738 lines
99 KiB
XML
738 lines
99 KiB
XML
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<mods:title id="3C0365FBB23DE0D42293E8A82F6455CA">New and revised taxa of Carboniferous spiriferides (Brachiopoda, Spiriferida) from the Donets Basin (Ukraine) and South Urals (Russia)</mods:title>
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<mods:namePart id="27E203E4E7A735BFA2043C1B801AA4D4">Poletaev, Vladyslav</mods:namePart>
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<mods:affiliation id="48B5A040BD5682C8ED045529170E1A1F">Institute of Geological Sciences of the National Academy of Sciences of Ukraine, 55 - b, Oles Honchar St., Kyiv, 01054, Ukraine.</mods:affiliation>
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<mods:nameIdentifier id="A37AAC8FDDE3C32EEFAC8470ED871F50" type="email">vipoletaev38@gmail.com</mods:nameIdentifier>
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<mods:title id="9D8764314056ACE83ADEC092C03105CC">European Journal of Taxonomy</mods:title>
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<mods:part id="6319DADE73FFCD12B3E96404A29C8407">
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<mods:date id="475F007D78A0C5853514F80D7A7E68CF">2024</mods:date>
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<mods:number id="B081B8B0F2A02072B1DF27B0737178EE">2024-11-04</mods:number>
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<treatment id="03BA8E19595BD200FDD7C88CFD05B13A" ID-DOI="http://doi.org/10.5281/zenodo.14052676" ID-Zenodo-Dep="14052676" LSID="urn:lsid:plazi:treatment:03BA8E19595BD200FDD7C88CFD05B13A" httpUri="http://treatment.plazi.org/id/03BA8E19595BD200FDD7C88CFD05B13A" lastPageId="14" lastPageNumber="145" pageId="11" pageNumber="143">
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<subSubSection id="C3096C84595BD205FDD7C88CFC1BB79A" box="[589,999,1189,1216]" pageId="11" pageNumber="143" type="nomenclature">
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<paragraph id="8BAC3F0F595BD205FDD7C88CFC1BB79A" blockId="11.[589,999,1189,1216]" box="[589,999,1189,1216]" pageId="11" pageNumber="143">
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Family
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||
<taxonomicName id="4C13448C595BD205FD3DC88CFC1BB79A" ID-CoL="7NK5X" authority="Poletaev, 2015" authorityName="Poletaev" authorityYear="2015" box="[679,999,1189,1216]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="11" pageNumber="143" phylum="Cnidaria" rank="family">
|
||
<emphasis id="B967E31D595BD205FD3DC88CFCCFB7E5" bold="true" box="[679,819,1189,1215]" pageId="11" pageNumber="143">Oceaniidae</emphasis>
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||
<bibRefCitation id="EF8242FE595BD205FCA0C88CFC1BB79A" author="Poletaev V. I." box="[826,999,1189,1216]" pageId="11" pageNumber="143" pagination="17 - 18" refId="ref12092" refString="Poletaev V. I. 2015. Novyy rod i semeystvo kamennougol'nykh spiriferid Yevropy. In: Gozhyk P. F. (ed.) Stratotypovi ta oporni rozrizy fanerozois'kykh vidkladiv Ukrainy: suchasnyi stan ta perspektyvy podal'shykh doslidzhen'. Materialy 36 sesii Paleontologichnogo tovarustva Ukrainy: 17 - 18. Kyiv." type="book chapter" year="2015">Poletaev, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3096C84595BD205FF27C8CFFE44B685" pageId="11" pageNumber="143" type="diagnosis">
|
||
<paragraph id="8BAC3F0F595BD205FF27C8CFFECAB65A" blockId="11.[189,310,1254,1280]" box="[189,310,1254,1280]" pageId="11" pageNumber="143">
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||
<heading id="D0E48863595BD205FF27C8CFFECAB65A" bold="true" box="[189,310,1254,1280]" fontSize="11" level="3" pageId="11" pageNumber="143" reason="3">
|
||
<emphasis id="B967E31D595BD205FF27C8CFFECAB65A" bold="true" box="[189,310,1254,1280]" pageId="11" pageNumber="143">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595BD205FF27C93CFE44B685" blockId="11.[189,1399,1301,1503]" pageId="11" pageNumber="143">Small or very small-sized, transverse, ventribiconvex shell; smooth and with two simple or compound opposed plica, dissected median parts of both valves on three parts. Fold and sulcus rarely have median opposite ribs; ventral interarea high or very high, flat, catacline or procline with strong vertical crenulations; hinge line denticulate; narrow and high delthyrium partly covered by pseudodeltidium; ornament of growth lamella only; ventral interior with low dental flanges; dorsal with ctenophoridium and short crural base.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3096C84595BD205FF27CA2FFB06B515" pageId="11" pageNumber="143" type="taxon_list">
|
||
<paragraph id="8BAC3F0F595BD205FF27CA2FFE71B57A" blockId="11.[189,397,1542,1568]" box="[189,397,1542,1568]" pageId="11" pageNumber="143">
|
||
<heading id="D0E48863595BD205FF27CA2FFE71B57A" bold="true" box="[189,397,1542,1568]" fontSize="11" level="3" pageId="11" pageNumber="143" reason="3">
|
||
<emphasis id="B967E31D595BD205FF27CA2FFE71B57A" bold="true" box="[189,397,1542,1568]" pageId="11" pageNumber="143">Genera included</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595BD205FF27CA1CFB06B515" blockId="11.[189,1274,1589,1616]" box="[189,1274,1589,1616]" pageId="11" pageNumber="143">
|
||
<taxonomicName id="4C13448C595BD205FF27CA1CFEAFB515" authorityName="Poletaev" authorityYear="2024" box="[189,339,1589,1615]" genus="Johnsoniana" pageId="11" pageNumber="149" rank="genus" status="nom. nov.">
|
||
<emphasis id="B967E31D595BD205FF27CA1CFEAFB515" box="[189,339,1589,1615]" italics="true" pageId="11" pageNumber="143">Johnsoniana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2545E66595BD205FEC0CA1FFE2DB50A" box="[346,465,1590,1616]" pageId="11" pageNumber="143">nom. nov.</taxonomicNameLabel>
|
||
, possibly
|
||
<taxonomicName id="4C13448C595BD205FDDCCA1CFC88B515" authority="Brunton, 1984" authorityName="Brunton" authorityYear="1984" box="[582,884,1589,1616]" class="Rhynchonellata" family="Verneuiliidae" genus="Minythyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595BD205FDDCCA1CFD3CB515" box="[582,704,1589,1615]" italics="true" pageId="11" pageNumber="143">Minythyra</emphasis>
|
||
<bibRefCitation id="EF8242FE595BD205FD52CA1FFC88B515" author="Brunton C. H. C." box="[712,884,1589,1616]" pageId="11" pageNumber="143" pagination="27 - 127" refId="ref9844" refString="Brunton C. H. C. 1984. Silicified brachiopods from the Visean of County Fermanagh, Ireland. (III). Rhynchonellids, spiriferids and terebratulids. Bulletin of the British Museum (Natural History) Geology 38 (2): 27 - 127. Available from https: // www. biodiversitylibrary. org / page / 2273038 [accessed 7 Oct. 2024]." type="journal article" year="1984">Brunton, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C13448C595BD205FC1ACA1CFB09B515" authority="Sun et al., 2004" authorityName="Sun" authorityYear="2004" box="[896,1269,1589,1616]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595BD205FC1ACA1CFBCEB515" box="[896,1074,1589,1615]" italics="true" pageId="11" pageNumber="143">Changshunella</emphasis>
|
||
<bibRefCitation id="EF8242FE595BD205FBA3CA1CFB09B515" author="Sun Y. & Balinski A. & Ma X. & Zhang Y." box="[1081,1269,1589,1616]" pageId="11" pageNumber="143" pagination="267 - 274" refId="ref12589" refString="Sun Y., Balinski A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267 - 274." type="journal article" year="2004">
|
||
Sun
|
||
<emphasis id="B967E31D595BD205FBF4CA1FFB55B515" box="[1134,1193,1589,1615]" italics="true" pageId="11" pageNumber="143">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3096C84595BD200FF27CA5EFD05B13A" lastPageId="14" lastPageNumber="146" pageId="11" pageNumber="143" type="discussion">
|
||
<paragraph id="8BAC3F0F595BD205FF27CA5EFED3B5CB" blockId="11.[189,303,1655,1681]" box="[189,303,1655,1681]" pageId="11" pageNumber="143">
|
||
<heading id="D0E48863595BD205FF27CA5EFED3B5CB" bold="true" box="[189,303,1655,1681]" fontSize="11" level="3" pageId="11" pageNumber="143" reason="3">
|
||
<emphasis id="B967E31D595BD205FF27CA5EFED3B5CB" bold="true" box="[189,303,1655,1681]" pageId="11" pageNumber="143">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595BD202FF27CA8CFE2DB2EA" blockId="11.[187,1399,1701,1974]" lastBlockId="12.[188,1401,265,432]" lastPageId="12" lastPageNumber="144" pageId="11" pageNumber="143">
|
||
The history of closely related genera and species previously included in the family
|
||
<taxonomicName id="4C13448C595BD205FB40CA8CFECEB5B8" authority="Schuchert" authorityName="Schuchert" authorityYear="1929" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="family">Verneuiliidae Schuchert</taxonomicName>
|
||
in Schuchert & Le Vene, 1929) is long and confusing.
