,439,465]" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
Söderström, 1920
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FB36FE2FFEB4A5AB" authority="Pillai, 1961" authorityName="Pillai" authorityYear="1961" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicristata">
<emphasis id="B95CA631A12AFF82FB36FE2FFA37A58F" box="[1265,1436,439,465]" italics="true" pageId="7" pageNumber="164">L. brevicristata</emphasis>
Pillai, 1961
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FEE9FE43FD13A5AB" authority="Nonato et al., 1986" authorityName="Nonato" authorityYear="1986" box="[302,696,475,501]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="branchiata">
<emphasis id="B95CA631A12AFF82FEE9FE43FE61A5AB" box="[302,458,475,501]" italics="true" pageId="7" pageNumber="164">L. branchiata</emphasis>
Nonato
<emphasis id="B95CA631A12AFF82FDF7FE43FDCEA5AB" box="[560,613,475,501]" italics="true" pageId="7" pageNumber="164">et al</emphasis>
., 1986
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FD00FE43FBEBA5AB" authority="Sikorski, 2003" authorityName="Sikorski" authorityYear="2003" box="[711,1088,475,501]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="shamrockensis">
<emphasis id="B95CA631A12AFF82FD00FE43FD7CA5AA" box="[711,727,475,500]" italics="true" pageId="7" pageNumber="164">L</emphasis>
.
<emphasis id="B95CA631A12AFF82FD2FFE43FC27A5AB" box="[744,908,475,501]" italics="true" pageId="7" pageNumber="164">shamrockensis</emphasis>
Sikorski, 2003
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FB88FE43FEC7A647" authority="Sigvaldadottir &amp; Desbruyeres, 2003" authorityName="Sigvaldadottir &amp; Desbruyeres" authorityYear="2003" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="asaccata">
<emphasis id="B95CA631A12AFF82FB88FE43FB7FA5AA" box="[1103,1236,475,500]" italics="true" pageId="7" pageNumber="164">L. asaccata</emphasis>
Sigvaldadottir &amp; Desbruyeres, 2003
</taxonomicName>
;
<taxonomicName id="4C2801A0A12AFF82FEBFFE67FD0EA647" authority="Sikorski, 2011" authorityName="Sikorski" authorityYear="2011" box="[376,677,511,537]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FEBFFE67FE52A647" box="[376,505,511,537]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
<bibRefCitation id="EFB907D2A12AFF82FDC7FE67FD0EA647" author="Sikorski, A. V." box="[512,677,511,537]" pageId="7" pageNumber="164" pagination="201 - 214" refId="ref8104" refString="Sikorski, A. V. (2011) Review of Laonice (Spionidae, Annelida) with remarks on several species and a description of a new species from South Africa. Italian Journal of Zoology, 78 (S 1), 201 - 214. https: // doi. org / 10.1080 / 11250003.2011.617218" type="journal article" year="2011">Sikorski, 2011</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C2801A0A12AFF82FD26FE67FB3AA647" authority="Sikorski &amp; Pavlova, 2016" authorityName="Sikorski &amp; Pavlova" authorityYear="2016" box="[737,1169,511,537]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cricketae">
<emphasis id="B95CA631A12AFF82FD26FE67FCCEA647" box="[737,869,511,537]" italics="true" pageId="7" pageNumber="164">L. cricketae</emphasis>
<bibRefCitation id="EFB907D2A12AFF82FCABFE67FB3AA647" author="Sikorski, A. &amp; Pavlova, L. V." box="[876,1169,511,537]" pageId="7" pageNumber="164" pagination="353 - 368" refId="ref8162" refString="Sikorski, A. &amp; Pavlova, L. V. (2016) Three new species of Laonice (Polychaete: Spionidae) from West and Southwest Africa. Zootaxa, 4097 (3), 353 - 368. https: // doi. org / 10.11646 / zootaxa. 4097.3.4" type="journal article" year="2016">Sikorski &amp; Pavlova, 2016</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FDBBFE04A04F" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00FDBBFE01A662" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,426,547,572]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00FDBBFE01A662" box="[199,426,547,572]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE74FDBBFDA5A663" bold="true" box="[435,526,547,573]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE74FDBBFDA5A663" box="[435,526,547,573]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, belongs to the
<emphasis id="B95CA631A12AFF82FD16FDBBFC20A663" box="[721,907,547,573]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FD16FDBBFC80A662" authorityName="Malmgren" authorityYear="1867" box="[721,811,547,572]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="genus">Laonice</taxonomicName>
cirrata
</emphasis>
