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<document id="C662981B099A8E9109D376A82D3A6F2C" ID-DOI="10.5252/zoosystema2024v46a25" ID-ISSN="1638-9387" ID-Zenodo-Dep="13907152" ID-ZooBank="D4AC1F89-AC34-43C4-9761-3F2015A02265" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728463297646" checkinUser="plazi" docAuthor="Segniagbeto, Gabriel Hoinsoudé, Mark-Oliver, Annemarie Ohler, Luca, Mark-Oliver Rödel, Alain, Luca Luiselli &amp; Dubois, Alain" docDate="2024" docId="03F187A4FFBFFFEFA9F1FBCEFEC9FE96" docLanguage="en" docName="zoosystema.2024.46.25.pdf" docOrigin="Zoosystema 46 (25)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a25.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Hoplobatrachus occipitalis Akloa" docType="treatment" docVersion="2" lastPageNumber="640" masterDocId="FFC8FFDCFFB4FFE2AA73FFBDFFEFFFCF" masterDocTitle="Amphibians of Togo: taxonomy, distribution and conservation status" masterLastPageNumber="670" masterPageNumber="631" pageNumber="640" updateTime="1728463900544" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="08ED8A055BD99760AFE3539E8A5EFA45">Amphibians of Togo: taxonomy, distribution and conservation status</mods:title>
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<mods:namePart id="D75194977B6C7FE999E408081728E232">Segniagbeto, Gabriel Hoinsoudé</mods:namePart>
<mods:affiliation id="270D4A185B65FAD01EC4E8660A14AA80">Laboratory of Ecology and Ecotoxicology, Faculty of Sciences, University of Lomé, BP 6057 Lomé, Togo, BP 1515, Lomé (Togo) and Institute for Development, Ecology, Conservation and Cooperation, Rome (Italy) gsegniagbeto @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="3A1B254D9612987BCE480DC1263D053C">Mark-Oliver, Annemarie Ohler</mods:namePart>
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<mods:namePart id="C54CBAD789851801A8A467C4CA44E107">Alain, Luca Luiselli</mods:namePart>
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<mods:namePart id="89AE856D90184CFDAC9EA9D19FEAC13B">Dubois, Alain</mods:namePart>
<mods:affiliation id="58B103BAB90AB646DA4EA489080E79E9">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE-PSL, Université des Antilles, 57 rue Cuvier, CP 30, F- 75005, Paris (France) alain. dubois @ mnhn. fr</mods:affiliation>
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<mods:title id="CD19FF1B218C405E08F096A8E846FE34">Zoosystema</mods:title>
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<mods:date id="883EF39239B9795C9AB33ADE4D09A2CF">2024</mods:date>
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<taxonomicName id="4C584D31FFBFFFE9A9F1FBCEFAB5FB40" ID-CoL="6MBTP" authority="(Gunther, 1859)" authorityName="Akloa" baseAuthorityName="Gunther" baseAuthorityYear="1859" box="[898,1370,1139,1167]" class="Amphibia" family="Dicroglossidae" genus="Hoplobatrachus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="640" phylum="Chordata" rank="species" species="occipitalis">
<emphasis id="B92CEAA0FFBFFFE9A9F1FBCEFB4FFB41" bold="true" box="[898,1184,1139,1166]" italics="true" pageId="11" pageNumber="640">Hoplobatrachus occipitalis</emphasis>
(Günther, 1859)
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(
<figureCitation id="13632A37FFBFFFE9AE35FB29FB79FB61" box="[1094,1174,1172,1198]" captionStart="FIG" captionStartId="12.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1869]" captionTargetId="figure-83@12.[698,1267,765,1318]" captionTargetPageId="12" captionText="FIG. 4. — Representatives of anuran species from Togo in life: A, Sclerophrys maculata (Hallowell, 1854) (Kalaré, Fazao Malfakassa); B, Sclerophrys regularis (Reuss, 1833) (Adjarala); C, Sclerophrys togoensis (Ahl, 1924) (Diguendue); D, Conraua derooi Hulselmans, 1972 (Yikpa-Dzigbe); E, Hoplobatrachus occipitalis (Günther, 1859) (Akloa); F, Hemisus marmoratus (Peters, 1854) (Kpalimé)." figureDoi="http://doi.org/10.5281/zenodo.13907160" httpUri="https://zenodo.org/record/13907160/files/figure.png" pageId="11" pageNumber="640">Fig. 4E</figureCitation>
)
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. —
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<collectingCountry id="F34F7622FFBFFFE9AE49FB70FB80FB2A" box="[1082,1135,1229,1253]" name="Togo" pageId="11" pageNumber="640">Togo</collectingCountry>
</emphasis>
<specimenCount id="9D5EFD3BFFBFFFE9AEFFFB70FB59FB27" box="[1164,1206,1229,1256]" count="1" pageId="11" pageNumber="640" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5EFD3BFFBFFFE9AEB5FB70FB14FB27" box="[1222,1275,1229,1256]" count="36" pageId="11" pageNumber="640" type="female">36 ♀</specimenCount>
; Bado; MNHN- RA-1979.1067-1072, MNHN-RA-1979.1074-1084, MNHN- RA-1979.1086-1103 •
<specimenCount id="9D5EFD3BFFBFFFE9AE61FABFFBD9FAD2" box="[1042,1078,1282,1309]" count="1" pageId="11" pageNumber="640" type="female">1 ♀</specimenCount>
; Kpalimé; MNHN-RA-2006.2178.