|
||
<bibRefCitation id="EF8242FE595BD205FC5ECAE1FB3DB5B9" author="Hall J. & Clarke J. M." box="[964,1217,1736,1763]" pageId="11" pageNumber="143" refId="ref10739" refString="Hall J. & Clarke J. M. 1893. An Introduction to the Study of the Genera of Paleozoic Brachiopoda: intended as a Handbook for the Use of Students. Palaeontology of New York 8 Pt. 2. James B. Lyon, Albany. https: // doi. org / 10.5962 / bhl. title. 15111" type="book" year="1893">Hall & Clarke (1893)</bibRefCitation>
|
||
established the genus
|
||
<taxonomicName id="4C13448C595BD205FE9DCAC5FE7CB45C" authorityName="Hall & Clarke" authorityYear="1893" box="[263,384,1772,1798]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595BD205FE9DCAC5FE7CB45C" box="[263,384,1772,1798]" italics="true" pageId="11" pageNumber="143">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
, which included two species:
|
||
<taxonomicName id="4C13448C595BD205FD43CAC5FAC4B45C" authority="d'Archiac & de Verneuil, 1842" authorityName="d'Archiac & de Verneuil" authorityYear="1842" box="[729,1336,1772,1798]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595BD205FD43CAC5FC30B45C" box="[729,972,1772,1798]" italics="true" pageId="11" pageNumber="143">Spirifer cheiropteryx</emphasis>
|
||
<bibRefCitation id="EF8242FE595BD205FC49CAC5FAC4B45C" author="d'Archiac V. E. & de Verneuil E. P." box="[979,1336,1772,1798]" pageId="11" pageNumber="143" pagination="303 - 410" refId="ref9672" refString="d'Archiac V. E. & de Verneuil E. P. 1842. On the fossils of the older deposits in the Rheinish Provinces. Transactions Geological Society 6 (2): 303 - 410. https: // doi. org / 10.1144 / transgslb. 6.2.303" type="journal article" year="1842">d’Archiac & de Verneuil, 1842</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Late Devonian of
|
||
<collectingCountry id="F3047F9F595BD205FE23CB26FDDAB473" box="[441,550,1807,1833]" name="Germany" pageId="11" pageNumber="143">Germany</collectingCountry>
|
||
and
|
||
<taxonomicName id="4C13448C595BD205FDFACB26FC2BB470" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" box="[608,983,1807,1834]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595BD205FDFACB26FCF2B473" box="[608,782,1807,1833]" italics="true" pageId="11" pageNumber="143">Spirifer oceani</emphasis>
|
||
<bibRefCitation id="EF8242FE595BD205FC8CCB26FC2BB470" author="d'Orbigny A." box="[790,983,1807,1834]" pageId="11" pageNumber="143" pagination="295 - 353" refId="ref11682" refString="d'Orbigny A. 1850. Memoire sur les Brachiopodes. II. Classification des Brachiopodes. Annales des Sciences Naturelles. Zoologie, Serie 3 e 13: 295 - 353. Available from https: // www. biodiversitylibrary. org / page / 13434796 [accessed 7 Oct. 2024]." type="book chapter" year="1850">d’Orbigny, 1850</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Viséan of
|
||
<collectingCountry id="F3047F9F595BD205FB24CB26FADBB473" box="[1214,1319,1807,1833]" name="Belgium" pageId="11" pageNumber="143">Belgium</collectingCountry>
|
||
. These species, according to the authors of the genus, share common characters: the shell is equibiconvex with each valve “divided externally by two strong divergent ridges into three depressed areas, one central and two lateral” (
|
||
<bibRefCitation id="EF8242FE595BD205FEC8CB51FD9BB4C9" author="Hall J. & Clarke J. M." box="[338,615,1912,1939]" pageId="11" pageNumber="143" refId="ref10739" refString="Hall J. & Clarke J. M. 1893. An Introduction to the Study of the Genera of Paleozoic Brachiopoda: intended as a Handbook for the Use of Students. Palaeontology of New York 8 Pt. 2. James B. Lyon, Albany. https: // doi. org / 10.5962 / bhl. title. 15111" type="book" year="1893">Hall & Clarke 1893: 58</bibRefCitation>
|
||
). Schuchert & Le Vene (1929) placed
|
||
<taxonomicName id="4C13448C595BD205FB81CB51FB68B4C8" authorityName="Hall & Clarke" authorityYear="1893" box="[1051,1172,1912,1938]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595BD205FB81CB51FB68B4C8" box="[1051,1172,1912,1938]" italics="true" pageId="11" pageNumber="143">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
in a new subfamily and in the family
|
||
<taxonomicName id="4C13448C595BD205FE10CBB5FDDAB4EC" authorityName="Schuchert" authorityYear="1929" box="[394,550,1948,1974]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
of the suborder
|
||
<taxonomicName id="4C13448C595BD205FD7CCBB5FC86B4EC" authorityName="Waagen" authorityYear="1883" box="[742,890,1948,1974]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="11" pageNumber="143" phylum="Brachiopoda" rank="subOrder" subOrder="Spiriferidina">Spiriferidina</taxonomicName>
|
||
(Schuchert in Schuchert & Le Vene 1929).
|
||
<bibRefCitation id="EF8242FE595CD202FF27CD20FE87B279" author="Tiazheva A. P." box="[189,379,265,291]" pageId="12" pageNumber="144" pagination="406 - 409" refId="ref12629" refString="Tiazheva A. P. 1960. Novyy vid devonskikh retikulyariid Urala. In: Markovskiy B. P. (ed.) Novyye vidy drevnikh rasteniy i bespozvonochnykh SSSR: 406 - 409. Vsecoyuznyi geologicheskiy institute, Leningrad." type="book chapter" year="1960">Tiazheva (1960)</bibRefCitation>
|
||
established
|
||
<taxonomicName id="4C13448C595CD202FD93CD20FCA9B279" authority="Tiazheva, 1960" authorityName="Tiazheva" authorityYear="1960" box="[521,853,265,291]" class="Rhynchonellata" family="Verneuiliidae" genus="Nuguschella" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FD93CD20FD60B279" box="[521,668,265,291]" italics="true" pageId="12" pageNumber="144">Nuguschella</emphasis>
|
||
<bibRefCitation id="EF8242FE595CD202FD38CD20FCA9B279" author="Tiazheva A. P." box="[674,853,265,291]" pageId="12" pageNumber="144" pagination="406 - 409" refId="ref12629" refString="Tiazheva A. P. 1960. Novyy vid devonskikh retikulyariid Urala. In: Markovskiy B. P. (ed.) Novyye vidy drevnikh rasteniy i bespozvonochnykh SSSR: 406 - 409. Vsecoyuznyi geologicheskiy institute, Leningrad." type="book chapter" year="1960">Tiazheva, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Middle Devonian of the Urals, whose
|
||
<typeStatus id="54A881AD595CD202FF26CD04FF13B21D" box="[188,239,301,327]" pageId="12" pageNumber="144">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C13448C595CD202FEC3CD05FD0BB21C" authority="Tiazheva, 1960" authorityName="Tiazheva" authorityYear="1960" box="[345,759,300,326]" class="Rhynchonellata" family="Verneuiliidae" genus="Nuguschella" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="polita">
|
||
<emphasis id="B967E31D595CD202FEC3CD05FDC6B21C" box="[345,570,300,326]" italics="true" pageId="12" pageNumber="144">Nuguschella polita</emphasis>
|
||
<bibRefCitation id="EF8242FE595CD202FDDBCD05FD0BB21C" author="Tiazheva A. P." box="[577,759,300,326]" pageId="12" pageNumber="144" pagination="406 - 409" refId="ref12629" refString="Tiazheva A. P. 1960. Novyy vid devonskikh retikulyariid Urala. In: Markovskiy B. P. (ed.) Novyye vidy drevnikh rasteniy i bespozvonochnykh SSSR: 406 - 409. Vsecoyuznyi geologicheskiy institute, Leningrad." type="book chapter" year="1960">Tiazheva, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
has a cruralium. The genus was placed in the family
|
||
<taxonomicName id="4C13448C595CD202FF25CD66FEA7B233" authorityName="Schuchert" authorityYear="1929" box="[191,347,335,361]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8242FE595CD202FE0BCD79FD77B230" author="Ivanova E. A." box="[401,651,335,362]" pageId="12" pageNumber="144" pagination="264 - 280" refId="ref10886" refString="Ivanova E. A. 1960. Otriad Spiriferida. In: Orlov Y. A. (ed.) Osnovy Paleontologii. Mshanki, Brachiopody. Vol. 7: 264 - 280. Izdatel'stvo Akademii Nauk SSSR, Moscow." type="book chapter" year="1960">Ivanova (1960: 272)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF8242FE595CD202FD54CD66FC9EB230" author="Pitrat C. W." box="[718,866,335,362]" pageId="12" pageNumber="144" refId="ref11797" refString="Pitrat C. W. 1965. Spiriferidina. In: Moore R. C. (ed.) Treatise on Invertebrate Paleontology, Part H, Brachiopoda: H 667 - H 728. Geological Society of America and University of Kansas, Lawrence." type="journal volume" year="1965">Pitrat (1965</bibRefCitation>
|
||
: H727), but they considered the systematic position of the family uncertain.
|
||
<bibRefCitation id="EF8242FE595CD202FDD6CD5AFCFEB2D6" author="Ivanova E. A." box="[588,770,370,397]" pageId="12" pageNumber="144" pagination="264 - 280" refId="ref10886" refString="Ivanova E. A. 1960. Otriad Spiriferida. In: Orlov Y. A. (ed.) Osnovy Paleontologii. Mshanki, Brachiopody. Vol. 7: 264 - 280. Izdatel'stvo Akademii Nauk SSSR, Moscow." type="book chapter" year="1960">Ivanova (1960)</bibRefCitation>
|
||
placed
|
||
<taxonomicName id="4C13448C595CD202FCF9CD5BFC20B2D6" authorityName="Hall & Clarke" authorityYear="1893" box="[867,988,370,396]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FCF9CD5BFC20B2D6" box="[867,988,370,396]" italics="true" pageId="12" pageNumber="144">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
separately in association with the “plicate ambocoeliids”.
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595CD202FF27CDFEFD0AB051" blockId="12.[188,1400,471,779]" pageId="12" pageNumber="144">
|
||
<bibRefCitation id="EF8242FE595CD202FF27CDFEFE44B2AB" author="Nalivkin D. V." box="[189,440,471,497]" pageId="12" pageNumber="144" refId="ref11601" refString="Nalivkin D. V. 1979. Brakhiopody turneyskogo yarusa Urala. Nauka, Leningrad." type="book" year="1979">Nalivkin (1979: 143)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C13448C595CD202FDA6CDFEFCF0B2AB" box="[572,780,471,497]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FDA6CDFEFCF0B2AB" box="[572,780,471,497]" italics="true" pageId="12" pageNumber="144">Verneuilia oceani</emphasis>
|
||
</taxonomicName>
|
||
from the late Tournaisian of the Urals as a member of the family
|
||
<taxonomicName id="4C13448C595CD202FEFDCDD3FDFFB14E" authorityName="Schuchert" authorityYear="1929" box="[359,515,506,532]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
and confidently assigned this family to the suborder
|
||
<taxonomicName id="4C13448C595CD202FB15CDD3FADBB14E" authorityName="Waagen" authorityYear="1883" box="[1167,1319,506,532]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="subOrder" subOrder="Spiriferidina">Spiriferidina</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF8242FE595CD202FAAECDD2FEB0B16D" author="Wang Y. & Jin Y. G. & Fang D. W." pageId="12" pageNumber="144" refId="ref12753" refString="Wang Y., Jin Y. G. & Fang D. W. 1966. Fossil Brachiopoda. Science Press, Beijing." type="book" year="1966">
|
||
Wang
|
||
<emphasis id="B967E31D595CD202FF27CE37FF04B162" box="[189,248,542,568]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
(1966)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF8242FE595CD202FE1FCE37FD94B162" author="Zhang C. & Zhang F. M. & Zhang Z. X. & Wang Z." box="[389,616,542,568]" pageId="12" pageNumber="144" pagination="262 - 386" refId="ref12998" refString="Zhang C., Zhang F. M., Zhang Z. X. & Wang Z. 1983. Phylum Brachiopoda. In: Paleontological Atlas of Northwestern China for Xinjiang Uygur Autonomous Region, Volume 2 (Upper Paleozoic): 262 - 386. Geological Publishing House, Beijing." type="book chapter" year="1983">
|
||
Zhang
|
||
<emphasis id="B967E31D595CD202FE4DCE37FDEEB162" box="[471,530,542,568]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
(1983)
|
||
</bibRefCitation>
|
||
also examined members of the
|
||
<taxonomicName id="4C13448C595CD202FC40CE37FB86B162" authorityName="Schuchert" authorityYear="1929" box="[986,1146,542,568]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
, but assigned them to the family
|
||
<taxonomicName id="4C13448C595CD202FEA7CE68FD85B101" authority="Frederiks, 1911" authorityName="Frederiks" authorityYear="1911" box="[317,633,577,603]" class="Rhynchonellata" family="Cyrtinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">
|
||
Cyrtinidae
|
||
<bibRefCitation id="EF8242FE595CD202FE5BCE68FD85B101" author="Frederiks G. N." box="[449,633,577,603]" pageId="12" pageNumber="144" pagination="1 - 11" refId="ref10453" refString="Frederiks G. N. 1911. Bemerkung uber einige Oberpalaeozoische Fosilien von Krasnoufimsk. Prilozhenie k Protokolam Zasedaniy Obshchestva Estestvoispytateley pri Imperatorskom Kazanskom Universitete 42 (269): 1 - 11." type="journal article" year="1911">Frederiks, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
on the basis of internal characters.