group due to the anteriorly fused prostomium and peristomium, the arrangement of the capillaries in the anterior part of the body in two vertical rows. However,
<taxonomicName id="4C2801A0A12AFF82FF50FDF3FED3A6DA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[151,376,619,644]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF50FDF3FED3A6DA" box="[151,376,619,644]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FEB8FDF3FE72A6DB" bold="true" box="[383,473,619,645]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FEB8FDF3FE72A6DB" box="[383,473,619,645]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from all other species by having the nuchal organs and caruncle extending up to chaetigers 1416 (in other species the nuchal organs and caruncle are either shorter (
<taxonomicName id="4C2801A0A12AFF82FB2DFD17FA3FA6F7" authorityName="Pillai" authorityYear="1961" box="[1258,1428,655,681]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicristata">
<emphasis id="B95CA631A12AFF82FB2DFD17FA3FA6F7" box="[1258,1428,655,681]" italics="true" pageId="7" pageNumber="164">L. brevicristata</emphasis>
</taxonomicName>
: to ch. 79,
<taxonomicName id="4C2801A0A12AFF82FED0FD2BFE05A693" authorityName="Nonato, Bolívar &amp; Lana" authorityYear="1986" box="[279,430,691,717]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="branchiate">
<emphasis id="B95CA631A12AFF82FED0FD2BFE05A693" box="[279,430,691,717]" italics="true" pageId="7" pageNumber="164">L. branchiate</emphasis>
</taxonomicName>
: ch. 12,
<taxonomicName id="4C2801A0A12AFF82FDD5FD2BFD7FA693" authorityName="Sikorski" authorityYear="2003" box="[530,724,691,717]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="shamrockensis">
<emphasis id="B95CA631A12AFF82FDD5FD2BFD7FA693" box="[530,724,691,717]" italics="true" pageId="7" pageNumber="164">L. shamrockensis</emphasis>
</taxonomicName>
: ch. 5,
<taxonomicName id="4C2801A0A12AFF82FCEEFD2BFC00A692" authorityName="Sigvaldadottir &amp; Desbruyeres" authorityYear="2003" box="[809,939,691,716]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="asaccata">
<emphasis id="B95CA631A12AFF82FCEEFD2BFC00A692" box="[809,939,691,716]" italics="true" pageId="7" pageNumber="164">L. asaccata</emphasis>
</taxonomicName>
: ch. 4,
<taxonomicName id="4C2801A0A12AFF82FBC7FD2BFB2DA693" authorityName="Sikorski &amp; Pavlova" authorityYear="2016" box="[1024,1158,691,717]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cricketae">
<emphasis id="B95CA631A12AFF82FBC7FD2BFB2DA693" box="[1024,1158,691,717]" italics="true" pageId="7" pageNumber="164">L. cricketae</emphasis>
</taxonomicName>
: caruncle to ch. 12 and nuchal organs to ch. 20), or longer (
<taxonomicName id="4C2801A0A12AFF82FDF3FD4FFD0AA6AE" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[564,673,727,752]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FDF3FD4FFD0AA6AE" box="[564,673,727,752]" italics="true" pageId="7" pageNumber="164">L. cirrata</emphasis>
</taxonomicName>
: to ch. 440,
<taxonomicName id="4C2801A0A12AFF82FC85FD4FFC74A6AF" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[834,991,727,753]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">
<emphasis id="B95CA631A12AFF82FC85FD4FFC74A6AF" box="[834,991,727,753]" italics="true" pageId="7" pageNumber="164">L. brevicornis</emphasis>
</taxonomicName>
: ch. 926,
<taxonomicName id="4C2801A0A12AFF82FBA5FD4FFAAFA6AF" authorityName="Soderstrom" authorityYear="1920" box="[1122,1284,727,753]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FBA5FD4FFAAFA6AF" box="[1122,1284,727,753]" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
</taxonomicName>
: ch. 635,
<taxonomicName id="4C2801A0A12AFF82FA41FD4FFF50A74B" authorityName="Sikorski" authorityYear="2011" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FA41FD4FFF50A74B" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
: ch. 3438). In addition, the notopodial postchaetal lamellae of chaetiger 1 are very small, subtriangular, shorter than the neurolamellae; the dorsal transverse ridges connecting the notopodial postchaetal lamellae on chaetigers 3748 are very low; the neuropodial postchaetal lamellae of chaetiger 1 are rectangular with rounded edges, oval on chaetigers 28 with earlobe-shaped and well developed ventral edges; the ventrolateral pouches appear from chaetigers 3548 up to chaetiger 57 (in the other species mentioned, these pouches start on much earlier chaetigers: between chaetigers 317, or are absent, except in