</paragraph>
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<smallCapsWord id="8D01A06EFFBFFFE9A95EFA8FFC5FFA87" baselines="1347,1348" box="[813,944,1330,1353]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Description" pageId="11" pageNumber="640">DESCRIPTION</smallCapsWord>
. — Very large frog (SVL
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), with robust and strong body, more or less dorso-ventrally flattened. Snout rounded. Head as long as broad (HW 38-41% SVL; HL 36-41% SVL). Tympanum very large and round (TYD 7-8% SVL). Hind legs strong, rather short (TL 44-47% SVL). Webbing complete, leaving no phalanges free. Back bearing numerous prominent short glandular folds, flanks with warts arranged longitudinally.
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. — The dorsal surface is characterised by its variable colouration (olive, dark gray brown, whitish and gray). The dorsal part of the thigh, tibia and forearm are brown to greenish with dark transverse spots. Dark spots of varying sizes are present on the posterior part of the thigh and tibia. The ventral side is clearer but usually shows dark blotches.
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. — The male is distinguished by a pair of vocal sacks laterally on the throat.
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. — The species
<taxonomicName id="4C584D31FFBFFFE9AE8AF943FC9AF8FF" authorityName="Akloa" baseAuthorityName="Gunther" baseAuthorityYear="1859" class="Amphibia" family="Dicroglossidae" genus="Hoplobatrachus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="640" phylum="Chordata" rank="species" species="occipitalis">
<emphasis id="B92CEAA0FFBFFFE9AE8AF943FC9AF8FF" italics="true" pageId="11" pageNumber="640">Hoplobatrachus occipitalis</emphasis>
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is very widely distributed in all regions of the country. It is both a savannah and forest form with its range extending from west to east Africa including the northern part of southern Africa. It frequents human dwellings and is present even in cities. Numerous observations were made in almost all localities visited. This species is exploited for human consumption in some restaurants in Lomé and some other cities (for very intense harvest of this species in neighbouring
<collectingCountry id="F34F7622FFBFFFE9A9C4F86CFC1BF826" box="[951,1012,2001,2025]" name="Benin" pageId="11" pageNumber="640">Benin</collectingCountry>
, see
<bibRefCitation id="EFC94B43FFBFFFE9AE6EF86CFB0CF825" author="MOHNEKE M. &amp; ONADEKO A. B. &amp; HIRSCHFELD M. &amp; RODEL M. - O." box="[1053,1251,2001,2026]" pageId="11" pageNumber="640" pagination="117 - 128" refId="ref36283" refString="MOHNEKE M., ONADEKO A. B., HIRSCHFELD M. &amp; RODEL M. - O. 2010. - Dried or fried: amphibians in local and regional food markets in West Africa. TRAFFIC Bulletin 22: 117 - 128." type="journal article" year="2010">
Mohneke
<emphasis id="B92CEAA0FFBFFFE9AE0EF86FFB44F826" box="[1149,1195,2001,2025]" italics="true" pageId="11" pageNumber="640">et al.</emphasis>
2010
</bibRefCitation>
). Some communities in the North hunt this species for family food. It was reported in
<collectingCountry id="F34F7622FFB9FFEFAAF6FF4FFF58FEC5" box="[133,183,242,266]" name="Togo" pageId="13" pageNumber="642">Togo</collectingCountry>
by
<bibRefCitation id="EFC94B43FFB9FFEFAAACFF4FFE97FEC5" author="BOURGAT R." box="[223,376,241,266]" pageId="13" pageNumber="642" pagination="597 - 624" refId="ref33022" refString="BOURGAT R. 1979. - Trematodes d'amphibiens du Togo. Bulletin du Museum national d'Histoire naturelle (4 e serie) 3: 597 - 624." type="journal article" year="1979">Bourgat (1979)</bibRefCitation>
from Atakpamé, Missahohé, Gadjagan, Kabou, Kanté, Katobodjo, Kovié, Niamtougou and Lomé, and by
<bibRefCitation id="EFC94B43FFB9FFEFAAF7FE9BFE99FEF1" author="SEGNIAGBETO G. H. &amp; DEKAWOLE J. K. &amp; KETOH G. K. &amp; DENDI D. &amp; LUISELLI L." box="[132,374,294,318]" pageId="13" pageNumber="642" refId="ref38700" refString="SEGNIAGBETO G. H., DEKAWOLE J. K., KETOH G. K., DENDI D. &amp; LUISELLI L. 2022. - Herpetofaunal diversity in a Dahomey Gap savannah of Togo (West Africa): effects of seasons on the populations of amphibians and reptiles. Diversity 14, 964." type="journal volume" year="2022">
Segniagbeto
<emphasis id="B92CEAA0FFB9FFEFAB73FE9AFEC1FEF1" box="[256,302,294,318]" italics="true" pageId="13" pageNumber="642">et al.</emphasis>
(2022)
</bibRefCitation>
from Tetetou to Nagbeto dam area along the Mono River.
</paragraph>
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