|
||
<bibRefCitation id="EF8242FE595CD202FB87CE68FAECB101" author="Brunton C. H. C." box="[1053,1296,577,603]" pageId="12" pageNumber="144" pagination="27 - 127" refId="ref9844" refString="Brunton C. H. C. 1984. Silicified brachiopods from the Visean of County Fermanagh, Ireland. (III). Rhynchonellids, spiriferids and terebratulids. Bulletin of the British Museum (Natural History) Geology 38 (2): 27 - 127. Available from https: // www. biodiversitylibrary. org / page / 2273038 [accessed 7 Oct. 2024]." type="journal article" year="1984">Brunton (1984: 101)</bibRefCitation>
|
||
selected specimen B7923 as the
|
||
<typeStatus id="54A881AD595CD202FE4BCE4CFDD2B125" box="[465,558,613,639]" pageId="12" pageNumber="144" type="neotype">neotype</typeStatus>
|
||
among three Belgian
|
||
<typeStatus id="54A881AD595CD202FCB4CE4CFC5CB125" box="[814,928,613,639]" pageId="12" pageNumber="144" type="topotype">topotypes</typeStatus>
|
||
from the NHM collection. He described and illustrated it for the first time as a
|
||
<typeStatus id="54A881AD595CD202FD3DCEA1FCF8B1F8" box="[679,772,648,674]" pageId="12" pageNumber="144" type="neotype">neotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C13448C595CD202FCACCEAEFBF5B1FB" box="[822,1033,647,673]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FCACCEAEFBF5B1FB" box="[822,1033,647,673]" italics="true" pageId="12" pageNumber="144">Verneuilia oceani</emphasis>
|
||
</taxonomicName>
|
||
. He re-established the family
|
||
<taxonomicName id="4C13448C595CD202FF27CE83FEA1B19E" authorityName="Schuchert" authorityYear="1929" box="[189,349,682,708]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
, in which he included the new Visean genus
|
||
<taxonomicName id="4C13448C595CD202FC1CCE83FBFCB19E" authorityName="Brunton" authorityYear="1984" box="[902,1024,682,708]" class="Rhynchonellata" family="Verneuiliidae" genus="Minythyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FC1CCE83FBFCB19E" box="[902,1024,682,708]" italics="true" pageId="12" pageNumber="144">Minythyra</emphasis>
|
||
</taxonomicName>
|
||
, but retained for
|
||
<taxonomicName id="4C13448C595CD202FB42CE83FAADB19E" authorityName="Hall & Clarke" authorityYear="1893" box="[1240,1361,682,708]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FB42CE83FAADB19E" box="[1240,1361,682,708]" italics="true" pageId="12" pageNumber="144">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
its Devonian type species
|
||
<taxonomicName id="4C13448C595CD202FE50CEE7FD42B1B2" authorityName="d'Archiac & de Verneuil" authorityYear="1842" box="[458,702,718,744]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595CD202FE50CEE7FD42B1B2" box="[458,702,718,744]" italics="true" pageId="12" pageNumber="144">Spirifer cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF8242FE595CD202FD56CEE7FC7DB1B2" author="Brunton C. H. C." box="[716,897,718,744]" pageId="12" pageNumber="144" pagination="27 - 127" refId="ref9844" refString="Brunton C. H. C. 1984. Silicified brachiopods from the Visean of County Fermanagh, Ireland. (III). Rhynchonellids, spiriferids and terebratulids. Bulletin of the British Museum (Natural History) Geology 38 (2): 27 - 127. Available from https: // www. biodiversitylibrary. org / page / 2273038 [accessed 7 Oct. 2024]." type="journal article" year="1984">Brunton (1984)</bibRefCitation>
|
||
doubtfully placed the family
|
||
<taxonomicName id="4C13448C595CD202FB40CEE7FA8AB1B2" authorityName="Schuchert" authorityYear="1929" box="[1242,1398,718,744]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
in the superfamily
|
||
<taxonomicName id="4C13448C595CD202FE03CED8FD0EB051" authority="Waagen, 1883" authorityName="Waagen" authorityYear="1883" box="[409,754,753,779]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="superFamily" superFamily="Reticulariacea">
|
||
Reticulariacea
|
||
<bibRefCitation id="EF8242FE595CD202FDD2CED8FD0EB051" author="Waagen W. H." box="[584,754,753,779]" pageId="12" pageNumber="144" pagination="391 - 546" refId="ref12671" refString="Waagen W. H. 1883. Salt Range fossils, vol. 1, part 4. Productus Limestone fossils, Brachiopoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, series 13 2: 391 - 546." type="book chapter" year="1883">Waagen, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595CD202FF27CF1BFD6DB73C" blockId="12.[188,1399,818,1126]" pageId="12" pageNumber="144">
|
||
<bibRefCitation id="EF8242FE595CD202FF27CF1BFE1CB017" author="Gourvennec R." box="[189,480,818,845]" pageId="12" pageNumber="144" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec (1994: 576)</bibRefCitation>
|
||
was the first to show pictures of the
|
||
<typeStatus id="54A881AD595CD202FC04CF1BFBF9B016" box="[926,1029,818,844]" pageId="12" pageNumber="144" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C13448C595CD202FBB5CF1BFABFB016" baseAuthorityName="d'Archiac & de Verneuil" baseAuthorityYear="1842" box="[1071,1347,818,844]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595CD202FBB5CF1BFABFB016" box="[1071,1347,818,844]" italics="true" pageId="12" pageNumber="144">Verneuilia cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
and some
|
||
<typeStatus id="54A881AD595CD202FE98CF7FFE88B02A" box="[258,372,854,880]" pageId="12" pageNumber="144" type="topotype">topotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C13448C595CD202FE0CCF7FFE03B035" box="[406,511,854,879]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FE0CCF7FFE03B035" box="[406,511,854,879]" italics="true" pageId="12" pageNumber="144">V. oceani</emphasis>
|
||
</taxonomicName>
|
||
. He emphasised the presence of a strong vertical crenulation on the interarea of
|
||
<taxonomicName id="4C13448C595CD202FF7BCF50FEB3B0C8" box="[225,335,889,914]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FF7BCF50FEB3B0C8" box="[225,335,889,914]" italics="true" pageId="12" pageNumber="144">V. oceani</emphasis>
|
||
</taxonomicName>
|
||
in contrast to
|
||
<taxonomicName id="4C13448C595CD202FD9DCF50FD47B0C9" baseAuthorityName="d'Archiac & de Verneuil" baseAuthorityYear="1842" box="[519,699,889,915]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595CD202FD9DCF50FD47B0C9" box="[519,699,889,915]" italics="true" pageId="12" pageNumber="144">V. cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
and concluded: “Detailed study of the available material now allows a clear definition and differentiation of the species
|
||
<taxonomicName id="4C13448C595CD202FC3FCFB5FBC4B0EC" baseAuthorityName="d'Archiac & de Verneuil" baseAuthorityYear="1842" box="[933,1080,924,950]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595CD202FC3FCFB5FBC4B0EC" box="[933,1080,924,950]" italics="true" pageId="12" pageNumber="144">cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C13448C595CD202FBE9CFB4FB3DB0EC" box="[1139,1217,925,950]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FBE9CFB4FB3DB0EC" box="[1139,1217,925,950]" italics="true" pageId="12" pageNumber="144">oceani</emphasis>
|
||
</taxonomicName>
|
||
.” (
|
||
<bibRefCitation id="EF8242FE595CD202FB7FCFB5FEC5B080" author="Gourvennec R." pageId="12" pageNumber="144" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec 1994: 581</bibRefCitation>
|
||
). He considered the superfamily
|
||
<taxonomicName id="4C13448C595CD202FD5ECF96FCAEB083" box="[708,850,959,985]" pageId="12" pageNumber="144" rank="superFamily" superFamily="Spiriferacea">Spiriferacea</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="4C13448C595CD202FC79CF96FB77B083" authorityName="Waagen" authorityYear="1883" box="[995,1163,959,985]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="superFamily" superFamily="Reticulariacea">Reticulariacea</taxonomicName>
|
||
as the higher taxon for
|
||
<taxonomicName id="4C13448C595CD202FF76CFCBFE99B0A6" authorityName="Hall & Clarke" authorityYear="1893" box="[236,357,994,1020]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FF76CFCBFE99B0A6" box="[236,357,994,1020]" italics="true" pageId="12" pageNumber="144">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
. At the same time,
|
||
<bibRefCitation id="EF8242FE595CD202FDFCCFCBFC44B0A6" author="Blodgett R. B. & Johnson J. G." box="[614,952,994,1020]" pageId="12" pageNumber="144" pagination="1240 - 1242" refId="ref9786" refString="Blodgett R. B. & Johnson J. G. 1994. First recognition of the genus Verneuilia Hall and Clarke (Brachiopoda, Spiriferida) from North America (West-Central Alaska). Journal of Paleontology 68 (6): 1240 - 1242. https: // doi. org / 10.1017 / S 0022336000034235" type="journal article" year="1994">Blodgett & Johnson (1994)</bibRefCitation>
|
||
described a new species
|
||
<taxonomicName id="4C13448C595CD202FB64CFCBFE89B745" authorityName="Blodgett & Johnson" authorityYear="1994" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="langenstrasseni">
|
||
<emphasis id="B967E31D595CD202FB64CFCBFE89B745" italics="true" pageId="12" pageNumber="144">Verneuilia langenstrasseni</emphasis>
|
||
</taxonomicName>
|
||
from the Middle Devonian of Alaska and placed the family
|
||
<taxonomicName id="4C13448C595CD202FBC0C82CFB06B745" authorityName="Schuchert" authorityYear="1929" box="[1114,1274,1029,1055]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
, including the genera
|
||
<taxonomicName id="4C13448C595CD202FED9C800FE46B719" authorityName="Hall & Clarke" authorityYear="1893" box="[323,442,1065,1091]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FED9C800FE46B719" box="[323,442,1065,1091]" italics="true" pageId="12" pageNumber="144">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C13448C595CD202FE50C800FDB8B719" authorityName="Brunton" authorityYear="1984" box="[458,580,1065,1091]" class="Rhynchonellata" family="Verneuiliidae" genus="Minythyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FE50C800FDB8B719" box="[458,580,1065,1091]" italics="true" pageId="12" pageNumber="144">Minythyra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C13448C595CD202FD18C800FCEFB719" authorityName="Tiazheva" authorityYear="1960" box="[642,787,1065,1091]" class="Rhynchonellata" family="Verneuiliidae" genus="Nuguschella" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FD18C800FCEFB719" box="[642,787,1065,1091]" italics="true" pageId="12" pageNumber="144">Nuguschella</emphasis>
|
||
</taxonomicName>
|
||
, in the superfamily
|
||
<taxonomicName id="4C13448C595CD202FB99C800FA8EB719" authority="George, 1931" authorityName="George" authorityYear="1931" box="[1027,1394,1065,1091]" pageId="12" pageNumber="144" rank="superFamily" superFamily="Ambocoelioidea">
|
||
Ambocoelioidea
|
||
<bibRefCitation id="EF8242FE595CD202FB54C800FA8EB719" author="George T. N." box="[1230,1394,1065,1091]" pageId="12" pageNumber="144" pagination="30 - 61" refId="ref10584" refString="George T. N. 1931. Ambocoelia Hall and certain similar British Spiriferidae. Quarterly Journal of the Geological Society of London 87: 30 - 61. https: // doi. org / 10.1144 / gsl. jgs. 1931.087.01 - 04.05" type="journal article" year="1931">George, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF8242FE595CD202FF25C865FE61B73C" author="Carter J. L. & Johnson J. G. & Gourvennec R. & Hongfei H." box="[191,413,1100,1126]" pageId="12" pageNumber="144" pagination="327 - 374" refId="ref10124" refString="Carter J. L., Johnson J. G., Gourvennec R. & Hongfei H. 1994. A revised classification of the spiriferid brachiopods. Annals of the Carnegie Museum 63: 327 - 374. Available from https: // www. biodiversitylibrary. org / part / 215817 [accessed 10 Oct. 2024]" type="journal article" year="1994">
|
||
Carter
|
||
<emphasis id="B967E31D595CD202FE94C864FEB4B73C" box="[270,328,1100,1126]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
(1994)
|
||
</bibRefCitation>
|
||
supported this view.