<taxonomicName id="4C2801A0A12AFF82FC89FC37FC78A797" authorityName="Sikorski &amp; Pavlova" authorityYear="2016" box="[846,979,943,969]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cricketae">
<emphasis id="B95CA631A12AFF82FC89FC37FC78A797" box="[846,979,943,969]" italics="true" pageId="7" pageNumber="164">L. cricketae</emphasis>
</taxonomicName>
where they start from chaetiger 40); the hooded hooks have a distal opening; and the pygidium has two small ventral papilliform cirri arranged close to each other, and five long cirri.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FB83FB23A023" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00FB83FE07A06A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,428,1051,1076]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00FB83FE07A06A" box="[199,428,1051,1076]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE72FB83FDB9A06B" bold="true" box="[437,530,1051,1077]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE72FB83FDB9A06B" box="[437,530,1051,1077]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is most similar to
<emphasis id="B95CA631A12AFF82FD3EFB83FB66A06B" box="[761,1229,1051,1077]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FD3EFB83FCC1A06A" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[761,874,1051,1076]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">L. cirrata</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FCBEFB83FBB0A06B" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[889,1051,1051,1077]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">L. brevicornis</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FBEEFB83FB66A06B" authorityName="Soderstrom" authorityYear="1920" box="[1065,1229,1051,1077]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">L. bahusiensis</taxonomicName>
</emphasis>
, and
<emphasis id="B95CA631A12AFF82FAD6FB83FA37A06B" box="[1297,1436,1051,1077]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FAD6FB83FA33A06B" authorityName="Sikorski" authorityYear="2011" box="[1297,1432,1051,1077]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">L. antipoda</taxonomicName>
,
</emphasis>
because they also have a long caruncle and nuchal organs. However,
<taxonomicName id="4C2801A0A12AFF82FC54FBA7FBD8A006" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[915,1139,1087,1112]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FC54FBA7FBD8A006" box="[915,1139,1087,1112]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FBBEFBA7FB7AA007" bold="true" box="[1145,1233,1087,1113]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FBBEFBA7FB7AA007" box="[1145,1233,1087,1113]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from these species by the morphological characters mentioned above, and by those mentioned below.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FB1FFEB4A19F" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00FB1FFE0DA0FE" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,422,1159,1184]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00FB1FFE0DA0FE" box="[199,422,1159,1184]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE6CFB1FFDA8A0FF" bold="true" box="[427,515,1159,1185]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE6CFB1FFDA8A0FF" box="[427,515,1159,1185]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, can be distinguished from
<emphasis id="B95CA631A12AFF82FCF7FB1FFB40A0FF" box="[816,1259,1159,1185]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FCF7FB1FFC37A0FE" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[816,924,1159,1184]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">L. cirrata</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FC61FB1FFBE9A0FF" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[934,1090,1159,1185]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">L. brevicornis</taxonomicName>
,
<taxonomicName id="4C2801A0A12AFF82FB8BFB1FFB40A0FF" authorityName="Soderstrom" authorityYear="1920" box="[1100,1259,1159,1185]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">L. bahusiensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C2801A0A12AFF82FADAFB1FFA36A0FF" authorityName="Sikorski" authorityYear="2011" box="[1309,1437,1159,1185]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FADAFB1FFA36A0FF" box="[1309,1437,1159,1185]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
by the shape and number of eyes, that in