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595CD202FF27C8A4FE60B41E" blockId="12.[187,1400,1165,1860]" pageId="12" pageNumber="144">
|
||
<bibRefCitation id="EF8242FE595CD202FF27C8A4FE7AB7FD" author="Sun Y. & Balinski A. & Ma X. & Zhang Y." box="[189,390,1165,1191]" pageId="12" pageNumber="144" pagination="267 - 274" refId="ref12589" refString="Sun Y., Balinski A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267 - 274." type="journal article" year="2004">
|
||
Sun
|
||
<emphasis id="B967E31D595CD202FF69C8A7FECCB7FD" box="[243,304,1165,1191]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
described a new genus
|
||
<taxonomicName id="4C13448C595CD202FD3CC8A4FCA4B7FD" box="[678,856,1165,1191]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FD3CC8A4FCA4B7FD" box="[678,856,1165,1191]" italics="true" pageId="12" pageNumber="144">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
whose
|
||
<typeStatus id="54A881AD595CD202FC29C8A7FC1AB7F2" box="[947,998,1166,1192]" pageId="12" pageNumber="144">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C13448C595CD202FBD7C8A7FA8BB7FD" authority="Sun et al., 2004" authorityName="Sun" authorityYear="2004" box="[1101,1399,1165,1192]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="yangi">
|
||
<emphasis id="B967E31D595CD202FBD7C8A7FB4EB7FD" box="[1101,1202,1166,1192]" italics="true" pageId="12" pageNumber="144">C. yangi</emphasis>
|
||
<bibRefCitation id="EF8242FE595CD202FB23C8A4FA8BB7FD" author="Sun Y. & Balinski A. & Ma X. & Zhang Y." box="[1209,1399,1165,1192]" pageId="12" pageNumber="144" pagination="267 - 274" refId="ref12589" refString="Sun Y., Balinski A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267 - 274." type="journal article" year="2004">
|
||
Sun
|
||
<emphasis id="B967E31D595CD202FB6BC8A7FAD0B7FD" box="[1265,1324,1165,1191]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
had a clearly denticulated hinge line and lacked dental plates. In order to place this genus in the family
|
||
<taxonomicName id="4C13448C595CD202FF27C8FDFEA1B7B4" authorityName="Schuchert" authorityYear="1929" box="[189,349,1236,1262]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
, the authors extended the family diagnosis by combining the characters of
|
||
<taxonomicName id="4C13448C595CD202FB5EC8FDFF02B64B" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="yangi">
|
||
<emphasis id="B967E31D595CD202FB5EC8FDFF02B64B" italics="true" pageId="12" pageNumber="144">Changshunella yangi</emphasis>
|
||
</taxonomicName>
|
||
and “
|
||
<taxonomicName id="4C13448C595CD202FEDCC8DEFDDEB64B" box="[326,546,1271,1297]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FEDCC8DEFE43B64B" box="[326,447,1271,1297]" italics="true" pageId="12" pageNumber="144">Verneuilia</emphasis>
|
||
”
|
||
<emphasis id="B967E31D595CD202FE4EC8D1FDDEB64B" box="[468,546,1272,1297]" italics="true" pageId="12" pageNumber="144">oceani</emphasis>
|
||
</taxonomicName>
|
||
, which are absent in the Devonian
|
||
<typeStatus id="54A881AD595CD202FC5FC8DEFC04B64B" box="[965,1016,1271,1297]" pageId="12" pageNumber="144">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C13448C595CD202FBE4C8DEFB0AB64B" authorityName="Hall & Clarke" authorityYear="1893" box="[1150,1270,1271,1297]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FBE4C8DEFB0AB64B" box="[1150,1270,1271,1297]" italics="true" pageId="12" pageNumber="144">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF8242FE595CD202FA9FC8DEFEF7B66E" author="Sun Y. & Balinski A. & Ma X. & Zhang Y." pageId="12" pageNumber="144" pagination="267 - 274" refId="ref12589" refString="Sun Y., Balinski A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267 - 274." type="journal article" year="2004">
|
||
Sun
|
||
<emphasis id="B967E31D595CD202FAA6C8D1FA8BB64B" box="[1340,1399,1271,1297]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
also failed to determine the superfamilial assignment of the family
|
||
<taxonomicName id="4C13448C595CD202FBB6C933FB34B66E" authorityName="Schuchert" authorityYear="1929" box="[1068,1224,1306,1332]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
and suggested that a new superfamily should be distinguished. Johnson (
|
||
<bibRefCitation id="EF8242FE595CD202FCF7C914FB70B60D" author="Carter J. L. & Johnson J. G. & Gourvennec R. & Hou H." box="[877,1164,1341,1367]" pageId="12" pageNumber="144" pagination="1689 - 1937" refId="ref10185" refString="Carter J. L., Johnson J. G., Gourvennec R. & Hou H. - f. 2006. Spiriferida. In: Moore R. C. (ed.) Treatise on Invertebrate Paleontology. Pt H, Revised Brachiopoda 5: 1689 - 1937. The Geological Society of America, The University of Kansas, Lawrence." type="book chapter" year="2006">
|
||
Carter
|
||
<emphasis id="B967E31D595CD202FC58C917FC01B60D" box="[962,1021,1341,1367]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
2006: 1746
|
||
</bibRefCitation>
|
||
) strongly disagreed with the extension of the family diagnosis and wrote: “The Visean species
|
||
<taxonomicName id="4C13448C595CD202FBAEC948FAC9B620" authority="(d'Orbigny)" baseAuthorityName="d'Orbigny" box="[1076,1333,1376,1402]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FBAEC948FB5CB620" box="[1076,1184,1377,1402]" italics="true" pageId="12" pageNumber="144">V. oceani</emphasis>
|
||
(d’Orbigny)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3047F9F595CD202FF24C9ADFEDFB6C4" box="[190,291,1412,1438]" name="Belgium" pageId="12" pageNumber="144">Belgium</collectingCountry>
|
||
is not congeneric with the
|
||
<typeStatus id="54A881AD595CD202FDFAC9ADFD6FB6C4" box="[608,659,1412,1438]" pageId="12" pageNumber="144">type</typeStatus>
|
||
species, bearing strong dental flanges, a finely striated cardinal process, lacing crural plates, in having a clearly denticulate ventral interarea with wide perideltidial areas, and possibly a weakly capillate microornament. It belongs in the
|
||
<taxonomicName id="4C13448C595CD202FB8FC9E3FB44B6BE" authorityName="Waagen" authorityYear="1883" box="[1045,1208,1482,1508]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="141" phylum="Brachiopoda" rank="superFamily" superFamily="Spiriferoidea">Spiriferoidea</taxonomicName>
|
||
.”. He concluded that these species should not only be placed in different genera, but also in other superfamilies. However, Johnson (
|
||
<bibRefCitation id="EF8242FE595CD202FEB1CA38FDBAB570" author="Carter J. L. & Johnson J. G. & Gourvennec R. & Hou H." box="[299,582,1552,1579]" pageId="12" pageNumber="144" pagination="1689 - 1937" refId="ref10185" refString="Carter J. L., Johnson J. G., Gourvennec R. & Hou H. - f. 2006. Spiriferida. In: Moore R. C. (ed.) Treatise on Invertebrate Paleontology. Pt H, Revised Brachiopoda 5: 1689 - 1937. The Geological Society of America, The University of Kansas, Lawrence." type="book chapter" year="2006">
|
||
Carter
|
||
<emphasis id="B967E31D595CD202FEE4CA38FE45B570" box="[382,441,1552,1578]" italics="true" pageId="12" pageNumber="144">et al.</emphasis>
|
||
2006: 1746
|
||
</bibRefCitation>
|
||
) did not attempt to identify or describe the new genus he had actually discovered.