<taxonomicName id="4C2801A0A12AFF82FDA6FB33FCE9A09A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[609,834,1195,1220]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FDA6FB33FCE9A09A" box="[609,834,1195,1220]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FC8FFB33FC09A09B" bold="true" box="[840,930,1195,1221]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FC8FFB33FC09A09B" box="[840,930,1195,1221]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, have the appearance of raccoon spots on the back of the prostomium; and by the last three pairs of branchiae that are so small, they are hidden in the parapodium. In addition,
<taxonomicName id="4C2801A0A12AFF82FEEFFB6BFDA6A152" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[296,525,1267,1292]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FEEFFB6BFDA6A152" box="[296,525,1267,1292]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FDDFFB6BFDDDA153" bold="true" box="[536,630,1267,1293]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FDDFFB6BFDDDA153" box="[536,630,1267,1293]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C2801A0A12AFF82FCD0FB6BFC2CA152" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[791,903,1267,1292]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FCD0FB6BFC2CA152" box="[791,903,1267,1292]" italics="true" pageId="7" pageNumber="164">L. cirrata</emphasis>
</taxonomicName>
by not having the dorsolateral margin of the notopodial postchaetal lamellae forming a small peak, but having small, triangular posterior notopodial lamellae.
<taxonomicName id="4C2801A0A12AFF82FF50FAA3FED3A10A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[151,376,1339,1364]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF50FAA3FED3A10A" box="[151,376,1339,1364]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FEB9FAA3FE7CA10B" bold="true" box="[382,471,1339,1365]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FEB9FAA3FE7CA10B" box="[382,471,1339,1365]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C2801A0A12AFF82FCC2FAA3FC0AA10B" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[773,929,1339,1365]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="brevicornis">
<emphasis id="B95CA631A12AFF82FCC2FAA3FC0AA10B" box="[773,929,1339,1365]" italics="true" pageId="7" pageNumber="164">L. brevicornis</emphasis>
</taxonomicName>
by short first two pair of branchiae which are slightly shorter than the notopodial postchaetal lamellae; the presence of dorsal transverse ridges connecting the notopodial postchaetal lamellae; and because the main fang of the hooded hooks is not accompanied at the base by apical teeth.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00FA53FEF7A35B" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
In addition,
<taxonomicName id="4C2801A0A12AFF82FE96FA53FD98A1BA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[337,563,1483,1508]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FE96FA53FD98A1BA" box="[337,563,1483,1508]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FDFDFA53FD3EA1BB" bold="true" box="[570,661,1483,1509]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FDFDFA53FD3EA1BB" box="[570,661,1483,1509]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C2801A0A12AFF82FC0EFA53FBC0A1BB" authorityName="Soderstrom" authorityYear="1920" box="[969,1131,1483,1509]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FC0EFA53FBC0A1BB" box="[969,1131,1483,1509]" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2801A0A12AFF82FB65FA53FA8EA1BB" authorityName="Sikorski" authorityYear="2011" box="[1186,1317,1483,1509]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FB65FA53FA8EA1BB" box="[1186,1317,1483,1509]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
by having a prostomium that is broadly rounded anteriorly, bicornate as a cap, instead of triangular, anteriorly wide, truncated or slightly incurved, sometimes broadly rounded; sabre chaetae from chaetigers 2428
<emphasis id="B95CA631A12AFF82FB44F98BFB09A272" box="[1155,1186,1555,1580]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
from ch.
<quantity id="4CD0D7C6A12AFF82FAD0F98BFAD7A273" box="[1303,1404,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="5.334" metricValueMin="2.54" pageId="7" pageNumber="164" unit="in" value="15.5" valueMax="21.0" valueMin="10.0">1021 in</quantity>
<emphasis id="B95CA631A12AFF82FA41F98BFE8AA20F" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FA41F98BFEB7A20F" authorityName="Soderstrom" authorityYear="1920" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">L. bahusiensis</taxonomicName>
,
</emphasis>
and ch.
<quantity id="4CD0D7C6A12AFF82FE4AF9AFFE5EA20F" box="[397,501,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.826" metricValueMin="3.556" pageId="7" pageNumber="164" unit="in" value="16.5" valueMax="19.0" valueMin="14.0">1419 in</quantity>
<emphasis id="B95CA631A12AFF82FDC7F9AFFCD5A20E" box="[512,894,1591,1617]" italics="true" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FDC7F9AFFD22A20F" authorityName="Sikorski" authorityYear="2011" box="[512,649,1591,1617]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">L. antipoda</taxonomicName>
.
<taxonomicName id="4C2801A0A12AFF82FD5FF9AFFCD5A20E" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[664,894,1591,1616]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">Laonice ayseniensis</taxonomicName>
</emphasis>
<emphasis id="B95CA631A12AFF82FC4EF9AFFC4CA20F" bold="true" box="[905,999,1591,1617]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FC4EF9AFFC4CA20F" box="[905,999,1591,1617]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C2801A0A12AFF82FA41F9AFFEB1A22B" authorityName="Soderstrom" authorityYear="1920" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="bahusiensis">
<emphasis id="B95CA631A12AFF82FA41F9AFFEB1A22B" italics="true" pageId="7" pageNumber="164">L. bahusiensis</emphasis>
</taxonomicName>
in having hooded hooks from chaetigers 3538
<emphasis id="B95CA631A12AFF82FCEFF9C3FCECA22A" box="[808,839,1627,1652]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
from ch. 1435, and by the number of teeth (tridentate with two teeth on the main fang, appearing bidentate in the lateral view
<emphasis id="B95CA631A12AFF82FC7CF9E7FC71A2C6" box="[955,986,1663,1688]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
five teeth (tridentate in the lateral view) with very small upper apical teeth.
<taxonomicName id="4C2801A0A12AFF82FDEFF93BFCA1A2E2" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[552,778,1699,1724]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FDEFF93BFCA1A2E2" box="[552,778,1699,1724]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FCD5F93BFCC7A2E3" bold="true" box="[786,876,1699,1725]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FCD5F93BFCC7A2E3" box="[786,876,1699,1725]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C2801A0A12AFF82FB31F93BFAD2A2E3" authorityName="Sikorski" authorityYear="2011" box="[1270,1401,1699,1725]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="B95CA631A12AFF82FB31F93BFAD2A2E3" box="[1270,1401,1699,1725]" italics="true" pageId="7" pageNumber="164">L. antipoda</emphasis>
</taxonomicName>
by the presence of nuchal tentacle, fewer pairs of branchiae (3437
<emphasis id="B95CA631A12AFF82FCBAF95FFC30A2BE" box="[893,923,1735,1760]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
3845); and very low (
<emphasis id="B95CA631A12AFF82FB6BF95FFB61A2BE" box="[1196,1226,1735,1760]" italics="true" pageId="7" pageNumber="164">vs.</emphasis>
prominent) dorsal transverse ridges.
</paragraph>
<paragraph id="8B977A23A12AFF82FF00F897FD87A32F" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="164">
Hartmann-Schröder (1965) recorded three species for
<collectingCountry id="F33F3AB3A12AFF82FCF1F897FCDCA377" box="[822,887,1807,1833]" name="Chile" pageId="7" pageNumber="164">Chile</collectingCountry>
:
<taxonomicName id="4C2801A0A12AFF82FC43F897FB9DA376" baseAuthorityName="M. Sars" baseAuthorityYear="1851" box="[900,1078,1807,1832]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="cirrata">
<emphasis id="B95CA631A12AFF82FC43F897FB9DA376" box="[900,1078,1807,1832]" italics="true" pageId="7" pageNumber="164">Laonice cirrata</emphasis>
</taxonomicName>
, and two subspecies
<taxonomicName id="4C2801A0A12AFF82FAE9F897FEB8A312" authorityName="Hartmann-Schroder" authorityYear="1965" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="subSpecies" species="cirrata" subSpecies="postcirrata">
<emphasis id="B95CA631A12AFF82FAE9F897FEB8A312" italics="true" pageId="7" pageNumber="164">L. cirrata postcirrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2801A0A12AFF82FE8AF8ABFDE8A312" box="[333,579,1843,1868]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="subSpecies" species="cirrata" subSpecies="praecirrata">
<emphasis id="B95CA631A12AFF82FE8AF8ABFDE8A312" box="[333,579,1843,1868]" italics="true" pageId="7" pageNumber="164">L. cirrata praecirrata</emphasis>
</taxonomicName>
. The first species is already discussed above; and the other two species are very different from the new species.