|
||
<bibRefCitation id="EF8242FE595CD202FED4CA1DFDC0B514" author="Poletaev V. I." box="[334,572,1587,1614]" pageId="12" pageNumber="144" pagination="17 - 18" refId="ref12092" refString="Poletaev V. I. 2015. Novyy rod i semeystvo kamennougol'nykh spiriferid Yevropy. In: Gozhyk P. F. (ed.) Stratotypovi ta oporni rozrizy fanerozois'kykh vidkladiv Ukrainy: suchasnyi stan ta perspektyvy podal'shykh doslidzhen'. Materialy 36 sesii Paleontologichnogo tovarustva Ukrainy: 17 - 18. Kyiv." type="book chapter" year="2015">Poletaev (2015: 17)</bibRefCitation>
|
||
established the new genus
|
||
<taxonomicName id="4C13448C595CD202FC17CA1DFC0CB514" authorityName="Peron & Lesueur" authorityYear="1810" box="[909,1008,1588,1614]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="12" pageNumber="144" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B967E31D595CD202FC17CA1DFC0CB514" box="[909,1008,1588,1614]" italics="true" pageId="12" pageNumber="144">Oceania</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C13448C595CD202FBA0CA1DFB10B517" authorityName="d'Orbigny" authorityYear="1850" box="[1082,1260,1588,1614]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FBA0CA1DFB10B517" box="[1082,1260,1588,1614]" italics="true" pageId="12" pageNumber="144">Spirifer oceani</emphasis>
|
||
</taxonomicName>
|
||
d`Orbigny, 1850 as the
|
||
<typeStatus id="54A881AD595CD202FEC1CA7EFE72B52B" box="[347,398,1623,1649]" pageId="12" pageNumber="144">type</typeStatus>
|
||
species. At the same time, he proposed to establish the new family
|
||
<taxonomicName id="4C13448C595CD202FB69CA7EFA8BB52B" authorityName="Poletaev" authorityYear="2015" box="[1267,1399,1623,1649]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="12" pageNumber="144" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
with the
|
||
<typeStatus id="54A881AD595CD202FEB6CA52FEA3B5CF" box="[300,351,1659,1685]" pageId="12" pageNumber="144">type</typeStatus>
|
||
genus
|
||
<taxonomicName id="4C13448C595CD202FE20CA52FDE1B5CF" authorityName="Peron & Lesueur" authorityYear="1810" box="[442,541,1659,1685]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="12" pageNumber="144" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B967E31D595CD202FE20CA52FDE1B5CF" box="[442,541,1659,1685]" italics="true" pageId="12" pageNumber="144">Oceania</emphasis>
|
||
</taxonomicName>
|
||
and, together with
|
||
<taxonomicName id="4C13448C595CD202FC88CA53FC70B5CE" authorityName="Brunton" authorityYear="1984" box="[786,908,1658,1684]" class="Rhynchonellata" family="Verneuiliidae" genus="Minythyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595CD202FC88CA53FC70B5CE" box="[786,908,1658,1684]" italics="true" pageId="12" pageNumber="144">Minythyra</emphasis>
|
||
</taxonomicName>
|
||
, to place this family in the superfamily
|
||
<taxonomicName id="4C13448C595CD202FF27CAB4FE50B5ED" authorityName="Ivanova" authorityYear="1972" box="[189,428,1693,1719]" pageId="12" pageNumber="144" rank="superFamily" superFamily="Paeckelmanelloidea">Paeckelmanelloidea</taxonomicName>
|
||
. However, the genus name
|
||
<taxonomicName id="4C13448C595CD202FD69CAB7FCAAB5E2" authorityName="Peron & Lesueur" authorityYear="1810" box="[755,854,1694,1720]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="12" pageNumber="144" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B967E31D595CD202FD69CAB7FCAAB5E2" box="[755,854,1694,1720]" italics="true" pageId="12" pageNumber="144">Oceania</emphasis>
|
||
</taxonomicName>
|
||
was already used by
|
||
<bibRefCitation id="EF8242FE595CD202FBCCCAB4FA8AB5ED" author="Peron F. & Lesueur C. A." box="[1110,1398,1693,1720]" pageId="12" pageNumber="144" pagination="325 - 366" refId="ref11736" refString="Peron F. & Lesueur C. A. 1810. Tableau des caracteres generiques et specifiques de toutes les especes de meduses conneus jusqu'a ce jour. Annales du Museum d'Histoire naturelle 14: 325 - 366. Available from https: // www. biodiversitylibrary. org / page / 3498981 [accessed 7 Oct. 2024]." type="journal article" year="1810">Péron & Lesueur (1810)</bibRefCitation>
|
||
and the family
|
||
<taxonomicName id="4C13448C595CD202FEE3CAE9FE01B580" authorityName="Poletaev" authorityYear="2015" box="[377,509,1728,1754]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="12" pageNumber="144" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
was established by Eschcheltz (1829). Both the genus and family names proposed by the author in 2015 are thus preoccupied (junior homonymy). Consequently, the genus with
|
||
<taxonomicName id="4C13448C595CD202FF27CB2EFE97B47B" authorityName="d'Orbigny" authorityYear="1850" box="[189,363,1799,1825]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FF27CB2EFE97B47B" box="[189,363,1799,1825]" italics="true" pageId="12" pageNumber="144">Spirifer oceani</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<typeStatus id="54A881AD595CD202FE0BCB2EFE38B47B" box="[401,452,1799,1825]" pageId="12" pageNumber="144">type</typeStatus>
|
||
species is here renamed as
|
||
<taxonomicName id="4C13448C595CD202FC99CB2EFC65B47B" authorityName="Poletaev" authorityYear="2024" box="[771,921,1799,1825]" genus="Johnsoniana" pageId="12" pageNumber="149" rank="genus" status="nom. nov.">
|
||
<emphasis id="B967E31D595CD202FC99CB2EFC65B47B" box="[771,921,1799,1825]" italics="true" pageId="12" pageNumber="144">Johnsoniana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2545E66595CD202FC3ACB2EFBE8B47B" box="[928,1044,1799,1825]" pageId="12" pageNumber="144">nom. nov.</taxonomicNameLabel>
|
||
and remains
|
||
<typeStatus id="54A881AD595CD202FB35CB2EFB1EB47B" box="[1199,1250,1799,1825]" pageId="12" pageNumber="144">type</typeStatus>
|
||
genus of the family
|
||
<taxonomicName id="4C13448C595CD202FE8ACB03FE64B41E" authorityName="Poletaev" authorityYear="2015" box="[272,408,1834,1860]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="12" pageNumber="144" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595CD203FF27CB42FC85B140" blockId="12.[187,1399,1899,1996]" lastBlockId="13.[188,1399,265,538]" lastPageId="13" lastPageNumber="145" pageId="12" pageNumber="144">
|
||
<bibRefCitation id="EF8242FE595CD202FF27CB42FE09B4DC" author="Mottequin B. & Poty E." box="[189,501,1899,1926]" pageId="12" pageNumber="144" pagination="351 - 371" refId="ref11541" refString="Mottequin B. & Poty E. 2022. Brachiopods from the historical type area of the Visean Stage (Carboniferous, Mississippian; Belgium) and the Vise fauna: preliminary remarks. Palaeobiodiversity and Palaeoenvironments 102: 351 - 371. https: // doi. org / 10.1007 / s 12549 - 021 - 00498 - 9" type="journal article" year="2022">Mottequin & Poty (2022)</bibRefCitation>
|
||
revised old European collections of brachiopods from the historical type locality of Visé, outlining the more correct geographic and stratigraphic positions of this fauna (including the age of “
|
||
<taxonomicName id="4C13448C595CD202FECACB9AFE2CB496" authorityName="d'Orbigny" authorityYear="1850" box="[336,464,1970,1996]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="12" pageNumber="144" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595CD202FECACB9AFE9EB496" box="[336,354,1971,1996]" italics="true" pageId="12" pageNumber="144">V</emphasis>
|
||
.”
|
||
<emphasis id="B967E31D595CD202FE18CB9AFE2CB496" box="[386,464,1971,1996]" italics="true" pageId="12" pageNumber="144">oceani</emphasis>
|
||
</taxonomicName>
|
||
) as latest Warnantian (Late Asbian) and published an image of the
|
||
<typeStatus id="54A881AD595CD202FA8ACB9BFA8BB496" box="[1296,1399,1970,1996]" pageId="12" pageNumber="144" type="topotype">topotype</typeStatus>
|
||
of “
|
||
<taxonomicName id="4C13448C595DD203FF77CD23FE91B279" authorityName="d'Orbigny" authorityYear="1850" box="[237,365,265,291]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595DD203FF77CD23FF03B279" box="[237,255,266,291]" italics="true" pageId="13" pageNumber="145">V</emphasis>
|
||
.”
|
||
<emphasis id="B967E31D595DD203FE85CD23FE91B279" box="[287,365,266,291]" italics="true" pageId="13" pageNumber="145">oceani</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF8242FE595DD203FE1ACD20FD54B279" author="Mottequin B. & Poty E." box="[384,680,265,292]" pageId="13" pageNumber="145" pagination="351 - 371" refId="ref11541" refString="Mottequin B. & Poty E. 2022. Brachiopods from the historical type area of the Visean Stage (Carboniferous, Mississippian; Belgium) and the Vise fauna: preliminary remarks. Palaeobiodiversity and Palaeoenvironments 102: 351 - 371. https: // doi. org / 10.1007 / s 12549 - 021 - 00498 - 9" type="journal article" year="2022">Mottequin & Poty 2022</bibRefCitation>
|
||
: fig. 11j).