</paragraph>
<paragraph id="8B977A23A12AFF8DFF00F8E3FDB9A663" blockId="7.[151,1437,150,2013]" lastBlockId="8.[151,1437,151,609]" lastPageId="8" lastPageNumber="165" pageId="7" pageNumber="164">
<taxonomicName id="4C2801A0A12AFF82FF00F8E3FE02A3CA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[199,425,1915,1940]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A12AFF82FF00F8E3FE02A3CA" box="[199,425,1915,1940]" italics="true" pageId="7" pageNumber="164">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A12AFF82FE77F8E3FDA1A3CB" bold="true" box="[432,522,1915,1941]" pageId="7" pageNumber="164">
<taxonomicNameLabel id="A26F1B4AA12AFF82FE77F8E3FDA1A3CB" box="[432,522,1915,1941]" pageId="7" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C2801A0A12AFF82FCFBF8E3FBB2A3CA" box="[828,1049,1915,1940]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A12AFF82FCFBF8E3FBB2A3CA" box="[828,1049,1915,1940]" italics="true" pageId="7" pageNumber="164">Laonice postcirrata</emphasis>
</taxonomicName>
in that the former has an occipital tentacle situated slightly forward of the first pair of notopodial lamellae, whilst
<taxonomicName id="4C2801A0A12AFF82FBC9F807FB0CA3E6" box="[1038,1191,1951,1976]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="7" pageNumber="164" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A12AFF82FBC9F807FB0CA3E6" box="[1038,1191,1951,1976]" italics="true" pageId="7" pageNumber="164">L. postcirrata</emphasis>
</taxonomicName>
has a tentacle located between the first and second pair of notopodial lamellae; the prostomium and peristomium are completely fused in the anterior part of the prostomium, unlike
<taxonomicName id="4C2801A0A125FF8DFDB6FF0FFCA1A4EE" box="[625,778,151,176]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFDB6FF0FFCA1A4EE" box="[625,778,151,176]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
where they are not fused. Another difference is in the shape and number of eyes:
<taxonomicName id="4C2801A0A125FF8DFEB9FF23FDF5A48A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[382,606,187,212]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFEB9FF23FDF5A48A" box="[382,606,187,212]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFDA3FF23FD17A48B" bold="true" box="[612,700,187,213]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFDA3FF23FD17A48B" box="[612,700,187,213]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, has the appearance of raccoon spots on the posterior region of the prostomium, whereas
<taxonomicName id="4C2801A0A125FF8DFE4AFF47FD8DA4A6" box="[397,550,223,248]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE4AFF47FD8DA4A6" box="[397,550,223,248]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
has two or four small eyes arranged in the same row, or in a square or trapezoid.
<taxonomicName id="4C2801A0A125FF8DFF50FE9BFEDCA542" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[151,375,259,284]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFF50FE9BFEDCA542" box="[151,375,259,284]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFEBAFE9BFE7DA543" bold="true" box="[381,470,259,285]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFEBAFE9BFE7DA543" box="[381,470,259,285]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, also has a caruncle and branchiae that reach chaetigers 1416 and 3538
<emphasis id="B95CA631A125FF8DFACDFE9BFA82A542" box="[1290,1321,259,284]" italics="true" pageId="8" pageNumber="165">vs.</emphasis>
ch. 2228 and 42, respectively, in
<taxonomicName id="4C2801A0A125FF8DFE63FEBFFD95A51E" box="[420,574,295,320]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE63FEBFFD95A51E" box="[420,574,295,320]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
. Furthermore, the shape of the anterior notopodial and neuropodial lamellae are different: in
<taxonomicName id="4C2801A0A125FF8DFE91FED3FD90A53A" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[342,571,331,356]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFE91FED3FD90A53A" box="[342,571,331,356]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFD82FED3FD09A53B" bold="true" box="[581,674,331,357]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFD82FED3FD09A53B" box="[581,674,331,357]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the notopodial lamellae have a broad subtriangular shape with a blunt tip, and the postchaetal neuropodial lamellae on chaetigers 28 are oval with well-developed, earlobe-shaped ventral edges, whilst in
<taxonomicName id="4C2801A0A125FF8DFE61FE0BFDEAA5F2" box="[422,577,403,428]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE61FE0BFDEAA5F2" box="[422,577,403,428]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
the notopodial lamellae have pointed tips, and the neuropodial lamellae are wide and subtriangular. Ventrolateral pouches appear from from chaetigers
<quantity id="4CD0D7C6A125FF8DFC2BFE2EFBFBA58F" box="[1004,1104,438,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0541" metricValueMax="1.2192" metricValueMin="0.889" pageId="8" pageNumber="165" unit="in" value="41.5" valueMax="48.0" valueMin="35.0">3548 in</quantity>
<taxonomicName id="4C2801A0A125FF8DFB9FFE2FFA91A58E" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[1112,1338,439,464]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFB9FFE2FFA91A58E" box="[1112,1338,439,464]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFA85FE2FFA37A58F" bold="true" box="[1346,1436,439,465]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFA85FE2FFA37A58F" box="[1346,1436,439,465]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B95CA631A125FF8DFF50FE43FF1DA5AA" box="[151,182,475,500]" italics="true" pageId="8" pageNumber="165">vs.</emphasis>
from ch.
<quantity id="4CD0D7C6A125FF8DFEEDFE42FE25A5AB" box="[298,398,474,501]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.81" metricValueMin="2.794" pageId="8" pageNumber="165" unit="in" value="13.0" valueMax="15.0" valueMin="11.0">1115 in</quantity>
<taxonomicName id="4C2801A0A125FF8DFE50FE43FCEEA5AB" authority="(Sirkorski et al., 2016)" baseAuthorityName="Sirkorski" baseAuthorityYear="2016" box="[407,837,475,501]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFE50FE43FD98A5AA" box="[407,563,475,500]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
(Sirkorski
<emphasis id="B95CA631A125FF8DFD71FE43FD5AA5AB" box="[694,753,475,501]" italics="true" pageId="8" pageNumber="165">et al.</emphasis>
, 2016)
</taxonomicName>
, and finally,
<taxonomicName id="4C2801A0A125FF8DFC1BFE43FB6BA5AA" authorityName="Delgado-Blas &amp; Díaz-Díaz &amp; Rozbaczylo" authorityYear="2024" box="[988,1216,475,500]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="ayseniensis" status="sp. nov.">
<emphasis id="B95CA631A125FF8DFC1BFE43FB6BA5AA" box="[988,1216,475,500]" italics="true" pageId="8" pageNumber="165">Laonice ayseniensis</emphasis>
</taxonomicName>
<emphasis id="B95CA631A125FF8DFB0EFE43FA8FA5AB" bold="true" box="[1225,1316,475,501]" pageId="8" pageNumber="165">
<taxonomicNameLabel id="A26F1B4AA125FF8DFB0EFE43FA8FA5AB" box="[1225,1316,475,501]" pageId="8" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has transdorsal ridges, and the neuropodial hooks and sabre chaetae appear from chaetigers 3538 and 2428
<emphasis id="B95CA631A125FF8DFAC4FE67FA8AA646" box="[1283,1313,511,536]" italics="true" pageId="8" pageNumber="165">vs.</emphasis>
ch. 31 and 16, respectively, in
<taxonomicName id="4C2801A0A125FF8DFEB6FDBBFDA0A662" box="[369,523,547,572]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="165" phylum="Annelida" rank="species" species="postcirrata">
<emphasis id="B95CA631A125FF8DFEB6FDBBFDA0A662" box="[369,523,547,572]" italics="true" pageId="8" pageNumber="165">L. postcirrata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B977A23A125FF8DFF00FDDFFC70A63F" blockId="8.[151,1437,151,609]" box="[199,987,583,609]" pageId="8" pageNumber="165">Differences compared to other similar species can be found in the key.</paragraph>
</subSubSection>
</treatment>
</document>