|
||
<bibRefCitation id="EF8242FE595DD203FCB3CD23FBF2B279" author="Wang Y. N. & Ma X. P. & Ebbighausen V. & Becker R. T." box="[809,1038,265,292]" pageId="13" pageNumber="145" pagination="19 - 35" refId="ref12778" refString="Wang Y. N., Ma X. P., Ebbighausen V. & Becker R. T. 2023. Spiriferide and Spiriferinide brachiopods from the Frasnian (Upper Devonian) of the Bergisches Land, Germany. Palaontologische Zeitschrift 97: 19 - 35. https: // doi. org / 10.1007 / s 12542 - 022 - 00635 - 5" type="journal article" year="2023">
|
||
Wang
|
||
<emphasis id="B967E31D595DD203FCEDCD23FC4AB279" box="[887,950,265,291]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
published the results of their research on the
|
||
<taxonomicName id="4C13448C595DD203FEEECD05FD75B21C" authority="King, 1946" authorityName="King" authorityYear="1946" box="[372,649,300,326]" class="Rhynchonellata" family="Spiriferidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="family">Spiriferidae King, 1946</taxonomicName>
|
||
fauna of the Frasnian (Late Devonian) Refrath Formation in the Bergisches Land of
|
||
<collectingCountry id="F3047F9F595DD203FE32CD79FDEAB230" box="[424,534,336,362]" name="Germany" pageId="13" pageNumber="145">Germany</collectingCountry>
|
||
, another historical type area for Western European Devonian fauna. They published the first transverse serial sections of two
|
||
<typeStatus id="54A881AD595DD203FC8CCD5AFC74B2D7" box="[790,904,371,397]" pageId="13" pageNumber="145" type="topotype">topotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C13448C595DD203FC35CD5BFB3EB2D6" baseAuthorityName="d'Archiac & de Verneuil" baseAuthorityYear="1842" box="[943,1218,370,396]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595DD203FC35CD5BFB3EB2D6" box="[943,1218,370,396]" italics="true" pageId="13" pageNumber="145">Verneuilia cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
, showing short dental plates and illustrating the ctenophoridium in the ventral valve. They also showed the absence of denticles on the hinge line of the ventral interarea, the crural base and short crus in the dorsal valve, rounded cardinal extremities of mature
|
||
<taxonomicName id="4C13448C595DD203FD1CCDF5FC6BB2AC" baseAuthorityName="d'Archiac & de Verneuil" baseAuthorityYear="1842" box="[646,919,476,502]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595DD203FD1CCDF5FC6BB2AC" box="[646,919,476,502]" italics="true" pageId="13" pageNumber="145">Verneuilia cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
and flank plications in young specimens. These are absent in the type species of
|
||
<emphasis id="B967E31D595DD203FD1DCDD6FC8EB143" box="[647,882,511,537]" italics="true" pageId="13" pageNumber="145">
|
||
<taxonomicName id="4C13448C595DD203FD1DCDD6FCE1B143" authorityName="Poletaev" authorityYear="2024" box="[647,797,511,537]" genus="Johnsoniana" pageId="13" pageNumber="149" rank="genus">Johnsoniana</taxonomicName>
|
||
<taxonomicName id="4C13448C595DD203FCBECE29FC8EB143" authorityName="d'Orbigny" authorityYear="1850" box="[804,882,512,537]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="oceani">oceani</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595DD203FF27CE68FAC5B7D4" blockId="13.[187,1399,577,1167]" pageId="13" pageNumber="145">
|
||
<bibRefCitation id="EF8242FE595DD203FF27CE68FE61B101" author="Wang Y. N. & Ma X. P. & Ebbighausen V. & Becker R. T." box="[189,413,577,603]" pageId="13" pageNumber="145" pagination="19 - 35" refId="ref12778" refString="Wang Y. N., Ma X. P., Ebbighausen V. & Becker R. T. 2023. Spiriferide and Spiriferinide brachiopods from the Frasnian (Upper Devonian) of the Bergisches Land, Germany. Palaontologische Zeitschrift 97: 19 - 35. https: // doi. org / 10.1007 / s 12542 - 022 - 00635 - 5" type="journal article" year="2023">
|
||
Wang
|
||
<emphasis id="B967E31D595DD203FE90CE6BFEBAB101" box="[266,326,577,603]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
proposed an emended diagnosis of the genus
|
||
<taxonomicName id="4C13448C595DD203FC53CE68FBBEB101" authorityName="Hall & Clarke" authorityYear="1893" box="[969,1090,577,603]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FC53CE68FBBEB101" box="[969,1090,577,603]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
and included short dental plates in the ventral valve of
|
||
<taxonomicName id="4C13448C595DD203FD8BCE4CFD3DB124" baseAuthorityName="d'Archiac & de Verneuil" baseAuthorityYear="1842" box="[529,705,612,638]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595DD203FD8BCE4CFD3DB124" box="[529,705,612,638]" italics="true" pageId="13" pageNumber="145">V. cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
, but not the denticulated hinge line of
|
||
<taxonomicName id="4C13448C595DD203FB1DCE4CFB0DB124" authorityName="d'Orbigny" authorityYear="1850" box="[1159,1265,613,638]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595DD203FB1DCE4CFB0DB124" box="[1159,1265,613,638]" italics="true" pageId="13" pageNumber="145">V. oceani</emphasis>
|
||
</taxonomicName>
|
||
. It remains unclear why
|
||
<bibRefCitation id="EF8242FE595DD203FEC9CEA1FDD1B1FB" author="Wang Y. N. & Ma X. P. & Ebbighausen V. & Becker R. T." box="[339,557,647,674]" pageId="13" pageNumber="145" pagination="19 - 35" refId="ref12778" refString="Wang Y. N., Ma X. P., Ebbighausen V. & Becker R. T. 2023. Spiriferide and Spiriferinide brachiopods from the Frasnian (Upper Devonian) of the Bergisches Land, Germany. Palaontologische Zeitschrift 97: 19 - 35. https: // doi. org / 10.1007 / s 12542 - 022 - 00635 - 5" type="journal article" year="2023">
|
||
Wang
|
||
<emphasis id="B967E31D595DD203FE07CEA1FE24B1FB" box="[413,472,647,673]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
did not exclude
|
||
<taxonomicName id="4C13448C595DD203FD6ACEAEFC62B1FB" authorityName="d'Orbigny" authorityYear="1850" box="[752,926,647,673]" class="Rhynchonellata" family="Spiriferidae" genus="Spirifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595DD203FD6ACEAEFC62B1FB" box="[752,926,647,673]" italics="true" pageId="13" pageNumber="145">Spirifer oceani</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C13448C595DD203FC7CCEAEFBA3B1FB" authorityName="Hall & Clarke" authorityYear="1893" box="[998,1119,647,673]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FC7CCEAEFBA3B1FB" box="[998,1119,647,673]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
, as its internal structure does not correspond to the revised diagnosis of the genus. They excluded
|
||
<taxonomicName id="4C13448C595DD203FBA6CE83FA8AB19E" authorityName="Blodgett & Johnson" authorityYear="1994" box="[1084,1398,682,708]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="langenstrasseni">
|
||
<emphasis id="B967E31D595DD203FBA6CE83FA8AB19E" box="[1084,1398,682,708]" italics="true" pageId="13" pageNumber="145">Verneuilia langenstrasseni</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C13448C595DD203FF66CEE7FE89B1B2" authorityName="Hall & Clarke" authorityYear="1893" box="[252,373,718,744]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FF66CEE7FE89B1B2" box="[252,373,718,744]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
because its internal structure does not correspond to the emended diagnosis of the genus. In my opinion, this species, together with
|
||
<emphasis id="B967E31D595DD203FD3DCED8FCD1B051" box="[679,813,753,779]" italics="true" pageId="13" pageNumber="145">Nugushella</emphasis>
|
||
<bibRefCitation id="EF8242FE595DD203FCAECED8FC16B051" author="Tiazheva A. P." box="[820,1002,753,779]" pageId="13" pageNumber="145" pagination="406 - 409" refId="ref12629" refString="Tiazheva A. P. 1960. Novyy vid devonskikh retikulyariid Urala. In: Markovskiy B. P. (ed.) Novyye vidy drevnikh rasteniy i bespozvonochnykh SSSR: 406 - 409. Vsecoyuznyi geologicheskiy institute, Leningrad." type="book chapter" year="1960">Tiazheva, 1960</bibRefCitation>
|
||
, should have been included under the new generic name in the group of so-called “plicated amboceleids” listed by
|
||
<bibRefCitation id="EF8242FE595DD203FBE7CF3CFAC9B074" author="Ivanova E. A." box="[1149,1333,788,815]" pageId="13" pageNumber="145" pagination="264 - 280" refId="ref10886" refString="Ivanova E. A. 1960. Otriad Spiriferida. In: Orlov Y. A. (ed.) Osnovy Paleontologii. Mshanki, Brachiopody. Vol. 7: 264 - 280. Izdatel'stvo Akademii Nauk SSSR, Moscow." type="book chapter" year="1960">Ivanova (1960)</bibRefCitation>
|
||
, only without
|
||
<taxonomicName id="4C13448C595DD203FE85CF1EFE64B00B" authorityName="Hall & Clarke" authorityYear="1893" box="[287,408,823,849]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FE85CF1EFE64B00B" box="[287,408,823,849]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
. The main difference between the internal structures of the Late Devonian (middle Frasnian)
|
||
<taxonomicName id="4C13448C595DD203FEAFCF73FDB5B02E" baseAuthorityName="d'Archiac & de Verneuil" baseAuthorityYear="1842" box="[309,585,858,884]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="cheiropteryx">
|
||
<emphasis id="B967E31D595DD203FEAFCF73FDB5B02E" box="[309,585,858,884]" italics="true" pageId="13" pageNumber="145">Verneuilia cheiropteryx</emphasis>
|
||
</taxonomicName>
|
||
(according to
|
||
<bibRefCitation id="EF8242FE595DD203FD6DCF72FC38B02F" author="Wang Y. N. & Ma X. P. & Ebbighausen V. & Becker R. T." box="[759,964,858,885]" pageId="13" pageNumber="145" pagination="19 - 35" refId="ref12778" refString="Wang Y. N., Ma X. P., Ebbighausen V. & Becker R. T. 2023. Spiriferide and Spiriferinide brachiopods from the Frasnian (Upper Devonian) of the Bergisches Land, Germany. Palaontologische Zeitschrift 97: 19 - 35. https: // doi. org / 10.1007 / s 12542 - 022 - 00635 - 5" type="journal article" year="2023">
|
||
Wang
|
||
<emphasis id="B967E31D595DD203FCDFCF72FC7DB02E" box="[837,897,858,884]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
2023
|
||
</bibRefCitation>
|
||
) and the Visean “
|
||
<taxonomicName id="4C13448C595DD203FB00CF73FA8AB02E" box="[1178,1398,858,885]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="oceani">
|
||
<emphasis id="B967E31D595DD203FB00CF73FAEFB02E" box="[1178,1299,858,884]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
”
|
||
<emphasis id="B967E31D595DD203FAB2CF72FA8AB02E" box="[1320,1398,859,884]" italics="true" pageId="13" pageNumber="145">oceani</emphasis>
|
||
</taxonomicName>
|
||
(according to
|
||
<bibRefCitation id="EF8242FE595DD203FEFECF57FDC5B0C2" author="Gourvennec R." box="[356,569,894,920]" pageId="13" pageNumber="145" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec 1994</bibRefCitation>
|
||
) confirms the necessity to separate the genus
|
||
<taxonomicName id="4C13448C595DD203FBFBCF57FB0BB0C2" authorityName="Poletaev" authorityYear="2024" box="[1121,1271,894,920]" genus="Johnsoniana" pageId="13" pageNumber="145" rank="genus" status="nom. nov.">
|
||
<emphasis id="B967E31D595DD203FBFBCF57FB0BB0C2" box="[1121,1271,894,920]" italics="true" pageId="13" pageNumber="145">Johnsoniana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2545E66595DD203FA9ACF57FA8BB0C2" box="[1280,1399,894,920]" pageId="13" pageNumber="145">nom. nov.</taxonomicNameLabel>
|
||
I agree with
|
||
<bibRefCitation id="EF8242FE595DD203FED4CF88FDDFB0E1" author="Wang Y. N. & Ma X. P. & Ebbighausen V. & Becker R. T." box="[334,547,929,955]" pageId="13" pageNumber="145" pagination="19 - 35" refId="ref12778" refString="Wang Y. N., Ma X. P., Ebbighausen V. & Becker R. T. 2023. Spiriferide and Spiriferinide brachiopods from the Frasnian (Upper Devonian) of the Bergisches Land, Germany. Palaontologische Zeitschrift 97: 19 - 35. https: // doi. org / 10.1007 / s 12542 - 022 - 00635 - 5" type="journal article" year="2023">
|
||
Wang
|
||
<emphasis id="B967E31D595DD203FE0ECF8BFE32B0E1" box="[404,462,929,955]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
that
|
||
<taxonomicName id="4C13448C595DD203FDC0CF88FD2FB0E1" authorityName="Hall & Clarke" authorityYear="1893" box="[602,723,929,955]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FDC0CF88FD2FB0E1" box="[602,723,929,955]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
cannot be placed in the
|
||
<taxonomicName id="4C13448C595DD203FC7DCF88FB51B0E1" authorityName="George" authorityYear="1931" box="[999,1197,929,955]" pageId="13" pageNumber="144" rank="superFamily" superFamily="Ambocoelioidea">Ambocoelioidea</taxonomicName>
|
||
. The
|
||
<typeStatus id="54A881AD595DD203FB73CF88FAE0B0E1" box="[1257,1308,929,955]" pageId="13" pageNumber="145">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C13448C595DD203FF47CFEDFEAAB084" authorityName="Hall & Clarke" authorityYear="1893" box="[221,342,964,990]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FF47CFEDFEAAB084" box="[221,342,964,990]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
has been well distinguished and described by
|
||
<bibRefCitation id="EF8242FE595DD203FCECCFEDFBA7B084" author="Gourvennec R." box="[886,1115,964,990]" pageId="13" pageNumber="145" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec (1994)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF8242FE595DD203FB0ECFECFA8FB084" author="Wang Y. N. & Ma X. P. & Ebbighausen V. & Becker R. T." box="[1172,1395,964,991]" pageId="13" pageNumber="145" pagination="19 - 35" refId="ref12778" refString="Wang Y. N., Ma X. P., Ebbighausen V. & Becker R. T. 2023. Spiriferide and Spiriferinide brachiopods from the Frasnian (Upper Devonian) of the Bergisches Land, Germany. Palaontologische Zeitschrift 97: 19 - 35. https: // doi. org / 10.1007 / s 12542 - 022 - 00635 - 5" type="journal article" year="2023">
|
||
Wang
|
||
<emphasis id="B967E31D595DD203FB45CFECFAE6B084" box="[1247,1306,964,990]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2023)
|
||
</bibRefCitation>
|
||
. Obviously, this genus has weakly plicate flanks, well-developed dental plates and lack of median septum in the ventral valve, non-denticulated hinge line, knob-like ctenophoridium and pair of short plate-like crural bases in the dorsal valve as well as microornament (after
|
||
<bibRefCitation id="EF8242FE595DD203FC39C807FB84B712" author="Gourvennec R." box="[931,1144,1070,1096]" pageId="13" pageNumber="145" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec 1994</bibRefCitation>
|
||
: pl. 1 fig. 6) with very fine, regularly spaced growth lines and randomly arranged tiny pits. All these data allow, in my opinion, to place
|
||
<emphasis id="B967E31D595DD203FEBAC85DFE5DB7D4" box="[288,417,1140,1166]" italics="true" pageId="13" pageNumber="145">Verneuillia</emphasis>
|
||
in the family
|
||
<taxonomicName id="4C13448C595DD203FDDCC85DFC48B7D4" authority="Gourvennec, 1994" authorityName="Gourvennec" authorityYear="1994" box="[582,948,1140,1166]" class="Rhynchonellata" family="Elythynidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="family">
|
||
Elythynidae
|
||
<bibRefCitation id="EF8242FE595DD203FD41C85DFC48B7D4" author="Gourvennec R." box="[731,948,1140,1166]" pageId="13" pageNumber="145" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
of the superfamily
|
||
<taxonomicName id="4C13448C595DD203FB03C85DFAC9B7D4" authorityName="Waagen" authorityYear="1883" box="[1177,1333,1140,1166]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="superFamily" superFamily="Martinioidea">Martinioidea</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595DD203FF27C89CFE93B7A9" blockId="13.[189,1398,1205,1267]" pageId="13" pageNumber="145">
|
||
The next question is what higher taxa could include the genus
|
||
<taxonomicName id="4C13448C595DD203FC0FC89CFBD7B795" authorityName="Poletaev" authorityYear="2024" box="[917,1067,1205,1231]" genus="Johnsoniana" pageId="13" pageNumber="145" rank="genus" status="nom. nov.">
|
||
<emphasis id="B967E31D595DD203FC0FC89CFBD7B795" box="[917,1067,1205,1231]" italics="true" pageId="13" pageNumber="145">Johnsoniana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2545E66595DD203FBA8C89FFB5BB78A" box="[1074,1191,1206,1232]" pageId="13" pageNumber="145">nom. nov.</taxonomicNameLabel>
|
||
and some closely related genera?
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595DD203FF27C933FC6FB580" blockId="13.[187,1399,1306,1755]" pageId="13" pageNumber="145">
|
||
Moderate to strong transverse contours of valves with high triangular area, acute or mucronate cardinal extremities and denticulate hinge line preclude the assignment of
|
||
<taxonomicName id="4C13448C595DD203FC2BC914FBBBB60D" authorityName="Poletaev" authorityYear="2024" box="[945,1095,1341,1367]" genus="Johnsoniana" pageId="13" pageNumber="149" rank="genus">
|
||
<emphasis id="B967E31D595DD203FC2BC914FBBBB60D" box="[945,1095,1341,1367]" italics="true" pageId="13" pageNumber="145">Johnsoniana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C13448C595DD203FBE7C914FAD3B60D" box="[1149,1327,1341,1367]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FBE7C914FAD3B60D" box="[1149,1327,1341,1367]" italics="true" pageId="13" pageNumber="145">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
to the superfamily
|
||
<taxonomicName id="4C13448C595DD203FED5C949FE17B620" authorityName="Waagen" authorityYear="1883" box="[335,491,1376,1402]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="superFamily" superFamily="Martinioidea">Martinioidea</taxonomicName>
|
||
. The absence of regular ribs on the flanks, smooth or with high median ridge sulcus and fold bordered by specific opposite plica do not allow to assign both genera to
|
||
<taxonomicName id="4C13448C595DD203FB4CC9ADFA8EB6C4" authorityName="Waagen" authorityYear="1883" box="[1238,1394,1412,1438]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="141" phylum="Brachiopoda" rank="superFamily" superFamily="Spiriferoidea">Spiriferoidea</taxonomicName>
|
||
, contrary to the opinion of
|
||
<bibRefCitation id="EF8242FE595DD203FE71C98EFD32B69B" author="Gourvennec R." box="[491,718,1447,1473]" pageId="13" pageNumber="145" pagination="573 - 581" refId="ref10655" refString="Gourvennec R. 1994. The genus Verneuilia Hall et Clarke, 1893 (Brachiopoda, Spiriferida). Geobios 27 (5): 573 - 581. https: // doi. org / 10.1016 / s 0016 - 6995 (94) 80253 - x" type="journal article" year="1994">Gourvennec (1994)</bibRefCitation>
|
||
and Johnson (
|
||
<bibRefCitation id="EF8242FE595DD203FCE9C98EFB70B69B" author="Carter J. L. & Johnson J. G. & Gourvennec R. & Hou H." box="[883,1164,1447,1473]" pageId="13" pageNumber="145" pagination="1689 - 1937" refId="ref10185" refString="Carter J. L., Johnson J. G., Gourvennec R. & Hou H. - f. 2006. Spiriferida. In: Moore R. C. (ed.) Treatise on Invertebrate Paleontology. Pt H, Revised Brachiopoda 5: 1689 - 1937. The Geological Society of America, The University of Kansas, Lawrence." type="book chapter" year="2006">
|
||
Carter
|
||
<emphasis id="B967E31D595DD203FC5EC981FC01B69B" box="[964,1021,1447,1473]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
2006: 1746
|
||
</bibRefCitation>
|
||
). On the other hand, most of the main characters of
|
||
<taxonomicName id="4C13448C595DD203FDADC9E3FD31B6BE" authorityName="Poletaev" authorityYear="2024" box="[567,717,1482,1508]" genus="Johnsoniana" pageId="13" pageNumber="149" rank="genus">
|
||
<emphasis id="B967E31D595DD203FDADC9E3FD31B6BE" box="[567,717,1482,1508]" italics="true" pageId="13" pageNumber="145">Johnsoniana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C13448C595DD203FC91C9E3FC41B6BE" box="[779,957,1482,1508]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FC91C9E3FC41B6BE" box="[779,957,1482,1508]" italics="true" pageId="13" pageNumber="145">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
in general are very close to or agree with the diagnosis of the superfamily
|
||
<taxonomicName id="4C13448C595DD203FD1AC9C4FB5FB55D" authority="(Carter et al. 2006: 1812)" baseAuthorityName="Carter et al. 2006:" baseAuthorityPageNumber="1812" baseAuthorityYear="1812" box="[640,1187,1517,1543]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="superFamily" superFamily="Paeckelmanelloidea">
|
||
Paeckelmanelloidea (
|
||
<bibRefCitation id="EF8242FE595DD203FCE0C9C4FB66B55D" author="Carter J. L. & Johnson J. G. & Gourvennec R. & Hou H." box="[890,1178,1517,1543]" pageId="13" pageNumber="145" pagination="1689 - 1937" refId="ref10185" refString="Carter J. L., Johnson J. G., Gourvennec R. & Hou H. - f. 2006. Spiriferida. In: Moore R. C. (ed.) Treatise on Invertebrate Paleontology. Pt H, Revised Brachiopoda 5: 1689 - 1937. The Geological Society of America, The University of Kansas, Lawrence." type="book chapter" year="2006">
|
||
Carter
|
||
<emphasis id="B967E31D595DD203FC57C9C7FBFBB55D" box="[973,1031,1517,1543]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
2006: 1812
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF8242FE595DD203FB35C9C4FA8AB55D" author="Sun Y. & Balinski A. & Ma X. & Zhang Y." box="[1199,1398,1517,1543]" pageId="13" pageNumber="145" pagination="267 - 274" refId="ref12589" refString="Sun Y., Balinski A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267 - 274." type="journal article" year="2004">
|
||
Sun
|
||
<emphasis id="B967E31D595DD203FB7FC9C7FADCB55D" box="[1253,1312,1517,1543]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
assumed the same for
|
||
<taxonomicName id="4C13448C595DD203FE24CA39FD44B570" box="[446,696,1552,1578]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="species" species="yangi">
|
||
<emphasis id="B967E31D595DD203FE24CA39FD44B570" box="[446,696,1552,1578]" italics="true" pageId="13" pageNumber="145">Changshunella yangi</emphasis>
|
||
</taxonomicName>
|
||
, but the presence of unusual opposed ribs on the ventral and dorsal valves of this species led them to reject the assignment of
|
||
<taxonomicName id="4C13448C595DD203FC34CA1DFB9CB514" box="[942,1120,1588,1614]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FC34CA1DFB9CB514" box="[942,1120,1588,1614]" italics="true" pageId="13" pageNumber="145">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C13448C595DD203FB1FCA1DFA88B514" baseAuthorityName="Carter et al. 2006:" baseAuthorityYear="1812" box="[1157,1396,1588,1614]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="149" phylum="Brachiopoda" rank="superFamily" superFamily="Paeckelmanelloidea">Paeckelmanelloidea</taxonomicName>
|
||
. The opposite
|
||
<typeStatus id="54A881AD595DD203FEC0CA7EFE71B52B" box="[346,397,1623,1649]" pageId="13" pageNumber="145">type</typeStatus>
|
||
of plica on both valves of
|
||
<taxonomicName id="4C13448C595DD203FD5BCA7EFC8FB52B" box="[705,883,1623,1649]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FD5BCA7EFC8FB52B" box="[705,883,1623,1649]" italics="true" pageId="13" pageNumber="145">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
links this genus to
|
||
<taxonomicName id="4C13448C595DD203FBC8CA7EFB37B52B" authorityName="Hall & Clarke" authorityYear="1893" box="[1106,1227,1623,1649]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FBC8CA7EFB37B52B" box="[1106,1227,1623,1649]" italics="true" pageId="13" pageNumber="145">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
. On this basis,
|
||
<bibRefCitation id="EF8242FE595DD203FF26CA53FE80B5CE" author="Sun Y. & Balinski A. & Ma X. & Zhang Y." box="[188,380,1658,1684]" pageId="13" pageNumber="145" pagination="267 - 274" refId="ref12589" refString="Sun Y., Balinski A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267 - 274." type="journal article" year="2004">
|
||
Sun
|
||
<emphasis id="B967E31D595DD203FF6ACA52FED5B5CE" box="[240,297,1658,1684]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
suggest that the above genera could form their own superfamily, namely Verneuilioidea. However, they prefer to include
|
||
<taxonomicName id="4C13448C595DD203FDD8CAB4FD08B5ED" box="[578,756,1693,1719]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FDD8CAB4FD08B5ED" box="[578,756,1693,1719]" italics="true" pageId="13" pageNumber="145">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
in their revised family
|
||
<taxonomicName id="4C13448C595DD203FB91CAB4FB5BB5ED" authorityName="Schuchert" authorityYear="1929" box="[1035,1191,1693,1719]" class="Rhynchonellata" family="Verneuiliidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="family">Verneuiliidae</taxonomicName>
|
||
and maintain the verneuiliids as a separate family in the suborder
|
||
<taxonomicName id="4C13448C595DD203FD6DCAE9FC73B580" authorityName="Waagen" authorityYear="1883" box="[759,911,1728,1754]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="subOrder" subOrder="Spiriferidina">Spiriferidina</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAC3F0F595DD200FF27CB2BFD05B13A" blockId="13.[188,1399,1794,1996]" lastBlockId="14.[187,1399,265,608]" lastPageId="14" lastPageNumber="146" pageId="13" pageNumber="145">
|
||
The pentagonal contours and the presence of extraordinary plica-bordered depressions in the central part of both valves of the biconvex shells of “
|
||
<taxonomicName id="4C13448C595DD203FD30CB0FFCF1B41A" authorityName="Peron & Lesueur" authorityYear="1810" box="[682,781,1830,1856]" class="Hydrozoa" family="Oceaniidae" genus="Oceania" kingdom="Animalia" order="Anthoathecata" pageId="13" pageNumber="145" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B967E31D595DD203FD30CB0FFCF1B41A" box="[682,781,1830,1856]" italics="true" pageId="13" pageNumber="145">Oceania</emphasis>
|
||
</taxonomicName>
|
||
” and
|
||
<taxonomicName id="4C13448C595DD203FCCDCB0CFC2DB465" authorityName="Brunton" authorityYear="1984" box="[855,977,1829,1855]" class="Rhynchonellata" family="Verneuiliidae" genus="Minythyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="145" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595DD203FCCDCB0CFC2DB465" box="[855,977,1829,1855]" italics="true" pageId="13" pageNumber="145">Minythyra</emphasis>
|
||
</taxonomicName>
|
||
previously seemed to me sufficient to distinguish a new family
|
||
<taxonomicName id="4C13448C595DD203FD88CB61FD6AB438" authorityName="Poletaev" authorityYear="2015" box="[530,662,1864,1890]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="13" pageNumber="145" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF8242FE595DD203FD40CB61FC79B439" author="Poletaev V. I." box="[730,901,1864,1891]" pageId="13" pageNumber="145" pagination="17 - 18" refId="ref12092" refString="Poletaev V. I. 2015. Novyy rod i semeystvo kamennougol'nykh spiriferid Yevropy. In: Gozhyk P. F. (ed.) Stratotypovi ta oporni rozrizy fanerozois'kykh vidkladiv Ukrainy: suchasnyi stan ta perspektyvy podal'shykh doslidzhen'. Materialy 36 sesii Paleontologichnogo tovarustva Ukrainy: 17 - 18. Kyiv." type="book chapter" year="2015">Poletaev 2015</bibRefCitation>
|
||
) and to include it very conventionally in the superfamily
|
||
<taxonomicName id="4C13448C595DD203FE12CB42FD8BB4DF" baseAuthorityName="Carter et al. 2006:" baseAuthorityYear="1812" box="[392,631,1899,1925]" pageId="13" pageNumber="145" rank="superFamily" superFamily="Paeckelmanelloidea">Paeckelmanelloidea</taxonomicName>
|
||
. In fact, the assignment of this family to
|
||
<taxonomicName id="4C13448C595DD203FB16CB42FA8BB4DF" baseAuthorityName="Carter et al. 2006:" baseAuthorityYear="1812" box="[1164,1399,1899,1925]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="149" phylum="Brachiopoda" rank="superFamily" superFamily="Paeckelmanelloidea">Paeckelmanelloidea</taxonomicName>
|
||
was wrong, because the characters of
|
||
<taxonomicName id="4C13448C595DD203FD08CBA6FCEAB4F3" authorityName="Poletaev" authorityYear="2015" box="[658,790,1935,1961]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="13" pageNumber="145" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
included smooth shells, which were not foreseen in the diagnosis of
|
||
<taxonomicName id="4C13448C595DD203FE39CB9BFD72B496" baseAuthorityName="Carter et al. 2006:" baseAuthorityYear="1812" box="[419,654,1970,1996]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="13" pageNumber="149" phylum="Brachiopoda" rank="superFamily" superFamily="Paeckelmanelloidea">Paeckelmanelloidea</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8242FE595DD203FD27CB9BFC8AB496" author="Ivanova E. A." box="[701,886,1970,1996]" pageId="13" pageNumber="145" pagination="28 - 42" refId="ref10929" refString="Ivanova E. A. 1972. Oshovnye zakonomernosty evolyutsyi spiriferid (Brachiopoda). Paleontologichesky Zhurnal 3: 28 - 42." type="journal article" year="1972">Ivanova (1972)</bibRefCitation>
|
||
. On the other hand, I agree with
|
||
<bibRefCitation id="EF8242FE595DD200FA9ECB9BFEF7B279" author="Sun Y. & Balinski A. & Ma X. & Zhang Y." lastPageId="14" lastPageNumber="146" pageId="13" pageNumber="145" pagination="267 - 274" refId="ref12589" refString="Sun Y., Balinski A., Ma X. & Zhang Y. 2004. New bizarre micro-spiriferid brachiopod from the Early Carboniferous of China. Acta Palaeontologica Polonica 49 (2): 267 - 274." type="journal article" year="2004">
|
||
Sun
|
||
<emphasis id="B967E31D595DD203FAA6CB9AFA8BB496" box="[1340,1399,1970,1996]" italics="true" pageId="13" pageNumber="145">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
and avoid distinguishing a new superfamily. It will be more reasonable to revise the diagnosis of
|
||
<taxonomicName id="4C13448C595ED200FF26CD05FE5BB21C" baseAuthorityName="Carter et al. 2006:" baseAuthorityYear="1812" box="[188,423,300,326]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="149" phylum="Brachiopoda" rank="superFamily" superFamily="Paeckelmanelloidea">Paeckelmanelloidea</taxonomicName>
|
||
to include “rare smooth forms with opposed ribs or plica on both valves”, which will make a valid inclusion of the family
|
||
<taxonomicName id="4C13448C595ED200FDF5CD66FD0FB233" authorityName="Poletaev" authorityYear="2015" box="[623,755,335,361]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="14" pageNumber="146" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
into this superfamily. The family
|
||
<taxonomicName id="4C13448C595ED200FB13CD66FAF1B233" authorityName="Poletaev" authorityYear="2015" box="[1161,1293,335,361]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="14" pageNumber="146" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
includes the genera
|
||
<emphasis id="B967E31D595ED200FED9CD5AFE36B2D6" box="[323,458,371,396]" italics="true" pageId="14" pageNumber="146">Jonsoniana</emphasis>
|
||
,
|
||
<taxonomicName id="4C13448C595ED200FE41CD5BFD71B2D6" box="[475,653,370,396]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595ED200FE41CD5BFD71B2D6" box="[475,653,370,396]" italics="true" pageId="14" pageNumber="146">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C13448C595ED200FD51CD5BFCB9B2D6" authorityName="Brunton" authorityYear="1984" box="[715,837,370,396]" class="Rhynchonellata" family="Verneuiliidae" genus="Minythyra" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595ED200FD51CD5BFCB9B2D6" box="[715,837,370,396]" italics="true" pageId="14" pageNumber="146">Minythyra</emphasis>
|
||
</taxonomicName>
|
||
, each of which has characters of the emended
|
||
<taxonomicName id="4C13448C595ED200FF27CDBFFE50B2EA" baseAuthorityName="Carter et al. 2006:" baseAuthorityYear="1812" box="[189,428,406,432]" class="Rhynchonellata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="149" phylum="Brachiopoda" rank="superFamily" superFamily="Paeckelmanelloidea">Paeckelmanelloidea</taxonomicName>
|
||
. The family excludes
|
||
<taxonomicName id="4C13448C595ED200FD5ACDBFFCC5B2EA" authorityName="Hall & Clarke" authorityYear="1893" box="[704,825,406,432]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595ED200FD5ACDBFFCC5B2EA" box="[704,825,406,432]" italics="true" pageId="14" pageNumber="146">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
because it has tooth plates and no denticulate hinge line. The smooth flanks and the opposite ribs or plica on both valves distinguish
|
||
<taxonomicName id="4C13448C595ED200FB69CD90FA8BB289" authorityName="Poletaev" authorityYear="2015" box="[1267,1399,441,467]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="14" pageNumber="146" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C13448C595ED200FE9ACDF5FE36B2AC" authorityName="Carter" authorityYear="1974" box="[256,458,476,502]" class="Rhynchonellata" family="Strophopleuridae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="family">Strophopleuridae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C13448C595ED200FD92CDF5FD1BB2AC" authorityName="Ivanova" authorityYear="1972" box="[520,743,476,502]" class="Rhynchonellata" family="Paeckelmanellidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="family">Paeckelmanellidae</taxonomicName>
|
||
, because the latter also have a ventral median septum, which is absent in
|
||
<taxonomicName id="4C13448C595ED200FE38CDD6FDD6B143" authorityName="Poletaev" authorityYear="2015" box="[418,554,511,537]" class="Hydrozoa" family="Oceaniidae" kingdom="Animalia" order="Anthoathecata" pageId="14" pageNumber="146" phylum="Cnidaria" rank="family">Oceaniidae</taxonomicName>
|
||
. Furthermore, the Frasnian (Late Devonian)
|
||
<taxonomicName id="4C13448C595ED200FBDACDD6FB45B143" authorityName="Hall & Clarke" authorityYear="1893" box="[1088,1209,511,537]" class="Rhynchonellata" family="Verneuiliidae" genus="Verneuilia" higherTaxonomySource="CoL" kingdom="Animalia" order="Spiriferida" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595ED200FBDACDD6FB45B143" box="[1088,1209,511,537]" italics="true" pageId="14" pageNumber="146">Verneuilia</emphasis>
|
||
</taxonomicName>
|
||
is not less than thirty million years older than the late Tournaisian to Visean
|
||
<taxonomicName id="4C13448C595ED200FCE4CE0BFBE8B166" authorityName="Poletaev" authorityYear="2024" box="[894,1044,546,572]" genus="Johnsoniana" pageId="14" pageNumber="149" rank="genus">
|
||
<emphasis id="B967E31D595ED200FCE4CE0BFBE8B166" box="[894,1044,546,572]" italics="true" pageId="14" pageNumber="146">Johnsoniana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C13448C595ED200FBD1CE0BFB01B166" box="[1099,1277,546,572]" genus="Changshunella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="146" phylum="Brachiopoda" rank="genus">
|
||
<emphasis id="B967E31D595ED200FBD1CE0BFB01B166" box="[1099,1277,546,572]" italics="true" pageId="14" pageNumber="146">Changshunella</emphasis>
|
||
</taxonomicName>
|
||
and could perhaps be a distant ancestor of the latter genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |